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Checkliste der Unterfamilie Scarabaeinae Latreille, 1802 (Pillendreher)
für die Region Ost-Afrika (Afrika)


Klasse
 
Insecta
Linnaeus, 1758
Freikiefler, Insekten, Kerbtiere, Kerfe
743
20447
107483
Familien
Gattungen
Arten
 
Taxom
 

Unterklasse
 
Pterygota
Lang, 1888
Fluginsekten
739
20429
107440
Familien
Gattungen
Arten
 
Taxom
 

Teilklasse
 
Neoptera
van der Wulp, 1890
Neuflügler
708
20177
106478
Familien
Gattungen
Arten
 
Taxom
 

Überordnung
 
Holometabola
Heider, 1889
Holometabolen Insekten
435
15998
93602
Familien
Gattungen
Arten
 
Taxom
 

Ordnung
 
Coleoptera
Linnaeus, 1758
Käfer
119
6705
38199
Familien
Gattungen
Arten
 
Taxom
 

Unterordnung
 
Polyphaga
Emery, 1886
108
6059
32317
Familien
Gattungen
Arten
 
Taxom
 

Teilordnung
 
Scarabaeiformia
Crowson, 1960
14
1053
7187
Familien
Gattungen
Arten
 
Taxom
 

Überfamilie
 
Scarabaeoidea
Latreille, 1802
Blatthornkäfer
14
1053
7187
Familien
Gattungen
Arten
 
Taxom
 
FamilieScarabaeidae
Latreille, 1802
Blatthornkäfer
951
6651
Gattungen
Arten
 
Taxom
 
UnterfamilieScarabaeinae
Latreille, 1802
Pillendreher
112
1894
Gattungen
Arten
 
Taxom
 


Gattungen für Ost-Afrika
112 Gattungen mit 1894 Arten
#ACDEGHIJKLMNOPSTUX
Acanthonitis Janssens, 20123ArtenTaxom
Afrodrepanus Krikken, 19212ArtenTaxom
Aleiantus Olsoufieff, 18984ArtenTaxom
Allogymnopleurus Janssens, 194113ArtenTaxom
Amietina Cambefort, 19892ArtenTaxom
Anachalcos Hope, 19016ArtenTaxom
Anonychonitis Janssens, 19501ArtTaxom
Apotolamprus Olsofieff, 200832ArtenTaxom
Apterepilissus Montreuil, 19584ArtenTaxom
Aptychonitis Janssens, 18951ArtTaxom
Arachnodes Westwood, 201163ArtenTaxom
Caccobius Thomson, 201437ArtenTaxom
Cambefortantus Paulian, 19758ArtenTaxom
Cambefortius Branco, 19362ArtenTaxom
Canthidium Erichson, 18673ArtenTaxom
Canthodimorpha Davis, Schol..., 19991ArtTaxom
Canthon Hoffmannsegg, 18403ArtenTaxom
Catharsiocopris1ArtTaxom
Catharsius Hope, 185774ArtenTaxom
Cheironitis van Lansberge, 19188ArtenTaxom
Circellium Latreille, 17811ArtTaxom
Cleptocaccobius Cambefort, 185710ArtenTaxom
Clypeodrepanus Krikken, 19533ArtenTaxom
Copridaspidus Bloucomont, 18781ArtTaxom
Copris Geoffroy, 191497ArtenTaxom
Coptodactyla Burmeister, 18771ArtTaxom
Coptorhina Hope, 18577ArtenTaxom
Cyptochirus Lesne, 18514ArtenTaxom
Delopleurus Erichson, 18577ArtenTaxom
Deltochilum Eschscholtz, 18931ArtTaxom
Diastellopalpus van Lansberge, 200022ArtenTaxom
Digitonthophagus Balthasar, 201711ArtenTaxom
Dorbignyolus Branco, 19131ArtTaxom
Drepanocerus Kirby, 185114ArtenTaxom
Drepanoplatynus Boucomont, 19211ArtTaxom
Eodrepanus Barbero, Pal..., 18774ArtenTaxom
Epactoides Olsoufieff, 197639ArtenTaxom
Epilissus Blanchard, 202242ArtenTaxom
Epirinus Reiche, 17751ArtTaxom
Euoniticellus Janssens, 195312ArtenTaxom
Euonthophagus Balthasar, 18975ArtenTaxom
Eusaproecius Branco, 19235ArtenTaxom
Garreta Janssens, 185717ArtenTaxom
Gilletellus Janssens, 19371ArtTaxom
Gymnopleurus Illiger, 184324ArtenTaxom
Haroldius Boucomont, 20102ArtenTaxom
Helictopleurus Orbigny, 189879ArtenTaxom
Heliocopris Hope, 201346ArtenTaxom
Heteroclitopus Péringuey, 19015ArtenTaxom
Heteronitis Gillet, 18913ArtenTaxom
Heterosyphus Paulian, 19751ArtTaxom
Hyalonthophagus Palestrini &..., 19109ArtenTaxom
Ixodina Roth, 09ArtenTaxom
Janssensantus Paulian, 19873ArtenTaxom
Kheper Janssens, 187121ArtenTaxom
Krikkenius Branco, 19821ArtTaxom
Latodrepanus Krikken, 19535ArtenTaxom
Liatongus Reitter, 191414ArtenTaxom
Litocopris Waterhouse, 19613ArtenTaxom
Lophodonitis Janssens, 20112ArtenTaxom
Madateuchus Paulian, 19531ArtTaxom
Megalonitis Janssens, 18553ArtenTaxom
Megaponerophilus Janssens, 19141ArtTaxom
Metacatharsius Paulian, 196543ArtenTaxom
Milichus Péringuey, 190411ArtenTaxom
Mimonthophagus Balthasar, 19054ArtenTaxom
Nanos Westwood, 201144ArtenTaxom
Neateuchus Gillet, 1911
0ArtenTaxom
Neonitis Péringuey, 19182ArtenTaxom
Neosaproecius Branco, 19901ArtTaxom
Neosisyphus Müller, 189116ArtenTaxom
Nesosisyphus Vinson, 19394ArtenTaxom
Nesovinsonia Martinez & P..., 19391ArtTaxom
Nitiocellus Branco, 19391ArtTaxom
Odontoloma Boheman, 19084ArtenTaxom
Oniticellus Serville, 18519ArtenTaxom
Onitis Fabricius, 2012107ArtenTaxom
Onthophagus Latreille, 1947Kotfresser562ArtenTaxom
Pachylomera Griffith & P..., 18742ArtenTaxom
Pachysoma MacLeay, 19071ArtTaxom
Panelus Lewis, 19751ArtTaxom
Paracanthon Balthasar, 19381ArtTaxom
Paronitis1ArtTaxom
Pedaria Laporte, 193953ArtenTaxom
Phalops Erichson, 200324ArtenTaxom
Phanaeus MacLeay, 18191ArtTaxom
Pinacopodius Branco, 20072ArtenTaxom
Pinacotarsus Harold, 19692ArtenTaxom
Platyonitis Janssens, 19742ArtenTaxom
Pleuronitis Lansberge, 20112ArtenTaxom
Proagoderus Lansberge, 185187ArtenTaxom
Pseudochironitis Ferreira, 18973ArtenTaxom
Pseudocopris Ferreira, 19102ArtenTaxom
Pseudosaproecius Balthasar, 18845ArtenTaxom
Sarophorus Erichson, 20033ArtenTaxom
Scaptocnemis Péringuey, 19011ArtTaxom
Scarabaeus Linnaeus, 190161ArtenTaxom
Sceliages Westwood, 18954ArtenTaxom
Sisyphus Latreille, 201762ArtenTaxom
Sphaerocanthon Olsoufieff, 1947
0ArtenTaxom
Stiptocnemis Branco, 19912ArtenTaxom
Stiptopodius Harold, 19018ArtenTaxom
Sukelus Branco, 19911ArtTaxom
Sulcodrepanus Krikken, 18401ArtTaxom
Tanzanolus Scholtz & Ho..., 19601ArtTaxom
Tibiodrepanus Krikken, 18401ArtTaxom
Tiniocellus Péringuey, 18513ArtenTaxom
Tomogonus d´Orbigny, 20125ArtenTaxom
Tragiscus Klug, 18551ArtTaxom
Tropidonitis Janssens, 18571ArtTaxom
Unidentis1ArtTaxom
Xinidium Harold, 18782ArtenTaxom


Artenliste für Ost-Afrika
1894 Arten und 79 Unterarten
Acanthonitis nicolasi Cambefort, 1995
Acanthonitis oberthuri Janssens, 1937
Acanthonitis spissus Josso, 2012
Afrodrepanus impressicollis Boheman, 1857
Afrodrepanus marshalli Boucomont, 1921
Aleiantus fiorii (Paulian, 1976)⇒ Epactoides fiorii Paulian, 1976
Aleiantus incertus (Lebis, 1953)⇒ Epactoides incertus Lebis, 1953
Aleiantus perinetanus (Paulian, 1975)⇒ Epactoides perinetanus Paulian, 1975
Aleiantus perrieri (Fairmaire, 1898)⇒ Epactoides perrieri Fairmaire, 1898
Allogymnopleurus aeneus Harold, 1875
Allogymnopleurus alluaudi Garreta, 1914
Allogymnopleurus anthracinus Klug, 1845
Allogymnopleurus chloris Klug, 1855
Allogymnopleurus histrio Balthasar, 1963
Allogymnopleurus indigaceus Reiche, 1847
Allogymnopleurus infranitens Fairmaire, 1887
Allogymnopleurus jeanneli Garreta, 1914
Allogymnopleurus signaticollis Waterhouse, 1890
Allogymnopleurus splendidus Bertoloni, 1849
Allogymnopleurus thalassinus Klug, 1855
Allogymnopleurus umbrinus Gerstaecker, 1871
Allogymnopleurus zavattarii Müller, 1941
Amietina cambeforti Branco, 1989
Amietina nyassalandica Branco, 1989
Anachalcos aurescens Bates, 1888
Anachalcos convexus Boheman, 1857
Anachalcos cupreus Fabricius, 1775
Anachalcos mfwangani ♰ R. Paulian, 1976
Anachalcos procerus Gerstaecker, 1871
Anachalcos spectabilis (Péringuey, 1901)⇒ Chalconotus procerus Gerstaecker, 1871
Anonychonitis freyi Janssens, 1950
Apotolamprus ambohitsitondronensis Lebis, 1953
Apotolamprus andranobeensis Montreuil & ..., 2012
Apotolamprus antsingyensis Montreuil, 2008
Apotolamprus bemarahaensis Montreuil, 2008
Apotolamprus cyanescens Brancsik, 1892
Apotolamprus darainaensis Montreuil, 2008
Apotolamprus densepunctatus Montreuil, 2008
Apotolamprus hagai Montreuil & ..., 2012
Apotolamprus hanskii Montreuil, 2005
Apotolamprus helenae Montreuil, 2004
Apotolamprus latipennis Boucomont, 1937
Apotolamprus mahajangaensis Montreuil, 2008
Apotolamprus manjarivoloensis Montreuil, 2008
Apotolamprus manomboensis Montreuil, 2008
Apotolamprus marojejyensis Montreuil, 2005
Apotolamprus metallicus Montreuil, 2008
Apotolamprus milloti Lebis, 1953
Apotolamprus nanoides Paulian, 1976
Apotolamprus orangeaensis Montreuil, 2008
Apotolamprus perinetensis Montreuil, 2008
Apotolamprus peyrierasi Paulian, 1985
Apotolamprus pseudomanomboensis Montreuil, 2008
Apotolamprus pseudoquadrimaculatus Montreuil & ..., 2012
Apotolamprus quadrimaculatus Lebis, 1953
Apotolamprus quadrinotatus Boucomont, 1937
Apotolamprus resplendens Montreuil, 2008
Apotolamprus sahatezaensis Montreuil & ..., 2012
Apotolamprus sericeus Paulian, 1976
Apotolamprus sinuatotibialis Montreuil, 2008
Apotolamprus sparsepunctatus Montreuil, 2008
Apotolamprus vadoni Paulian, 1976
Apotolamprus zombitsyensis Montreuil, 2008
Apterepilissus analavelonae Paulian, 1975
Apterepilissus centralis Paulian, 1976
Apterepilissus ovalis Felsche, 1911
Apterepilissus sericeus Lebis, 1958
Aptychonitis anomalus Gestro, 1895
Arachnodes abadiei Lebis, 1953
Arachnodes aeneoviridis Boucomont, 1937
Arachnodes ampasindavae Paulian, 1976
Arachnodes andriai Paulian, 1976
Arachnodes angulosus Boucomont, 1937
Arachnodes bicolor Montreuil, 2011
Arachnodes biimpressus Lebis, 1953
Arachnodes brunnipes Castelnau, 1840
Arachnodes delaunayi Lebis, 1953
Arachnodes delphinensis Lebis, 1953
Arachnodes dichrous Paulian, 1976
Arachnodes didiensis Lebis, 1953
Arachnodes dieganus Paulian, 1976
Arachnodes fairmairei Boucomont, 1937
Arachnodes fulgens Kunckel, 1887
Arachnodes fusconitens Fairmaire, 1899
Arachnodes fuscovirens Fairmaire, 1889
Arachnodes globuloides Paulian, 1976
Arachnodes globulosus (Kunckel, 1887)⇒ Arachnodes globuloides Paulian, 1976
Arachnodes grossepunctatus Paulian, 1986
Arachnodes hanskii Montreuil, 2003
Arachnodes histeroides Castelnau, 1840
Arachnodes insularis Paulian, 1976
Arachnodes jeanneli Lebis, 1953
Arachnodes kelifelyi Paulian, 1976
Arachnodes luctuosus Boucomont, 1937
Arachnodes mahafalyensis Montreuil, 2006
Arachnodes manaitrai Montreuil, 2006
Arachnodes manaitrai kirindyensis Montreuil, 2006
Arachnodes manaitrai manaitrai
Arachnodes manomboensis Montreuil, 2006
Arachnodes mantillerii Montreuil, 2003
Arachnodes micans Lebis, 1953
Arachnodes minutus Paulian, 1976
Arachnodes morati Paulian, 1975
Arachnodes morio Harold, 1879
Arachnodes neonitidus Montreuil, 2011
Arachnodes nitidus Castelnau, 1840
Arachnodes nosybeensis Lebis, 1961
Arachnodes oberthuri Boucomont, 1937
Arachnodes philippi Lebis, 1953
Arachnodes pierrettae Lebis, 1953
Arachnodes pillula Paulian, 1976
Arachnodes pusillus Lebis, 1953
Arachnodes refulgens Fairmaire, 1901
Arachnodes renaudi Lebis, 1958
Arachnodes robinsoni Boucomont, 1937
Arachnodes rubrolimbatus Boucomont, 1937
Arachnodes rubrotinctus Lebis, 1953
Arachnodes sakarahae Paulian, 1976
Arachnodes saprinoides Fairmaire, 1889
Arachnodes semichalceus Lebis, 1953
Arachnodes seminitidus Lebis, 1953
Arachnodes seyrigi Lebis, 1953
Arachnodes soganus Paulian, 1976
Arachnodes splendidus Lebis, 1953
Arachnodes tibialis Lebis, 1956
Arachnodes variolosus Lebis, 1953
Arachnodes vicinus Castelnau, 1840
Arachnodes viettei Montreuil, 2011
Arachnodes vieui Paulian, 1976
Arachnodes vigilans Lebis, 1953
Arachnodes zombitsyensis Montreuil, 2011
Caccobius atomarius (Balthasar, 1935)⇒ Caccobius brevis Waterhouse, 1875
Caccobius auberti d´Orbigny, 1902
Caccobius brevis Waterhouse, 1875
Caccobius cabellai Walter, 2019
Caccobius castaneus (Klug, 1855)⇒ Caccocnemus castaneus Klug, 1855
Caccobius cavatus d´Orbigny, 1908
Caccobius convexifrons (Raffray, 1877)⇒ Caccophilus convexifrons Raffray, 1877
Caccobius corniceps d´Orbigny, 1913
Caccobius croceocinctus d´Orbigny, 1913
Caccobius curvicornis Walter & Cam..., 1977
Caccobius cuspidiger d´Orbigny, 1913
Caccobius discrepans Péringuey, 1901
Caccobius ferrugineus Fahraeus, 1857
Caccobius fuliginosus (Roth, 1851)⇒ Caccophilus reticuliger d´Orbigny, 1904
Caccobius gananensis d´Orbigny, 1904
Caccobius globaticeps d´Orbigny, 1905
Caccobius histerinus Fahraeus, 1857
Caccobius inconspicuus (Fahraeus, 1857)⇒ Caccophilus fuliginosus Roth, 1851
Caccobius inops Péringuey, 1901
Caccobius jossoi Walter, 2014
Caccobius krikkeni Cambefort, 1980
Caccobius longipennis d´Orbigny, 1904
Caccobius megaponerae (Brauns, 1914)⇒ Megaponerophilus megaponerae Brauns, 1914
Caccobius meridionalis Boucomont, 1914
Caccobius nigritulus Klug, 1855
Caccobius obtusus Fahraeus, 1857
Caccobius ocellipennis d´Orbigny, 1913
Caccobius pentagonus d´Orbigny, 1908
Caccobius postlulatus
Caccobius postlutatus d´Orbigny, 1905
Caccobius pseudocorniceps Josso, 2018
Caccobius reticuliger d´Orbigny, 1904
Caccobius semiluteus d´Orbigny, 1905
Caccobius signatipennis Harold, 1867
Caccobius tuberifrons d´Orbigny, 1902
Caccobius ugandicus Frey, 1975
Caccobius zambiensis Walter, 2014
Cambefortantus ankaratrae Paulian, 1975
Cambefortantus blanci Paulian, 1975
Cambefortantus helenae Montreuil, 2008
Cambefortantus micros Paulian, 1986
Cambefortantus myops Lebis, 1953
Cambefortantus pauliani Montreuil, 2008
Cambefortantus ranomafanensis Montreuil, 2008
Cambefortantus ranomandryi Paulian, 1975
Cambefortius dudleyi Branco, 1989
Cambefortius turneri Boucomont, 1936
Canthidium gerstaeckeri Harold, 1867
Canthidium muticum Boheman, 1858
Canthidium smaragdinum Harold, 1867
Canthodimorpha lawrencei Davis, Schol..., 1999
Canthon (Boreocanthon) praticola Le Conte, 1859
Canthon dives Harold, 1868
Canthon rutilans (Castelnau, 1840)⇒ Francmonrosia rutilans Castelnau, 1840
Catharsiocopris quadrituberculatus Balthasar, 1967
Catharsius achates Olivier, 1789
Catharsius adamastor Gillet, 1932
Catharsius aegeus Génier, 2017
Catharsius alpheus Gillet, 1932
Catharsius bradshawi Lansberge, 1887
Catharsius brittoni Ferreira, 1963
Catharsius brutus Harold, 1880
Catharsius calaharicus Kolbe, 1893
Catharsius chinai Ferreira, 1960
Catharsius coptorhinaformis Josso & Prév..., 2009
Catharsius disseptus (Péringuey, 1901)⇒ Catharsius calaharicus Kolbe, 1893
Catharsius dubius (Péringuey, 1901)⇒ Catharsius molossus Linnaeus, 1758
Catharsius duciformis Ferreira, 1959
Catharsius dux Harold, 1878
Catharsius erechtheus Felsche, 1907
Catharsius fastidiosus Thomson, 1858
Catharsius folicornis Josso & Prév..., 2019
Catharsius furcillatus Guérin-Ménev..., 1847
Catharsius furcillatus furcillatus Guérin, 1847
Catharsius furcillatus laeviplaga G. Müller, 1941
Catharsius gorilla Thomson, 1858
Catharsius guineensis Ferreira, 1963
Catharsius haafi Ferreira, 1960
Catharsius haroldi Takano, 2021
Catharsius harpagus Harold, 1877
Catharsius heros Boheman, 1860
Catharsius hertli Ferreira, 1964
Catharsius jacksoni Waterhouse, 1891
Catharsius juheli Josso, 2011
Catharsius karlwerneri Moretto, 2008
Catharsius kolbei Felsche, 1907
Catharsius laticeps Boheman, 1857
Catharsius latifossa Müller, 1941
Catharsius longiceps Gillet, 1907
Catharsius marcellus Kolbe, 1893
Catharsius melancholicus (Boheman, 1860)⇒ Catharsius longiceps Gillet, 1907
Catharsius minutus (Lansberge, 1882)⇒ Metacatharsius minutus Lansberge, 1882
Catharsius mirabilis Felsche, 1901
Catharsius molossus Linnaeus, 1758Dickkopf
Catharsius morettoi Josso & Prév..., 2009
Catharsius mossambicanus Ferreira, 1960
Catharsius mulanjeensis Josso & Prév..., 2019
Catharsius neptunus Kolbe, 1893
Catharsius ninus Gillet, 1918
Catharsius oedipus Felsche, 1907
Catharsius omoensis Müller, 1941
Catharsius pallas Génier, 2017
Catharsius pandion Harold, 1877
Catharsius parafastidiosus Ferreira, 1971
Catharsius phidias (Olivier, 1789)⇒ Catharsius platicornis Ferreira, 1960
Catharsius philus Kolbe, 1893
Catharsius pithecius Fabricius, 1775Indianische Schulteraale
Catharsius platicornis Ferreira, 1960
Catharsius platycerus Klug, 1855
Catharsius pseudocongolensis Ferreira, 1963
Catharsius pseudoplaticornis Josso & Prév..., 2019
Catharsius pseudovitulus Müller, 1941
Catharsius punctatus Josso & Prév..., 2019
Catharsius rhinoceros Klug, 1855
Catharsius satyrus Kolbe, 1893
Catharsius scopas Felsche, 1907
Catharsius sesostris Waterhouse, 1888
Catharsius severini Gillet, 1907
Catharsius simillimus Ferreira, 1960
Catharsius somalicus Gillet, 1918
Catharsius spectabilis Gillet, 1918
Catharsius stricticornis Josso & Prév..., 2019
Catharsius stuhlmanni Kolbe, 1893
Catharsius takanoi Josso & Prév..., 2019
Catharsius tricornutus DeGeer, 1778
Catharsius tridens Josso & Prév..., 2009
Catharsius ugandicus Gillet, 1918
Catharsius ulysses Boheman, 1857
Catharsius vitulus Boheman, 1857
Cheironitis dentifemoralis Ferreira, 1976
Cheironitis flabellatus Boucomont, 1923
Cheironitis hungaricus Herbst, 1789
Cheironitis imitator Balthasar, 1963
Cheironitis muelleri Janssens, 1943
Cheironitis osiridis Reiche, 1856
Cheironitis ungaricus ungaricus Herbst, 1789
Cheironitis viridicans Gillet, 1918
Circellium bacchus Fabricius, 1781Riesen-Pillendreher
Cleptocaccobius biceps d´Orbigny, 1905
Cleptocaccobius convexifrons Raffray, 1877
Cleptocaccobius humilis d´Orbigny, 1902
Cleptocaccobius postlutatus d´Orbigny, 1905
Cleptocaccobius schaedlei d´Orbigny, 1902
Cleptocaccobius semiluteus d´Orbigny, 1905
Cleptocaccobius seminulum Klug, 1855
Cleptocaccobius signaticollis d´Orbigny, 1902
Cleptocaccobius uniseries d´Orbigny, 1905
Cleptocaccobius viridicollis Fahraeus, 1857
Clypeodrepanus digitatus Krikken, 2009
Clypeodrepanus striatus Boucomont, 1921
Clypeodrepanus strigatus Janssens, 1953
Copridaspidus peregrinus Harold, 1878
Copris achates (Péringuey, 1901)⇒ Copris cornifrons Boheman, 1860
Copris afgoi Cambefort, 1992
Copris algol Nguyen-phung, 1989
Copris amabilis Kolbe, 1914
Copris amyntor (Klug, 1855)⇒ Copris harrisi Waterhouse, 1891
Copris anceus Olivier, 1789
Copris antares Bacescu & Ca..., 1947
Copris armiger Gillet, 1910
Copris atropolitus Gillet, 1910
Copris bootes Klug, 1855
Copris brittoni Ferreira, 1962
Copris caelatus Fabricius, 1794
Copris camerunus Felsche, 1904
Copris carmelita Fabricius, 1801
Copris cassius Péringuey, 1901
Copris complexus Nguyen-phung, 1989
Copris coriarius Gillet, 1907
Copris cornifrons Boheman, 1860
Copris corniger (Sahlberg, 1823)⇒ Onthophagus taurus Schreber, 1759
Copris cribricollis (Gillet, 1910)⇒ Catharsius pithecius Fabricius, 1775
Copris curlettii Moretto, 2011
Copris decellei Cambefort, 1992
Copris denticulatus Nguyen-phung, 1988
Copris diversus Waterhouse, 1891
Copris dracunculus Fereira, 1959
Copris dudleyi Cambefort, 1992
Copris elpheneroides Felsche, 1910
Copris elphenor Klug, 1855
Copris evanidus Klug, 1855
Copris fallaciosus Gillet, 1907
Copris fidius Olivier, 1789
Copris gazellarum Gillet, 1918
Copris gilleti Kolbe, 1907
Copris gracilis Waterhouse, 1891
Copris graueri (Kolbe, 1914)⇒ Copris camerunus Felsche, 1904
Copris harrisi Waterhouse, 1891
Copris hartli Frey, 1975
Copris humilis Gillet, 1908
Copris hybridus Gillet, 1910
Copris inhalatus Quedenfeldt, 1884
Copris inhalatus grandis Nguyen-phung..., 1986
Copris inhalatus inhalatus
Copris inhalatus perturbator Péringuey, 1901
Copris inhalatus sanctaeluciae Nguyen-phung..., 1986
Copris insidiosus Péringuey, 1901
Copris integer (Reiche, 1847)⇒ Copris fallaciosus Gillet, 1907
Copris interioris Kolbe, 1897
Copris jacchus Lang, 1971
Copris jahi Nguyen-phung..., 1986
Copris jucundus Gillet, 1932
Copris kasanka Josso & Prév..., 2016
Copris laiiformis Nguyen-phung..., 1987
Copris laioides Boucomont, 1932
Copris laius angusticornis Nguyen-phung..., 1987
Copris leakeyorum ♰ R. Paulian, 1976
Copris macer Péringuey, 1901
Copris martinae Nguyen-phung, 1988
Copris martinae martinae
Copris martinae ntchisi Moretto, 2011
Copris mesacanthus Harold, 1878
Copris mesacanthus mesacanthus
Copris misellus Péringuey, 1901
Copris moffartsi Gillet, 1907
Copris montivagus (Gillet, 1908)⇒ Copris harrisi Waterhouse, 1891
Copris morettoi Josso & Prév..., 2020
Copris morphaeus Gillet, 1932
Copris mourgliai Zelenka, 1993
Copris muticus (Boheman, 1857)⇒ Litocopris muticus Boheman, 1857
Copris nepos Gillet, 1908
Copris obesus Boheman, 1857
Copris orion Klug, 1835
Copris orion orion
Copris orion overlaeti Balthasar, 1961
Copris orphanus (Guérin-Ménev..., 1847)⇒ Copris evanidus Klug, 1855
Copris prevosti Josso, 2011
Copris pseudobootes Ferreira, 1962
Copris puncticollis Boheman, 1857
Copris punctiventris Waterhouse, 1891
Copris ruandanus Balthasar, 1961
Copris rugosus (Gillet, 1908)⇒ Copris inhalatus Quedenfeldt, 1884
Copris sarpedon Harold, 1868
Copris scorpio Zelenka, 1993
Copris serius Nguyen-phung, 1987
Copris sexdentatus (Thunberg, 1818)⇒ Copris sarpedon Harold, 1868
Copris simonettai Cambefort, 1992
Copris singularis Nguyen-phung..., 1986
Copris sphaeropterus Harold, 1877
Copris subsidens Péringuey, 1901
Copris typhoeus Gerstaecker, 1884
Copris urus (Boheman, 1857)⇒ Catharsius molossus Linnaeus, 1758
Copris usambaricus Gillet, 1908
Copris vankhaii Nguyen-phung, 1988
Copris vankhaii genopunctatus Nguyen-phung, 1988
Copris vankhaii vankhaii
Copris vankhalii Nguyen-phung, 1988
Copris vilhenai Ferreira, 1962
Copris wiesei Kolbe, 1914
Coptodactyla subaenea Harold, 1877
Coptorhina auspicata Péringuey, 1901
Coptorhina davidi Janssens, 1939
Coptorhina excavata Frolov, Akhm..., 2008
Coptorhina klugi Hope, 1833
Coptorhina klugii Hope, 1833
Coptorhina nitefacta Gillet, 1932
Coptorhina nitidipennis Boheman, 1857
Cyptochirus ambiguus (Kirby, 1828)⇒ Drepanocerus trogiformis Roth, 1851
Cyptochirus decellei Simonis & Zu..., 1980
Cyptochirus distinctus Janssens, 1953
Cyptochirus trogiformis Roth, 1851
Delopleurus darrenmanni Frolov, 2014
Delopleurus fossatus Frolov, 2014
Delopleurus gilletti Janssens, 1939
Delopleurus krikkeni Frolov, 2014
Delopleurus naviauxi Frolov & Cam..., 2014
Delopleurus pubescens Frolov, 2014
Delopleurus pullus Boheman, 1857
Deltochilum enceladus Kolbe, 1893
Diastellopalpus basilobatus d´Orbigny, 1902
Diastellopalpus cambeforti Josso & Pré..., 2000
Diastellopalpus dudleyi Cambefort, 1980
Diastellopalpus fuelleborni Kolbe, 1901
Diastellopalpus gilleti d´Orbigny, 1908
Diastellopalpus ikelenge Josso & Pré..., 2014
Diastellopalpus infernalis Lansberge, 1887
Diastellopalpus johnstoni Waterhouse, 1885
Diastellopalpus laevibasis d´Orbigny, 1902
Diastellopalpus lamellicollis Quedenfeldt, 1884
Diastellopalpus metriogonus (d´Orbigny, 1914)⇒ Diastellopalpus neavei d´Orbigny, 1914
Diastellopalpus monapoides Bates, 1888
Diastellopalpus murrayi Harold, 1868
Diastellopalpus neavei d´Orbigny, 1914
Diastellopalpus quinquedens Bates, 1888
Diastellopalpus semirubidus d´Orbigny, 1913
Diastellopalpus sulciger Kolbe, 1893
Diastellopalpus thomsoni Bates, 1888
Diastellopalpus thomsoni malawiensis Josso & Pré..., 2000
Diastellopalpus thomsoni thomsoni
Diastellopalpus tridens (Fairmaire, 1891)⇒ Diastellopalpus thomsoni Bates, 1888
Diastellopalpus walteri Josso & Pré..., 2000
Digitonthophagus aksumensis Génier, 2017
Digitonthophagus biflagellatus Génier, 2017
Digitonthophagus catta Fabricius, 1787
Digitonthophagus eucatta Génier, 2017
Digitonthophagus falciger Génier, 2017
Digitonthophagus fimator Génier, 2017
Digitonthophagus gazella Fabricius, 1787
Digitonthophagus lusinganus Orbigny, 1905
Digitonthophagus petilus Génier, 2017
Digitonthophagus sahelicus Moretto, 2017
Digitonthophagus ulcerosus Génier, 2017
Dorbignyolus mirecribratus d´Orbigny, 1913
Drepanocerus abyssinicus (Roth, 1851)⇒ Ixodina abyssinica Roth, 1851
Drepanocerus ambiguus Kirby, 1828
Drepanocerus bechynei (Janssens, 1953)⇒ Eodrepanus bechynei Janssens, 1953
Drepanocerus fastiditus (Péringuey, 1901)⇒ Eodrepanus fastiditus Péringuey, 1901
Drepanocerus kirbyi Kirby, 1828
Drepanocerus laticollis (Fahraeus, 1857)⇒ Latodrepanus laticollis Fahraeus, 1857
Drepanocerus marshalli (Boucomont, 1921)⇒ Afrodrepanus marshalli Boucomont, 1921
Drepanocerus orientalis Krikken, 2009
Drepanocerus parallelus (Raffray, 1877)⇒ Eodrepanus parallelus Raffray, 1877
Drepanocerus patrizii Boucomont, 1923
Drepanocerus striatus (Boucomont, 1921)⇒ Clypeodrepanus striatus Boucomont, 1921
Drepanocerus strigatus (Janssens, 1953)⇒ Clypeodrepanus strigatus Janssens, 1953
Drepanocerus sulcicollis (Castelnau, 1840)⇒ Sulcodrepanus sulcicollis Castelnau, 1840
Drepanocerus trogiformis Roth, 1851
Drepanoplatynus gilleti Boucomont, 1921
Eodrepanus bechynei Janssens, 1953
Eodrepanus fastiditus Péringuey, 1901
Eodrepanus paolae Barbero, Pal..., 2009
Eodrepanus parallelus Raffray, 1877
Epactoides andringitrae Paulian, 1975
Epactoides ankasokae Paulian, 1975
Epactoides basilewskyi Balthasar, 1960
Epactoides betschi Paulian, 1975
Epactoides costatus Paulian, 1992
Epactoides emkeni Montreuil, 2017
Epactoides femoralis Montreuil, 2005
Epactoides fiorii Paulian, 1976
Epactoides frontalis Montreuil, 2003
Epactoides giganteus Rossini, Vaz..., 2021
Epactoides hanskii Montreuil, 2005
Epactoides helenae Montreuil, 2005
Epactoides humberti Paulian, 1975
Epactoides humeralis Paulian, 1976
Epactoides hyphydroides Lebis, 1953
Epactoides incertus Lebis, 1953
Epactoides jounii Wirta & Mont..., 2008
Epactoides lacustris Paulian, 1976
Epactoides lissus Lebis, 1953
Epactoides madecassus Paulian, 1936
Epactoides mahaboi Paulian, 1976
Epactoides major Paulian, 1991
Epactoides mananarae Paulian, 1976
Epactoides mangabeensis Wirta & Mont..., 2008
Epactoides masoalae Paulian, 1976
Epactoides olsoufieffi Lebis, 1953
Epactoides paradoxus Paulian, 1976
Epactoides pauliani Lebis, 1953
Epactoides perinetanus Paulian, 1975
Epactoides perrieri Fairmaire, 1898
Epactoides rahagai Montreuil, 2005
Epactoides sambavae Paulian, 1976
Epactoides semiaeneus Lebis, 1953
Epactoides spinicollis Montreuil, 2003
Epactoides tiinae Montreuil, 2005
Epactoides vadoni Montreuil, 2005
Epactoides vagecarinatus Lebis, 1953
Epactoides viridicollis Montreuil, 2005
Epactoides vondrozoi Paulian, 1976
Epilissus alluaudi Lebis, 1953
Epilissus andranobeensis Montreuil, 2011
Epilissus antoetrae Paulian, 1975
Epilissus apotolamproides Lebis, 1961
Epilissus armipes Lebis, 1953
Epilissus bemarahaensis Montreuil & ..., 2022
Epilissus colasi Paulian, 1935
Epilissus cuprarius Fairmaire, 1889
Epilissus cuprarius discretus Montreuil, 2006
Epilissus cuprarius serindicus Paulian, 1976
Epilissus delphinensis Lebis, 1953
Epilissus discretus Montreuil, 2006
Epilissus emmae Lebis, 1953
Epilissus emmae emmae Lebis, 1953
Epilissus emmae obscuripennis Montreuil, 2006
Epilissus fantamattii Montreuil, 2011
Epilissus fuscovirens (Fairmaire, 1889)⇒ Arachnodes fuscovirens Fairmaire, 1889
Epilissus genieri Montreuil, 2006
Epilissus guillaumeti Paulian, 1975
Epilissus hova Künckel d’..., 1887
Epilissus mantasoae Paulian, 1976
Epilissus micheli Lebis, 1953
Epilissus morio (Harold, 1879)⇒ Arachnodes morio Harold, 1879
Epilissus morio hova Künckel d’He..., 1887
Epilissus morio morio Harold, 1879
Epilissus morio serindicus Paulian, 1976
Epilissus planatus Montreuil, 2006
Epilissus prasinus Klug, 1832
Epilissus robustus Montreuil & ..., 2022
Epilissus ruteri Lebis, 1953
Epilissus semicribrosus (Fairmaire, 1898)⇒ Nanos semicribrosus Fairmaire, 1898
Epilissus semilissus Montreuil & ..., 2022
Epilissus sikorai Paulian, 1977
Epilissus splendidus (Fairmaire, 1889)⇒ Arachnodes splendidus Lebis, 1953
Epilissus splendidus delphinensis Lebis, 1953
Epilissus splendidus descarpentriesi Lebis, 1953
Epilissus splendidus splendidus Fairmaire, 1889
Epilissus striatoides Paulian, 1976
Epilissus striatus Castelnau, 1840
Epilissus superbus Montreuil & ..., 2022
Epilissus viridis Klug, 1832
Epilissus vohemarensis Montreuil & ..., 2022
Epirinus flagellatus Fabricius, 1775
Euoniticellus africanus Harold, 1873
Euoniticellus fumigatus Boucomont, 1923
Euoniticellus inaequalis Reiche, 1850
Euoniticellus intermedius Reiche, 1848
Euoniticellus kawanus Janssens, 1939
Euoniticellus nasicornis Reiche, 1849
Euoniticellus pallens Olivier, 1789
Euoniticellus parvus Kraatz, 1895
Euoniticellus perniger Cambefort & ..., 1980
Euoniticellus tibatensis Kolbe, 1904
Euoniticellus triangulatus Harold, 1873
Euoniticellus zumpti Janssens, 1953
Euonthophagus carbonarius Klug, 1855
Euonthophagus consanguineus Müller, 1944
Euonthophagus consobrinus Péringuey, 1901
Euonthophagus jeanneli jeanneli
Euonthophagus maindroni d´Orbigny, 1897
Eusaproecius dudleyi Branco, 1992
Eusaproecius laticeps d´Orbigny, 1908
Eusaproecius magnigena d´Orbigny, 1913
Eusaproecius optatus Péringuey, 1901
Eusaproecius tinantae Boucomont, 1923
Garreta australugens Davis & Desc..., 2018
Garreta azareus
Garreta azureus Fabricius, 1801
Garreta caffer Fahraeus, 1857
Garreta crenulatus Kolbe, 1895
Garreta dejeani Castelnau, 1840
Garreta fastiditus Harold, 1867
Garreta laetus laetus
Garreta laetus olivaceus Quedenfeldt, 1884
Garreta lugens Fairmaire, 1891
Garreta malleolus Kolbe, 1895
Garreta matabelensis Janssens, 1938
Garreta nitens Olivier, 1789
Garreta nyassicus Kolbe, 1897
Garreta sumptuosus Castelnau, 1840
Garreta unicolor Fahraeus, 1857
Garreta wahlbergi Fahraeus, 1857
Gilletellus collarti Janssens, 1937
Gymnopleurus aenescens Wiedemann, 1821
Gymnopleurus bicolor (Latreille, 1823)⇒ Gymnopleurus leei Fabricius, 1792
Gymnopleurus coerulescens Olivier, 1789
Gymnopleurus foricarius Garreta, 1914
Gymnopleurus fulgidus Olivier, 1789
Gymnopleurus humanus MacLeay, 1821
Gymnopleurus humeralis Klug, 1855
Gymnopleurus ignitus Klug, 1855
Gymnopleurus jacksoni Waterhouse, 1890
Gymnopleurus laevicollis (Castelnau, 1840)⇒ Gymnopleurus purpureus Garreta, 1914
Gymnopleurus leei Fabricius, 1792
Gymnopleurus moerens Kolbe, 1895
Gymnopleurus plicatulus Fairmaire, 1890
Gymnopleurus pumilus (Reiche, 1847)⇒ Gymnopleurus virens Erichson, 1843
Gymnopleurus purpureus Garreta, 1914
Gymnopleurus pustulatus Kolbe, 1895
Gymnopleurus reichei Waterhouse, 1890
Gymnopleurus rhodesianus Balthasar, 1963
Gymnopleurus ruandensis Janssens, 1938
Gymnopleurus sericeifrons Fairmaire, 1887
Gymnopleurus somaliensis Lansberge, 1882
Gymnopleurus thelwalli Waterhouse, 1890
Gymnopleurus thoracicus Harold, 1868
Gymnopleurus virens Erichson, 1843
Haroldius modestus Janssens, 1953
Haroldius stefanii Montreuil, 2010
Helictopleurus aeneoniger Lebis, 1960
Helictopleurus ambiguus Paulian & Ca..., 1991
Helictopleurus aurocupreus Fairmaire, 1901
Helictopleurus basicornis Fairmaire, 1904
Helictopleurus bivittatus d´Orbigny, 1915
Helictopleurus cambeforti Montreuil, 2005
Helictopleurus carbonarius Lebis, 1960
Helictopleurus clouei Harold, 1869
Helictopleurus coruscus d´Orbigny, 1915
Helictopleurus cribricollis Lebis, 1960
Helictopleurus dorbignyi Montreuil, 2005
Helictopleurus fasciolatus Fairmaire, 1898
Helictopleurus fasciolatus fasciolatus
Helictopleurus fasciolatus obscurus Lebis, 1960
Helictopleurus fasciolatus pseudofasciolatus Montreuil, 2007
Helictopleurus fasciolatus suturalis Montreuil, 2012
Helictopleurus fissicollis Fairmaire, 1895
Helictopleurus fissicollis australis Montreuil, 2007
Helictopleurus fissicollis fissicollis
Helictopleurus fissicollis morettoi Montreuil, 2005
Helictopleurus fulgens Lebis, 1960
Helictopleurus fulgens fulgens
Helictopleurus fulgens purpuricollis Lebis, 1960
Helictopleurus fungicola Fairmaire, 1899
Helictopleurus fungicola peyrierasi Paulian & Ca..., 1991
Helictopleurus furcicornis Lebis, 1960
Helictopleurus gibbicollis Fairmaire, 1895
Helictopleurus giganteus Harold, 1869
Helictopleurus halffteri Balthasar, 1964
Helictopleurus hanskii Montreuil, 2005
Helictopleurus heidiae Montreuil, 2007
Helictopleurus hypocrita Balthasar, 1941
Helictopleurus infimus Fairmaire, 1901
Helictopleurus littoralis Montreuil, 2005
Helictopleurus marsyas Olivier, 1789
Helictopleurus minutus Montreuil, 2007
Helictopleurus neoamplicollis Krell, 2000
Helictopleurus neuter Fairmaire, 1898
Helictopleurus nicollei Lebis, 1960
Helictopleurus niger d´Orbigny, 1915
Helictopleurus nigricans d´Orbigny, 1915
Helictopleurus nigritulus Lebis, 1960
Helictopleurus nitidicollis Lebis, 1960
Helictopleurus patricii Paulian, 1937
Helictopleurus perpunctatus Balthasar, 1963
Helictopleurus perrieri Fairmaire, 1898
Helictopleurus peyrierasi (Paulian & Ca..., 1991)⇒ Helictopleurus fungicola peyrierasi Paulian & Ca..., 1991
Helictopleurus pluristriatus d´Orbigny, 1915
Helictopleurus politicollis Fairmaire, 1902
Helictopleurus praerugosus Fairmaire, 1898
Helictopleurus pseudoniger Montreuil, 2005
Helictopleurus quadripunctatus Olivier, 1789
Helictopleurus rubricollis Lebis, 1960
Helictopleurus rudicollis Fairmaire, 1898
Helictopleurus semimetallicus Fairmaire, 1888
Helictopleurus seminiger d´Orbigny, 1915
Helictopleurus semivirens d´Orbigny, 1915
Helictopleurus sericeus Lebis, 1960
Helictopleurus sicardi Paulian, 1975
Helictopleurus sinuaticornis Fairmaire, 1898
Helictopleurus sinuatocornis Fairmaire, 1898
Helictopleurus splendidicollis Fairmaire, 1893
Helictopleurus splendidicollis similis Montreuil, 2007
Helictopleurus splendidicollis splendidicollis
Helictopleurus splendidus d´Orbigny, 1915
Helictopleurus steineri Paulian & Ca..., 1991
Helictopleurus subretusus Fairmaire, 1895
Helictopleurus sumptuosus Harold, 1873
Helictopleurus tamatavensis Montreuil, 2005
Helictopleurus tristis (d´Orbigny, 1915)⇒ Helictopleurus quadripunctatus Olivier, 1789
Helictopleurus undatus Olivier, 1789
Helictopleurus unifasciatus Fairmaire, 1901
Helictopleurus vadoni Lebis, 1960
Helictopleurus viettei Paulian & Ca..., 1984
Helictopleurus villiersi Paulian & Ca..., 1984
Helictopleurus viridans Fairmaire, 1901
Helictopleurus viridans pseudoviridans Montreuil, 2007
Helictopleurus viridans viridans
Helictopleurus viridiflavus Fairmaire, 1898
Heliocopris alatus Felsche, 1910
Heliocopris anadematus Gillet, 1908
Heliocopris andersoni Bates, 1868
Heliocopris antenor (Olivier, 1789)⇒ Heliocopris colossus Bates, 1868
Heliocopris atropos Boheman, 1860
Heliocopris beccarii Harold, 1871
Heliocopris biimpressus Kolbe, 1893
Heliocopris colossus Bates, 1868
Heliocopris corniculatus Janssens, 1939
Heliocopris cuneifer Lesne, 1906
Heliocopris densissa Roth, 1851
Heliocopris dilloni Guérin-Ménev..., 1847
Heliocopris dolosus Janssens, 1939
Heliocopris erycoides Felsche, 1907
Heliocopris eryx ikelenge Moretto, 2017
Heliocopris faunus Boheman, 1857
Heliocopris felschei Kolbe, 1904
Heliocopris fischeri Schäfer, 2019
Heliocopris furcithorax Müller, 1941
Heliocopris gigas Linnaeus, 1758Ägyptische Riese
Heliocopris gigas erythraeus Müller, 1941
Heliocopris gigas gigas
Heliocopris hamadryas Fabricius, 1775
Heliocopris hamifer Harold, 1878
Heliocopris haroldi Kolbe, 1893
Heliocopris hermes Gillet, 1911
Heliocopris hunteri Waterhouse, 1891
Heliocopris japetus Klug, 1855
Heliocopris kolbei Felsche, 1901
Heliocopris letiranti Moretto, 2021
Heliocopris marshalli Péringuey, 1901
Heliocopris minos Gillet, 1907
Heliocopris myrmidon Kolbe, 1893
Heliocopris neptuniformis Felsche, 1907
Heliocopris neptunoides Janssens, 1939
Heliocopris neptunus Boheman, 1857
Heliocopris pirmal Fabricius, 1798
Heliocopris quinqueangulatus Janssens, 1939
Heliocopris samson Harold, 1878
Heliocopris sirius (Gillet, 1925)⇒ Heliocopris hamifer Harold, 1878
Heliocopris smithi Moretto, 2017
Heliocopris solitarius Kolbe, 1893
Heliocopris staudingeri Kolbe, 1893
Heliocopris stroehlei Moretto, 2014
Heliocopris sylvanus Gillet, 1925
Heliocopris werneri Moretto & Mi..., 2013
Heteroclitopus collarti Janssens, 1939
Heteroclitopus foveatus Boucomont, 1923
Heteroclitopus foveolatus Branco & Josso, 2007
Heteroclitopus herteli Ferreira, 1969
Heteroclitopus remipes Péringuey, 1901
Heteronitis castelnaui Harold, 1862
Heteronitis ragazzii Fairmaire, 1893
Heteronitis tricornutus Fairmaire, 1891
Heterosyphus sicardi Paulian, 1975
Hyalonthophagus alcedo Orbigny, 1913
Hyalonthophagus alcyon Klug, 1855
Hyalonthophagus alcyonides Orbigny, 1913
Hyalonthophagus hyalcyon Palestrini &..., 1989
Hyalonthophagus mixtifrons Orbigny, 1910
Hyalonthophagus novalcedo Josso, 2021
Hyalonthophagus pseudovirens Orbigny, 1913
Hyalonthophagus virens d´Orbigny, 1913
Hyalonthophagus viridiceps Orbigny, 1910
Ixodina abyssinica Roth, 1851
Ixodina abyssinica abyssinica
Ixodina abyssinica tangana Endrödi, 1971
Ixodina abyssinica tanganus Endrodi, 1971
Ixodina freyi Janssens, 1953
Ixodina kovacsi Endrödi, 1976
Ixodina saegeri Balthasar, 1963
Ixodina szunyoghyi Endrödi, 1971
Ixodina szunyoghyi szunyoghyi
Janssensantus alatus Josso, 2022
Janssensantus inexpectatus Paulian, 1976
Janssensantus pauliani Scholtz & Ho..., 1987
Kheper aegyptiorum Latreille, 1823
Kheper aegyptiorum aegyptiorum
Kheper aegyptiorum purpurascens Gerstaecker, 1871
Kheper aeratus Gerstaecker, 1871
Kheper asceticus Gillet, 1907
Kheper clericus Boheman, 1857
Kheper cupreus Castelnau, 1840
Kheper cuvieri MacLeay, 1821
Kheper festivus Harold, 1868
Kheper festivus wellsi Moretto, 1998
Kheper laevistriatus Fairmaire, 1893
Kheper lamarcki MacLeay, 1821
Kheper nigroaeneus Boheman, 1857
Kheper platynotus Bates, 1888
Kheper porosus Bates, 1888
Kheper prodigiosus Erichson, 1843
Kheper pustulosus Gerstaecker, 1871
Kheper rolciki Pokorný & Zí..., 2015
Kheper subaeneus Harold, 1869
Kheper vanderkelleni Lansberge, 1886
Kheper venerabilis Harold, 1871
Krikkenius smeenki Krikken, 1982
Latodrepanus caelatus Gerstaecker, 1871
Latodrepanus laticollis Fahraeus, 1857
Latodrepanus nicolasi Barbero, Pal..., 2009
Latodrepanus pulvinarius Balthasar, 1963
Latodrepanus schimperi Janssens, 1953
Liatongus amitinus Boucomont, 1921
Liatongus arrowi Boucomont, 1921
Liatongus aterrimus Boucomont, 1921
Liatongus ferreirae Balthasar, 1964
Liatongus fulvostriatus d´Orbigny, 1916
Liatongus militaris Castelnau, 1840
Liatongus raffrayi Lansberge, 1886
Liatongus schoutedeni Boucomont, 1920
Liatongus sjostedti Felsche, 1904
Liatongus spathulatus Roth, 1851
Liatongus taurus Boucomont, 1920
Liatongus testudo Lansberge, 1886
Liatongus tuberculicollis Felsche, 1909
Liatongus urus Kolbe, 1914
Litocopris muticus Boheman, 1857
Litocopris punctiventris Waterhouse, 1891
Litocopris ruandanus Balthasar, 1961
Lophodonitis carinatus Felsche, 1907
Lophodonitis minettii Moretto, 2011
Madateuchus viettei (Paulian, 1953)⇒ Scarabaeus viettei Paulian, 1953
Megalonitis bohemani Lansberge, 1875
Megalonitis bohemanni Lansberge, 1875
Megalonitis gigas Bertoloni, 1855
Megaponerophilus megaponerae Brauns, 1914
Metacatharsius abortivus Fairmaire, 1891
Metacatharsius anderseni Waterhouse, 1891
Metacatharsius anomalus Felsche, 1910
Metacatharsius auberti Fairmaire, 1891
Metacatharsius benadirensis Müller, 1947
Metacatharsius brevicosta Müller, 1947
Metacatharsius clypeolatus Balthasar, 1940
Metacatharsius convexiusculus Shipp, 1897
Metacatharsius dentinum Ferreira, 1964
Metacatharsius duplicatus Müller, 1941
Metacatharsius exiguus Boheman, 1860
Metacatharsius ferreirae Balthasar, 1965
Metacatharsius gardoensis Balthasar, 1941
Metacatharsius holyi Balthasar, 1965
Metacatharsius imitator Balthasar, 1965
Metacatharsius inermis Castelnau, 1840
Metacatharsius janssensi Frey, 1963
Metacatharsius latifrons Harold, 1868
Metacatharsius lomii Müller, 1941
Metacatharsius minutus Lansberge, 1882
Metacatharsius modestus Balthasar, 1965
Metacatharsius nubiensis Harold, 1878
Metacatharsius omoensis Müller, 1941
Metacatharsius opacus Waterhouse, 1891
Metacatharsius ovulum Balthasar, 1940
Metacatharsius patrizii Müller, 1947
Metacatharsius peleus Olivier, 1789
Metacatharsius pernitidus Müller, 1941
Metacatharsius politus Paulian, 1934
Metacatharsius pollicatus Harold, 1878
Metacatharsius pseudoopacus Ferreira, 1965
Metacatharsius pumilionis Wallengren, 1881
Metacatharsius pusio Kolbe, 1908
Metacatharsius rochai Ferreira, 1964
Metacatharsius rosinae Balthasar, 1970
Metacatharsius rusingae ♰ R. Paulian, 1976
Metacatharsius seminulum Balthasar, 1940
Metacatharsius simulator Balthasar, 1939
Metacatharsius tenellus Müller, 1947
Metacatharsius troglodytes (Boheman, 1857)⇒ Metacatharsius anderseni Waterhouse, 1891
Metacatharsius tuberifrons Fairmaire, 1884
Metacatharsius useramus Kolbe, 1893
Metacatharsius zuluanus Balthasar, 1965
Milichus apicalis (Fahraeus, 1857)⇒ Milichus rhodesianus Péringuey, 1904
Milichus boucomonti Cambefort, 1996
Milichus dudleyae Cambefort, 1996
Milichus dudleyi Cambefort, 1996
Milichus dudleyorum Cambefort, 1996
Milichus inaequalis Boucomont, 1928
Milichus inaequalis inaequalis
Milichus jugatus d´Orbigny, 1907
Milichus lecourti Nicolas & Ca..., 1994
Milichus picticollis Gerstaecker, 1871
Milichus rhodesianus Péringuey, 1904
Mimonthophagus ambiguus Péringuey, 1901
Mimonthophagus anomalus Klug, 1855
Mimonthophagus hinnulus Klug, 1832
Mimonthophagus limbibasis d´Orbigny, 1905
Nanos agaboides Boucomont, 1937
Nanos andreiae Montreuil & ..., 2014
Nanos ankaranae Paulian, 1976
Nanos antsalovaensis Montreuil & ..., 2011
Nanos antsihanakensis Lebis, 1953
Nanos ater Castelnau, 1840
Nanos bemarahaensis Montreuil & ..., 2014
Nanos bicoloratus Lebis, 1953
Nanos bimaculatus Kunckel, 1887
Nanos binotatus Lebis, 1953
Nanos clypeatus Castelnau, 1840
Nanos constricticollis Montreuil & ..., 2014
Nanos dubitatus Lebis, 1953
Nanos fusconitens Fairmaire, 1899
Nanos hanskii Montreuil & ..., 2007
Nanos humbloti Lebis, 1953
Nanos humeralis Paulian, 1975
Nanos incertus Paulian, 1976
Nanos magnus Montreuil & ..., 2014
Nanos manomboensis Montreuil & ..., 2007
Nanos manongorivoensis Montreuil & ..., 2014
Nanos marojejyensis Montreuil & ..., 2014
Nanos minutus Castelnau, 1840
Nanos mirjae Montreuil & ..., 2014
Nanos mixtus Montreuil & ..., 2014
Nanos nitens Lebis, 1953
Nanos nitens nitens
Nanos nitens pseudonitens Lebis, 1953
Nanos occidentalis Paulian, 1976
Nanos peyrierasi (Paulian, 1976)⇒ Apotolamprus peyrierasi Paulian, 1985
Nanos pseudofusconitens Montreuil & ..., 2014
Nanos pseudominutus Montreuil & ..., 2014
Nanos pseudorubromaculatus Montreuil & ..., 2014
Nanos pseudoviettei Montreuil & ..., 2014
Nanos punctatus Boucomont, 1937
Nanos pygmaeus Castelnau, 1840
Nanos ranomafanaensis Montreuil & ..., 2014
Nanos rubromaculatus Kunckel d´He..., 1887
Nanos rubrosignatus Lebis, 1953
Nanos semicribrosus Fairmaire, 1898
Nanos sinuatipes Boucomont, 1937
Nanos vadoni Lebis, 1953
Nanos viettei Paulian, 1976
Nanos viridissimus Montreuil & ..., 2011
Neonitis nigritiae Gillet, 1918
Neonitis rhodesiae Gillet, 1918
Neosaproecius nyassalandicus Branco, 1990
Neosisyphus astriatus Schafer & Fi..., 2001
Neosisyphus basilewskyi Haaf, 1956
Neosisyphus calcaratus Klug, 1855
Neosisyphus confrater Kolbe, 1914
Neosisyphus fortuitus Péringuey, 1901
Neosisyphus infuscatus Klug, 1855
Neosisyphus mirabilis Arrow, 1927
Neosisyphus penicillatus Harold, 1880
Neosisyphus quadricollis Gory, 1833
Neosisyphus rubripes Péringuey, 1901
Neosisyphus rubrus Paschalidis, 1974
Neosisyphus rugosus Gory, 1833
Neosisyphus setiger Roth, 1851
Neosisyphus spinipes Thunberg, 1818
Neosisyphus tibialis Raffray, 1877
Neosisyphus trochantericus Fairmaire, 1891
Nesosisyphus pygmaeus Vinson, 1946
Nesosisyphus regnardi Alluaud, 1898
Nesosisyphus rotundatus Vinson, 1946
Nesosisyphus vicinus Vinson, 1939
Nesovinsonia vinsoni Paulian, 1939
Nitiocellus collarti Janssens, 1939
Odontoloma metasternale d´Orbigny, 1902
Odontoloma multipunctatum Howden & Sch..., 1987
Odontoloma pauxillum Boheman, 1857
Odontoloma quadridens d´Orbigny, 1908
Oniticellus egregius Klug, 1855
Oniticellus formosus (Chevrolat, 1830)⇒ Oniticellus pictus Hausmann, 1807
Oniticellus inaequalis (Reiche, 1847)⇒ Euoniticellus inaequalis Reiche, 1850
Oniticellus intermedius (Reiche, 1848)⇒ Euoniticellus intermedius Reiche, 1848
Oniticellus militaris (Castelnau, 1840)⇒ Liatongus militaris Castelnau, 1840
Oniticellus pictus Hausmann, 1807
Oniticellus planatus Castelnau, 1840
Oniticellus pseudoplanatus Balthasar, 1964
Oniticellus spinipes (Roth, 1851)⇒ Tiniocellus spinipes Roth, 1851
Onitis abyssinicus (Reiche, 1847)⇒ Onitis lycophron Klug, 1855
Onitis adelphus Kolbe, 1914
Onitis adriani Ferreira, 1976
Onitis aeneomicans Gillet, 1918
Onitis aerarius Harold, 1878
Onitis aeruginosus Klug, 1855
Onitis aethiops Lesne, 1906
Onitis affinis Felsche, 1911
Onitis albertcollarti Génier & Mor..., 2019
Onitis alexis (Klug, 1835)⇒ Cheironitis ungaricus ungaricus Herbst, 1789
Onitis alexis alexis
Onitis anthracinus Felsche, 1907
Onitis archettii Janssens, 1943
Onitis artuosus Gillet, 1909
Onitis autumnalis Davis, 1986
Onitis bispicticollis Krikken, 1977
Onitis bredoi Janssens, 1949
Onitis brucemillerorum Josso & Mont..., 2017
Onitis caffer Boheman, 1857
Onitis camerunus (Gillet, 1933)⇒ Onitis cryptodus Gillet, 1918
Onitis camiadei Josso, 2012
Onitis cerrutii Müller, 1949
Onitis chalceus Lansberge, 1875
Onitis confusus Boheman, 1860
Onitis coxalis Gillet, 1918
Onitis crenatus Reiche, 1847
Onitis cryptodus Gillet, 1918
Onitis cupreus Castelnau, 1840
Onitis curvipes Lansberge, 1875
Onitis deceptor Péringuey, 1901
Onitis denticoxa Müller, 1941
Onitis dimidiatus Felsche, 1907
Onitis dispar Péringuey, 1896
Onitis excavaticollis Balthasar, 1942
Onitis fabricii Roth, 1851
Onitis femoralis Josso, 2012
Onitis fractipes Boucomont, 1923
Onitis fulgidus (Klug, 1855)⇒ Pleuronitis fulgidus Castelnau, 1840
Onitis fulmineus Janssens, 1943
Onitis gazanus Arrow, 1909
Onitis granicollis Felsche, 1907
Onitis inflaticollis Felsche, 1907
Onitis insuetus (Kolbe, 1914)⇒ Onitis sulcipennis Felsche, 1907
Onitis inversidens Lansberge, 1875
Onitis janssenii Gomes Alves, 1954
Onitis kasanka Josso, 2012
Onitis keniensis Gillet, 1933
Onitis kingstoni Krikken, 1977
Onitis laminosus Ferreira, 1976
Onitis lamnifer Gillet, 1909
Onitis lobipes Felsche, 1907
Onitis lycophron Klug, 1855
Onitis malleatus Janssens, 1937
Onitis marshalli Janssens, 1939
Onitis mendax Gillet, 1918
Onitis menieri Cambefort, 1981
Onitis meruensis Ferreira, 1977
Onitis meyeri Kolbe, 1891
Onitis minutus Lansberge, 1875
Onitis monstrosus Felsche, 1910
Onitis montreuili Josso, 2012
Onitis mossambicensis Ferreira, 1976
Onitis mraci Moretto, 2021
Onitis nemoralis Gillet, 1932
Onitis nubiensis Lansberge, 1875
Onitis obenbergeri Balthasar, 1942
Onitis orthopus Lansberge, 1875
Onitis paradoxus (Boheman, 1857)⇒ Tropidonitis paradoxus Boheman, 1857
Onitis paramniszechi Ferreira, 1976
Onitis parvulus Balthasar, 1963
Onitis pecuarius Lansberge, 1875
Onitis perbrincki Ferreira, 1958
Onitis perturbator Péringuey, 1901
Onitis picticollis Boheman, 1857
Onitis pinheyi Ferreira, 1977
Onitis politus Lansberge, 1875
Onitis popei Ferreira, 1977
Onitis proximus Ferreira, 1977
Onitis pseudojansenii Krikken, 1977
Onitis pseudoorthopus Ferreira, 1976
Onitis pseudosetosus Ferreira, 1976
Onitis pumilio Balthasar, 1963
Onitis reichei Lansberge, 1875
Onitis retrodentatus Gillet, 1932
Onitis robustus Boheman, 1857
Onitis rothi Lansberge, 1875
Onitis setosus Lansberge, 1875
Onitis shoensis (Reiche, 1847)⇒ Onitis vicinus Lansberge, 1875
Onitis similis Felsche, 1911
Onitis sphinx Fabricius, 1775
Onitis spinicrus Fairmaire, 1893
Onitis subcrenatus Kolbe, 1897
Onitis sulcipennis Felsche, 1907
Onitis sulcipennis naromorus Krikken, 1977
Onitis sulcipennis sulcipennis
Onitis tanzaniensis Ferreira, 1977
Onitis thalassinus Lansberge, 1875
Onitis thoracicus Ferreira, 1976
Onitis tortuosus Houston, 1983
Onitis trochantericus Ferreira, 1976
Onitis tumidus Gillet, 1918
Onitis uncinatus Klug, 1855
Onitis vanderkelleni Lansberge, 1886
Onitis vicinus Lansberge, 1875
Onitis viridulus (Boheman, 1857)⇒ Onitis pecuarius Lansberge, 1875
Onitis westermanni Lansberge, 1886
Onitis zambiensis Josso, 2012
Onthophagus (Afrostrandius) bimetallicus d´Orbigny, 1907
Onthophagus (Afrostrandius) plebejus Klug, 1855
Onthophagus (Afrostrandius) pseudoplebejus Moretto, 2008
Onthophagus tigrinus kageraensis Moretto, 2009
Onthophagus (Digitonthophagus) gazella (Fabricius, 1787)⇒ Digitonthophagus gazella Fabricius, 1787
Onthophagus (Euonthophagus) carbonarius Klug, 1855
Onthophagus (Euonthophagus) consobrinus Péringuey, 1901
Onthophagus (Hyalonthophagus) hyalcyon Palestrini &..., 1989
Onthophagus (Onthophagus) alternans (Raffray, 1877)⇒ Onthophagus taurus Schreber, 1759
Onthophagus (Onthophagus) depressus Harold, 1871
Onthophagus (Onthophagus) maculatus Fabricius, 1801
Onthophagus (Onthophagus) margaritifer d´Orbigny, 1898
Onthophagus (Onthophagus) seabrai Gomes Alves, 1944
Onthophagus (Onthophagus) subalternans d´Orbigny, 1902
Onthophagus (Onthophagus) taurus Schreber, 1759Oechslein
Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790)⇒ Onthophagus similis Scriba, 1790
Onthophagus (Palaeonthophagus) lemur Fabricius, 1781
Onthophagus marginalis przewalski Kabakov, 2006
Onthophagus (Palaeonthophagus) petrovitzi (Frey, 1975)⇒ Onthophagus petrovitzianus Pittino, 2006
Onthophagus (Palaeonthophagus) petrovitzianus Pittino, 2006
Onthophagus (Palaeonthophagus) ponticus Harold, 1883
Onthophagus (Palaeonthophagus) semicornis Panzer, 1798Halbhörniger Kotkäfer
Onthophagus (Paleonthophagus) similis Scriba, 1790
Onthophagus (Proagoderus) alcedo d´Orbigny, 1913
Onthophagus (Proagoderus) alcyon (Klug, 1855)⇒ Onthophagus alcedo d´Orbigny, 1913
Onthophagus (Proagoderus) alcyonides d´Orbigny, 1913
Onthophagus (Proagoderus) aureiceps d´Orbigny, 1902
Onthophagus (Proagoderus) fossidorsis d´Orbigny, 1905
Onthophagus (Proagoderus) mixtifrons d´Orbigny, 1910
Onthophagus (Proagoderus) pegesimallus Marcus, 1917
Onthophagus (Proagoderus) pseudovirens d´Orbigny, 1913
Onthophagus (Proagoderus) viridiceps d´Orbigny, 1910
Onthophagus obliquus shimba (Cambefort, 1980)⇒ Onthophagus shimba Cambefort, 1980
Onthophagus abas Balthasar, 1946
Onthophagus abeillei d´Orbigny, 1897
Onthophagus abruptus d´Orbigny, 1913
Onthophagus abyssinicus Gillet, 1925
Onthophagus acutus d´Orbigny, 1908
Onthophagus aeneoniger d´Orbigny, 1913
Onthophagus aeneopiceus d´Orbigny, 1902
Onthophagus aequatus Péringuey, 1901
Onthophagus aequepubens d´Orbigny, 1905
Onthophagus aereidorsis d´Orbigny, 1902
Onthophagus aeremicans d´Orbigny, 1904
Onthophagus aeruginosus Roth, 1851
Onthophagus aesopus Lansberge, 1882
Onthophagus aethiopicus d´Orbigny, 1902
Onthophagus aguatus
Onthophagus akamineae Moretto & Gé..., 2020
Onthophagus alainmonforti Josso, 2022
Onthophagus albicomus d´Orbigny, 1908
Onthophagus albipodex d´Orbigny, 1902
Onthophagus aliceae Josso, 2019
Onthophagus ambiquus
Onthophagus amoenus d´Orbigny, 1908
Onthophagus amplipennis d´Orbigny, 1905
Onthophagus angulicornis (Mulsant, 1842)⇒ Onthophagus semicornis Panzer, 1798
Onthophagus antecornutus Josso, 2019
Onthophagus apiciosus d´Orbigny, 1902
Onthophagus apicosus
Onthophagus arcifer d´Orbigny, 1904
Onthophagus areolatus d´Orbigny, 1905
Onthophagus asperatus d´Orbigny, 1905
Onthophagus aspernatus d´Orbigny, 1907
Onthophagus asperrimus d´Orbigny, 1902
Onthophagus astigma d´Orbigny, 1904
Onthophagus athiensis d´Orbigny, 1913
Onthophagus atricolor d´Orbigny, 1907
Onthophagus atridorsis d´Orbigny, 1902
Onthophagus atrofasciatus d´Orbigny, 1905
Onthophagus atrosplendens Frey, 1961
Onthophagus auriculatus (Klug, 1855)⇒ Onthophagus latestriatus d´Orbigny, 1908
Onthophagus axillaris (Boheman, 1860)⇒ Onthophagus criniger d´Orbigny, 1904
Onthophagus babaulti d´Orbigny, 1915
Onthophagus beiranus Péringuey, 1908
Onthophagus bellus d´Orbigny, 1905
Onthophagus bergeri Frey, 1975
Onthophagus bicallifrons d´Orbigny, 1902
Onthophagus bicallosus Klug, 1855
Onthophagus bicavifrons d´Orbigny, 1902
Onthophagus bicolor (Raffray, 1877)⇒ Onthophagus flavolimbatus Klug, 1855
Onthophagus bicristiger d´Orbigny, 1913
Onthophagus bidens Olivier, 1789
Onthophagus bifidicornis d´Orbigny, 1902
Onthophagus bifidus Reiche, 1849
Onthophagus bifrons d´Orbigny, 1905
Onthophagus bimarginatus d´Orbigny, 1902
Onthophagus binodis Thunberg, 1818
Onthophagus binodosus d´Orbigny, 1908
Onthophagus binodulus d´Orbigny, 1913
Onthophagus biplagiatus Thomson, 1858
Onthophagus birugatus d´Orbigny, 1902
Onthophagus bisignatus d´Orbigny, 1913
Onthophagus bituber d´Orbigny, 1904
Onthophagus bituberans d´Orbigny, 1905
Onthophagus bituberculatus (Olivier, 1789)⇒ Onthophagus acutus d´Orbigny, 1908
Onthophagus bituberifrons Josso, 2018
Onthophagus blanchardi Harold, 1869
Onthophagus boops d´Orbigny, 1905
Onthophagus bovimus
Onthophagus bovinus Péringuey, 1892
Onthophagus breviceps d´Orbigny, 1902
Onthophagus brevisetis d´Orbigny, 1915
Onthophagus brunellii Müller, 1942
Onthophagus bufonidius Josso, 2019
Onthophagus calliger d´Orbigny, 1913
Onthophagus cancer Lansberge, 1886
Onthophagus carcharias Harold, 1875
Onthophagus carinicollis Raffray, 1877
Onthophagus carinidorsis d´Orbigny, 1908
Onthophagus carinifer d´Orbigny, 1904
Onthophagus carpophilus Pereira & Ha..., 1961
Onthophagus catta Fabricius, 1787
Onthophagus cavifrons Harold, 1886
Onthophagus cervus (Fabricius, 1798)⇒ Onthophagus falsus Gillet, 1925
Onthophagus chalcostolus d´Orbigny, 1902
Onthophagus chirindaensis Josso, 2022
Onthophagus chirindanus d´Orbigny, 1913
Onthophagus chitipaensis Josso, 2019
Onthophagus chlorophanus d´Orbigny, 1902
Onthophagus choanicus d´Orbigny, 1904
Onthophagus cicer Balthasar, 1964
Onthophagus cincticollis d´Orbigny, 1902
Onthophagus cinctipennis Quedenfeldt, 1884
Onthophagus cineraceus d´Orbigny, 1902
Onthophagus clavisetis d´Orbigny, 1905
Onthophagus clitellarius d´Orbigny, 1908
Onthophagus clusifrons d´Orbigny, 1905
Onthophagus comatulus d´Orbigny, 1905
Onthophagus concavifrons d´Orbigny, 1904
Onthophagus confertus Péringuey, 1908
Onthophagus confusus Boucomont, 1932
Onthophagus conradsi Balthasar, 1937
Onthophagus contiguicornis d´Orbigny, 1913
Onthophagus convexipennis Dierkens, 2022
Onthophagus convexus d´Orbigny, 1908
Onthophagus corniculatus (Reiche, 1847)⇒ Onthophagus senegalensis d´Orbigny, 1902
Onthophagus corniculiger d´Orbigny, 1913
Onthophagus cornutus Ferreira, 1971
Onthophagus coronatus d´Orbigny, 1902
Onthophagus costilatus d´Orbigny, 1908
Onthophagus costulicollis Frey, 1975
Onthophagus crassicollis Boucomont, 1913
Onthophagus creber d´Orbigny, 1905
Onthophagus cribellum d´Orbigny, 1902
Onthophagus cribripennis d´Orbigny, 1902
Onthophagus criniger d´Orbigny, 1904
Onthophagus cristatus d´Orbigny, 1905
Onthophagus croesulus Bates, 1888
Onthophagus crucenotatus d´Orbigny, 1905
Onthophagus cruentatus Klug, 1855
Onthophagus cupreopinguis Josso, 2022
Onthophagus curvilamina Müller, 1942
Onthophagus cyaneiceps d´Orbigny, 1913
Onthophagus cyaneoniger d´Orbigny, 1902
Onthophagus davisi Frey, 1975
Onthophagus debilis d´Orbigny, 1905
Onthophagus decedens Péringuey, 1904
Onthophagus decedeus
Onthophagus declivicollis d´Orbigny, 1902
Onthophagus decolor d´Orbigny, 1902
Onthophagus dedecor Wallengren, 1881
Onthophagus deflexus d´Orbigny, 1908
Onthophagus delahayei Josso, 2011
Onthophagus delicatulus Raffray, 1877
Onthophagus delphinensis d´Orbigny, 1913
Onthophagus densatus Frey, 1963
Onthophagus densepunctatus Frey, 1963
Onthophagus denticulatus d´Orbigny, 1902
Onthophagus depilatus d´Orbigny, 1913
Onthophagus depilis d´Orbigny, 1902
Onthophagus deterrens Péringuey, 1901
Onthophagus dicax Balthasar, 1966
Onthophagus dicella Bates, 1888
Onthophagus dinoderus d´Orbigny, 1913
Onthophagus discovirens d´Orbigny, 1902
Onthophagus discretus Péringuey, 1901
Onthophagus dispar Péringuey, 1901
Onthophagus distichus Roth, 1851
Onthophagus ditus (Péringuey, 1901)⇒ Onthophagus fungicola d´Orbigny, 1908
Onthophagus diversus Reiche, 1847
Onthophagus dlabolai Balthasar, 1964
Onthophagus dohertyi d´Orbigny, 1902
Onthophagus dudleyianus Josso, 2019
Onthophagus ebenicolor d´Orbigny, 1902
Onthophagus ebenus Péringuey, 1888
Onthophagus echinus Boucomont, 1914
Onthophagus elegans Klug, 1832
Onthophagus emeritus Péringuey, 1901
Onthophagus erectinasus d´Orbigny, 1902
Onthophagus estelleae Josso, 2022
Onthophagus evae Balthasar, 1944
Onthophagus exasperatus (Gerstaecker, 1871)⇒ Onthophagus latissimus d´Orbigny, 1902
Onthophagus excisiceps d´Orbigny, 1902
Onthophagus exiguus (Raffray, 1877)⇒ Onthophagus rubens d´Orbigny, 1902
Onthophagus exilis d´Orbigny, 1913
Onthophagus extensicollis d´Orbigny, 1907
Onthophagus extensus Harold, 1878
Onthophagus extraneus Josso, 2021
Onthophagus falcarius Balthasar, 1966
Onthophagus fallaciosus Raffray, 1877
Onthophagus fallax d´Orbigny, 1913
Onthophagus fasciolatus Boucomont, 1924
Onthophagus filicornis Harold, 1873
Onthophagus fimetarius Roth, 1851
Onthophagus flavibasis d´Orbigny, 1904
Onthophagus flavipennis d´Orbigny, 1905
Onthophagus flavipleurus Dierkens, 2022
Onthophagus flavocinctus Frey, 1975
Onthophagus flavolimbatus Klug, 1855
Onthophagus flexifrons d´Orbigny, 1908
Onthophagus foraminosus d´Orbigny, 1902
Onthophagus fossibasis d´Orbigny, 1908
Onthophagus fossulatus d´Orbigny, 1908
Onthophagus francoisi Josso, 2013
Onthophagus frontalis Raffray, 1877
Onthophagus fulvocinctus d´Orbigny, 1902
Onthophagus fumatus d´Orbigny, 1913
Onthophagus fungicola d´Orbigny, 1908
Onthophagus furcatoides (Lansberge, 1886)⇒ Onthophagus neofurcatus Goidanich, 1926
Onthophagus furculifer d´Orbigny, 1905
Onthophagus fuscatus d´Orbigny, 1908
Onthophagus fuscidorsis (d´Orbigny, 1902)⇒ Onthophagus fuscatus d´Orbigny, 1908
Onthophagus fuscivestis d´Orbigny, 1902
Onthophagus galeatus Boucomont, 1919
Onthophagus ganalensis Gestro, 1895
Onthophagus gemellatus Raffray, 1877
Onthophagus geminatus d´Orbigny, 1907
Onthophagus gibbus d´Orbigny, 1913
Onthophagus giuseppecarpanetoi Tagliaferri ..., 2012
Onthophagus gradivus Balthasar, 1966
Onthophagus granosus d´Orbigny, 1913
Onthophagus granulifer (Harold, 1886)⇒ Onthophagus ponticus Harold, 1883
Onthophagus granulifrons Frey, 1961
Onthophagus granulopygus Dierkens, 2022
Onthophagus granulum d´Orbigny, 1904
Onthophagus graphicus Wallengren, 1881
Onthophagus gravearmatus Balthasar, 1944
Onthophagus haafi Frey, 1963
Onthophagus helciatus Harold, 1871
Onthophagus henus
Onthophagus hericius d´Orbigny, 1902
Onthophagus herus Péringuey, 1901
Onthophagus hildebrandti Harold, 1878
Onthophagus hinnulus (Klug, 1832)⇒ Mimonthophagus hinnulus Klug, 1832
Onthophagus hirtellus Frey, 1957
Onthophagus hirticulus d´Orbigny, 1915
Onthophagus hirtipodex d´Orbigny, 1904
Onthophagus horrens d´Orbigny, 1908
Onthophagus horridus d´Orbigny, 1908
Onthophagus hyaena Fabricius, 1801
Onthophagus ikelengeensis Josso, 2018
Onthophagus illotus Péringuey, 1901
Onthophagus imbellis d´Orbigny, 1905
Onthophagus impressicollis Boheman, 1860
Onthophagus impunctatus d´Orbigny, 1904
Onthophagus impurus Harold, 1868
Onthophagus inaequalis d´Orbigny, 1902
Onthophagus incantatus Balthasar, 1967
Onthophagus includens d´Orbigny, 1905
Onthophagus inclusus d´Orbigny, 1905
Onthophagus indigus Péringuey, 1901
Onthophagus ineptus Harold, 1871
Onthophagus inermiceps d´Orbigny, 1905
Onthophagus inermicollis d´Orbigny, 1908
Onthophagus infuscatus Klug, 1845
Onthophagus insignis (Péringuey, 1896)⇒ Proagoderus fossidorsis d´Orbigny, 1905
Onthophagus interruptus (Raffray, 1877)⇒ Onthophagus leanus Goidanich, 1926
Onthophagus interstitialis (Fahraeus, 1857)⇒ Euonthophagus carbonarius Klug, 1855
Onthophagus intonsus d´Orbigny, 1913
Onthophagus investis d´Orbigny, 1904
Onthophagus iringaensis Dierkens, 2022
Onthophagus jacksoni d´Orbigny, 1913
Onthophagus jeanneli d´Orbigny, 1913
Onthophagus jocquei Josso, 2019
Onthophagus jugicola d´Orbigny, 1902
Onthophagus juncticornis d´Orbigny, 1908
Onthophagus juniperensis Josso, 2019
Onthophagus juvencus Klug, 1835
Onthophagus kaomaensis Josso, 2022
Onthophagus kapangaensis Dierkens, 2022
Onthophagus karlwerneri Josso, 2022
Onthophagus katangensis Frey, 1961
Onthophagus kavirondi (d´Orbigny, 1913)⇒ Onthophagus kavirondus d´Orbigny, 1915
Onthophagus kavirondus d´Orbigny, 1915
Onthophagus kingstoni Cambefort, 1980
Onthophagus kirki Frey, 1975
Onthophagus kirokanus Frey, 1975
Onthophagus koma Matsumura, 1937
Onthophagus kyleensis Frey, 1975
Onthophagus laceratus (Gerstaecker, 1871)⇒ Onthophagus depressus Harold, 1871
Onthophagus lacustris Harold, 1877
Onthophagus laevatus d´Orbigny, 1902
Onthophagus laeviceps d´Orbigny, 1902
Onthophagus laevidorsis d´Orbigny, 1907
Onthophagus laevissimus d´Orbigny, 1905
Onthophagus lagnyi Paulian, 1937
Onthophagus lamelliger (Gerstaecker, 1871)⇒ Onthophagus ochropygus d´Orbigny, 1902
Onthophagus laminidorsis d´Orbigny, 1902
Onthophagus latepunctatus d´Orbigny, 1913
Onthophagus latestriatus d´Orbigny, 1908
Onthophagus latevittatus d´Orbigny, 1905
Onthophagus latipennis d´Orbigny, 1897
Onthophagus latissimus d´Orbigny, 1902
Onthophagus leanus Goidanich, 1926
Onthophagus lemekensis d´Orbigny, 1915
Onthophagus leroyi d´Orbigny, 1902
Onthophagus leucopygus (Harold, 1867)⇒ Onthophagus albipodex d´Orbigny, 1902
Onthophagus lilongweensis Dierkens, 2022
Onthophagus liodermus d´Orbigny, 1913
Onthophagus liopterus Harold, 1880
Onthophagus lituratus Roth, 1851
Onthophagus liwondeensis Josso, 2019
Onthophagus lomii Müller, 1942
Onthophagus longiceps d´Orbigny, 1904
Onthophagus loroi Balthasar, 1941
Onthophagus loveni Gillet, 1928
Onthophagus lugens (Fahraeus, 1857)⇒ Onthophagus fimetarius Roth, 1851
Onthophagus lugubris (Fahraeus, 1857)⇒ Onthophagus lituratus Roth, 1851
Onthophagus lunatus Harold, 1868
Onthophagus lutaticollis d´Orbigny, 1907
Onthophagus mabuensis Daniel, Strü..., 2023
Onthophagus maculosus d´Orbigny, 1908
Onthophagus malevolus Balthasar, 1964
Onthophagus marginipennis Josso, 2018
Onthophagus marshalli d´Orbigny, 1908
Onthophagus martaponsae
Onthophagus martelli Frey, 1972
Onthophagus martellii Frey, 1972
Onthophagus masaicus d´Orbigny, 1905
Onthophagus melanocephalus Klug, 1845
Onthophagus meruanus d´Orbigny, 1908
Onthophagus merus Péringuey, 1901
Onthophagus metalliger d´Orbigny, 1913
Onthophagus millingeni d´Orbigny, 1900
Onthophagus minans d´Orbigny, 1913
Onthophagus minettianus Josso, 2019
Onthophagus minutus (Hausmann, 1807)⇒ Onthophagus centricornis Fabricius, 1798
Onthophagus miricollis Frey, 1962
Onthophagus miricornis d´Orbigny, 1902
Onthophagus miscellaneus d´Orbigny, 1908
Onthophagus miscellus d´Orbigny, 1905
Onthophagus misellus d´Orbigny, 1907
Onthophagus mixticeps d´Orbigny, 1905
Onthophagus mixtidorsis d´Orbigny, 1905
Onthophagus mocquerysi d´Orbigny, 1902
Onthophagus monodon Fahraeus, 1857
Onthophagus montivagus d´Orbigny, 1902
Onthophagus morettoi Josso, 2022
Onthophagus morosus Gerstaecker, 1871
Onthophagus mugheseensis Josso, 2019
Onthophagus murphyi Josso, 2019
Onthophagus mwenseensis Dierkens, 2022
Onthophagus naevuliger d´Orbigny, 1908
Onthophagus nanus (Harold, 1878)⇒ Onthophagus waterhousei Gillet, 1927
Onthophagus nchilaensis Josso, 2022
Onthophagus neavei d´Orbigny, 1914
Onthophagus neofurcatus Goidanich, 1926
Onthophagus neostenocerus Goidanich, 1926
Onthophagus nigriceps Raffray, 1877
Onthophagus nigricornis Fairmaire, 1887
Onthophagus nigrinus Paulian, 1937
Onthophagus nigriventris d´Orbigny, 1902
Onthophagus nigroflavus Josso, 2022
Onthophagus niloticus Harold, 1879
Onthophagus nimbatus d´Orbigny, 1905
Onthophagus nitidulus Klug, 1845
Onthophagus nudatus d´Orbigny, 1913
Onthophagus nummelini Moretto & Gé..., 2020
Onthophagus nyikaensis Josso, 2018
Onthophagus obtusicornis (Fahraeus, 1857)⇒ Onthophagus marginalis przewalski Kabakov, 2006
Onthophagus ocellifer d´Orbigny, 1902
Onthophagus ochreatus d´Orbigny, 1897
Onthophagus ochropygus d´Orbigny, 1902
Onthophagus okahandjanus Balthasar, 1974
Onthophagus olidus Balthasar, 1964
Onthophagus omostigma d´Orbigny, 1902
Onthophagus orthocerus Thomson, 1858
Onthophagus ovulum Gerstaecker, 1871
Onthophagus pallidipennis (Fahraeus, 1857)⇒ Onthophagus rasipennis d´Orbigny, 1908
Onthophagus paravinctus Tagliaferri ..., 2012
Onthophagus parcenotatus d´Orbigny, 1913
Onthophagus parcepictus d´Orbigny, 1908
Onthophagus parenthesis Boucomont, 1913
Onthophagus parisii (Balthasar, 1941)⇒ Onthophagus lemur Fabricius, 1781
Onthophagus parumnotatus Fahraeus, 1857
Onthophagus parvifrons (d´Orbigny, 1907)⇒ Onthophagus semiasper d´Orbigny, 1902
Onthophagus paucigranosus Lansberge, 1886
Onthophagus pauxillus d´Orbigny, 1902
Onthophagus perniger Boucomont, 1930
Onthophagus peyrierasi Paulian, 1985
Onthophagus philippemorettoi Josso, 2022
Onthophagus picatus d´Orbigny, 1902
Onthophagus pictipodex d´Orbigny, 1902
Onthophagus pilicollis d´Orbigny, 1902
Onthophagus pilosus (Fahraeus, 1857)⇒ Onthophagus echinus Boucomont, 1914
Onthophagus pinguis Gerstaecker, 1871
Onthophagus pipitzi Ancey, 1883
Onthophagus placens Péringuey, 1904
Onthophagus planiceps (MacLeay, 1886)⇒ Onthophagus pinguis Gerstaecker, 1871
Onthophagus planidorsis Dierkens, 2022
Onthophagus plicatifrons d´Orbigny, 1908
Onthophagus politissimus d´Orbigny, 1908
Onthophagus pollicatus Harold, 1879
Onthophagus polyedrus d´Orbigny, 1905
Onthophagus polyodon d´Orbigny, 1913
Onthophagus polystigma d´Orbigny, 1902
Onthophagus posticicornis d´Orbigny, 1913
Onthophagus praedatus Harold, 1862
Onthophagus praestans Péringuey, 1901
Onthophagus praetortus Péringuey, 1904
Onthophagus probus Péringuey, 1901
Onthophagus producticollis d´Orbigny, 1908
Onthophagus pronocarinatus Josso, 2019
Onthophagus proteus d´Orbigny, 1902
Onthophagus protuberans Frey, 1955
Onthophagus pseudobidens d´Orbigny, 1915
Onthophagus pseudobufonidius Josso, 2019
Onthophagus pseudocribripennis Josso, 2019
Onthophagus pseudofimetarius Balthasar, 1946
Onthophagus pseudosanguineus Walter & Cam..., 1977
Onthophagus pseudosellatus Balthasar, 1964
Onthophagus pseudovinctus Tagliaferri ..., 2012
Onthophagus pugionatus Fahraeus, 1857
Onthophagus pulchellus d´Orbigny, 1905
Onthophagus pullus (Roth, 1851)⇒ Onthophagus parenthesis Boucomont, 1913
Onthophagus quadricuspis d´Orbigny, 1908
Onthophagus quadrilunatus d´Orbigny, 1902
Onthophagus quadrimaculatus Raffray, 1877
Onthophagus quadrinodosus Fahraeus, 1857
Onthophagus quadrinotatus d´Orbigny, 1905
Onthophagus ragazzii d´Orbigny, 1904
Onthophagus rasipennis d´Orbigny, 1908
Onthophagus rectilamina d´Orbigny, 1902
Onthophagus renejeanneli Dierkens, 2022
Onthophagus reticulatus d´Orbigny, 1902
Onthophagus reticulundaticeps Josso, 2022
Onthophagus rhodesianus Frey, 1975
Onthophagus rhynchophorus Péringuey, 1904
Onthophagus rhyncophorus
Onthophagus robertminettii Josso, 2022
Onthophagus rohwedderi Moretto, 2017
Onthophagus rojkoffianus Josso, 2018
Onthophagus rostrifer (Reitter, 1893)⇒ Onthophagus gibbulus Pallas, 1781
Onthophagus rotundatus d´Orbigny, 1905
Onthophagus ruandanus Frey, 1971
Onthophagus rubefactus (d´Orbigny, 1913)⇒ Onthophagus tessulatus Harold, 1871
Onthophagus rubens d´Orbigny, 1902
Onthophagus rubenticollis d´Orbigny, 1902
Onthophagus rubidus d´Orbigny, 1913
Onthophagus rubricatus d´Orbigny, 1902
Onthophagus rufobasalis Fairmaire, 1887
Onthophagus rufocastaneus Fairmaire, 1887
Onthophagus rufolimbatus d´Orbigny, 1913
Onthophagus rufonotatus d´Orbigny, 1902
Onthophagus rugidorsis d´Orbigny, 1913
Onthophagus rugosissimus d´Orbigny, 1913
Onthophagus rugulipennis Fairmaire, 1887
Onthophagus sansibaricus Harold, 1878
Onthophagus sapphirinus (Fahraeus, 1857)⇒ Proagoderus aureiceps d´Orbigny, 1902
Onthophagus scaber Roth, 1851
Onthophagus scaberrimus d´Orbigny, 1904
Onthophagus schaufussi Harold, 1867
Onthophagus sculptilis Gerstaecker, 1871
Onthophagus secundarius Roth, 1851
Onthophagus sellatulus d´Orbigny, 1902
Onthophagus sellatus Klug, 1845
Onthophagus semiaratus d´Orbigny, 1902
Onthophagus semiasper d´Orbigny, 1902
Onthophagus semiflavus (Boheman, 1860)⇒ Onthophagus flavolimbatus Klug, 1855
Onthophagus semilaevis d´Orbigny, 1913
Onthophagus senegalensis d´Orbigny, 1902
Onthophagus sericeicollis Boucomont, 1928
Onthophagus setosus Fahraeus, 1857
Onthophagus shimba Cambefort, 1980
Onthophagus sidama Gestro, 1895
Onthophagus signatus Fahraeus, 1857
Onthophagus simillimus d´Orbigny, 1913
Onthophagus simoni d´Orbigny, 1902
Onthophagus simplex Raffray, 1877
Onthophagus simpliciceps d´Orbigny, 1905
Onthophagus simulator d´Orbigny, 1905
Onthophagus sinuosicollis d´Orbigny, 1905
Onthophagus smeenki Cambefort, 1980
Onthophagus snoflaki Balthasar, 1944
Onthophagus somalicola Balthasar, 1941
Onthophagus sophiae Nicolas, 2006
Onthophagus splendidoides Josso, 2013
Onthophagus splendidus Boucomont, 1932
Onthophagus sprecherae Moretto, 2013
Onthophagus spurcatus d´Orbigny, 1905
Onthophagus stellio Erichson, 1843
Onthophagus stenocerus (Harold, 1867)⇒ Onthophagus neostenocerus Goidanich, 1926
Onthophagus sticticus Harold, 1867
Onthophagus stictus d´Orbigny, 1913
Onthophagus stigmosus d´Orbigny, 1902
Onthophagus stillatus d´Orbigny, 1904
Onthophagus stomachosus Krell, 2009
Onthophagus stuhlmanni d´Orbigny, 1908
Onthophagus subcinctus d´Orbigny, 1913
Onthophagus subdivisus d´Orbigny, 1908
Onthophagus subhumeralis d´Orbigny, 1902
Onthophagus subocellatus d´Orbigny, 1905
Onthophagus subrugosus d´Orbigny, 1905
Onthophagus suffusus Klug, 1855
Onthophagus sugillatus Klug, 1855
Onthophagus sulcipennis d´Orbigny, 1902
Onthophagus sutiliceps d´Orbigny, 1902
Onthophagus taboranus d´Orbigny, 1902
Onthophagus talpa Fahraeus, 1857
Onthophagus tanganus d´Orbigny, 1913
Onthophagus teitanicus d´Orbigny, 1902
Onthophagus tenax Balthasar, 1964
Onthophagus tenuigraniger d´Orbigny, 1913
Onthophagus tersicollis Müller, 1947
Onthophagus tersipennis d´Orbigny, 1902
Onthophagus tersus d´Orbigny, 1913
Onthophagus tesseratus d´Orbigny, 1908
Onthophagus tessulatus Harold, 1871
Onthophagus thomsoni Bates, 1888
Onthophagus tonsus d´Orbigny, 1902
Onthophagus transcaspicus Koenig, 1888
Onthophagus trapezicornis d´Orbigny, 1902
Onthophagus traversii d´Orbigny, 1904
Onthophagus tricariniger d´Orbigny, 1902
Onthophagus trifidisetis d´Orbigny, 1905
Onthophagus trinodosus Fahraeus, 1857
Onthophagus trinominatus Goidanich, 1926
Onthophagus tubericollis (Raffray, 1877)⇒ Onthophagus bidens Olivier, 1789
Onthophagus tumidulus Gerstaecker, 1871
Onthophagus turneri Boucomont, 1936
Onthophagus ugandaensis Dierkens, 2022
Onthophagus ugandicus Frey, 1975
Onthophagus ukerewensis Balthasar, 1937
Onthophagus umbilicatus d´Orbigny, 1908
Onthophagus undaticeps d´Orbigny, 1902
Onthophagus unicarina d´Orbigny, 1904
Onthophagus ursinus d´Orbigny, 1902
Onthophagus ursus Boucomont, 1926
Onthophagus usambaricus Paulian, 1937
Onthophagus valentineae Josso, 2019
Onthophagus variatus d´Orbigny, 1902
Onthophagus variegatus (Fabricius, 1798)⇒ Onthophagus filicornis Harold, 1873
Onthophagus variegranosus d´Orbigny, 1905
Onthophagus varius d´Orbigny, 1913
Onthophagus vassei d´Orbigny, 1908
Onthophagus vatovai Müller, 1942
Onthophagus venzoi Balthasar, 1941
Onthophagus verae Josso, 2013
Onthophagus verrucosus d´Orbigny, 1902
Onthophagus versutus Péringuey, 1901
Onthophagus verticalis (Fahraeus, 1857)⇒ Onthophagus praedatus Harold, 1862
Onthophagus vinctoides Frey, 1957
Onthophagus vinctus Erichson, 1843
Onthophagus violaceoviridis Paulian, 1937
Onthophagus virens (d´Orbigny, 1913)⇒ Onthophagus alcyonides d´Orbigny, 1913
Onthophagus virescens Harold, 1867
Onthophagus waterhousei Gillet, 1927
Onthophagus xanthopterus d´Orbigny, 1908
Onthophagus xanthopygus d´Orbigny, 1908
Onthophagus zambundaticeps Josso, 2022
Onthophagus zavattarii Müller, 1947
Pachylomera femoralis Kirby, 1828
Pachylomera opaca Lansberge, 1874
Pachysoma bennigseni Felsche, 1907
Panelus rhodesiensis Paulian, 1975
Paracanthon trichonotulus Balthasar, 1938
Paronitis excavaticollis Balthasar, 1942
Pedaria acuta Josso & Prév..., 2015
Pedaria aethiopica Josso & Prév..., 2003
Pedaria alternans Waterhouse, 1890
Pedaria antoinei Josso & Prév..., 2009
Pedaria armata Raffray, 1877
Pedaria barrei Josso & Prév..., 2006
Pedaria biseria Josso & Prév..., 2003
Pedaria brancoi Josso & Prév..., 2003
Pedaria cambeforti Josso & Prév..., 2003
Pedaria catherinae Josso & Prév..., 2009
Pedaria cicatricosus (Péringuey, 1901)⇒ Sarophorus cicatricosus Péringuey, 1901
Pedaria conformis Péringuey, 1901
Pedaria cylindrica Fahraeus, 1857
Pedaria decorsei (Boucomont, 1922)⇒ Pedaria taylori Waterhouse, 1890
Pedaria demeyeri Josso & Prév..., 2003
Pedaria dentata Raffray, 1877
Pedaria dudleyi Josso & Prév..., 2003
Pedaria genierorum Josso & Prév..., 2015
Pedaria gigantea Josso & Prév..., 2003
Pedaria hanae Balthasar, 1963
Pedaria insularis Kolbe, 1914
Pedaria intermedia Boucomont, 1922
Pedaria jacksoni Waterhouse, 1890
Pedaria jossoi Génier, 2017
Pedaria juhellegrandi Josso & Prév..., 2003
Pedaria manni Josso & Prév..., 2015
Pedaria minetti Josso & Prév..., 2009
Pedaria minima Josso & Prév..., 2003
Pedaria montreuili Josso & Prév..., 2015
Pedaria murphyi Josso & Prév..., 2003
Pedaria murphyi murphyi
Pedaria murphyi septentrionalis Josso & Prév..., 2009
Pedaria ovata Boucomont, 1922
Pedaria puncticollis Waterhouse, 1890
Pedaria quirimbasensis Josso & Prév..., 2015
Pedaria raffrayi Josso & Prév..., 2006
Pedaria reichenbachi Josso & Prév..., 2006
Pedaria renwarti Cambefort, 1982
Pedaria rohani Boucomont, 1922
Pedaria rosemariae Josso & Prév..., 2009
Pedaria segregis Péringuey, 1901
Pedaria segrestis
Pedaria smrzi Josso & Prév..., 2003
Pedaria spinipennis Josso & Prév..., 2003
Pedaria spinosa Boucomont, 1922
Pedaria sudrei Josso & Prév..., 2003
Pedaria tanganyika Josso & Prév..., 2003
Pedaria taylori Waterhouse, 1890
Pedaria tenebrosa Balthasar, 1937
Pedaria tibialis Josso & Prév..., 2006
Pedaria tuberculigera Waterhouse, 1890
Pedaria werneri Josso & Prév..., 2003
Pedaria wittei Janssens, 1939
Phalops ardea Klug, 1855
Phalops aurifrons Fairmaire, 1884
Phalops barbicornis Lansberge, 1883
Phalops beccarii Harold, 1875
Phalops boschas Klug, 1855
Phalops djuricus Kuntzen, 1913
Phalops dregei Harold, 1867
Phalops faccai Müller, 1941
Phalops fimbriatus Klug, 1835
Phalops flavocinctus Klug, 1855
Phalops gallanus d´Orbigny, 1904
Phalops glauningi Kuntzen, 1913
Phalops guttulatus Fairmaire, 1887
Phalops inermis Lansberge, 1883
Phalops laminifrons Fairmaire, 1882
Phalops lutatus d´Orbigny, 1902
Phalops princeps d´Orbigny, 1908
Phalops rufosignatus (Lansberge, 1885)⇒ Phalops dregei Harold, 1867
Phalops sinuaticeps d´Orbigny, 1902
Phalops smaragdinus Harold, 1875
Phalops tricuspis d´Orbigny, 1908
Phalops tuberosus d´Orbigny, 1905
Phalops wittei Harold, 1867
Phalops zuninoi Barbero & Pa..., 2003
Phanaeus (Phanaeus) vindex MacLeay, 1819
Pinacopodius meruensis Krikken, 1982
Pinacopodius praestolatus Branco & Josso, 2007
Pinacotarsus dohrni Harold, 1875
Pinacotarsus proximus Ferreira, 1969
Platyonitis oberthueri Janssens, 1942
Platyonitis smeenkorum Krikken, 1974
Pleuronitis fulgidus Castelnau, 1840
Pleuronitis minettii Josso, 2011
Proagoderus aciculatus Fahraeus, 1857
Proagoderus albicapillus d´Orbigny, 1902
Proagoderus amblychromatus Marcus, 1917
Proagoderus armicollis d´Orbigny, 1904
Proagoderus atriclaviger d´Orbigny, 1905
Proagoderus atrosetosus d´Orbigny, 1902
Proagoderus auratus Fabricius, 1801
Proagoderus aureiceps d´Orbigny, 1902
Proagoderus biarmatus d´Orbigny, 1908
Proagoderus bicallosus Klug, 1855
Proagoderus boranus Gestro, 1895
Proagoderus bottegi Gestro, 1895
Proagoderus brucei (d´Orbigny, 1902)⇒ Onthophagus chalcostolus d´Orbigny, 1902
Proagoderus brucei brucei
Proagoderus brucei cyanostolus d´Orbigny, 1902
Proagoderus brucei njombeensis Moretto, 2004
Proagoderus camiadei Josso, 2014
Proagoderus chalcostolus d´Orbigny, 1902
Proagoderus chrysopes Bates, 1888
Proagoderus cyanopterus d´Orbigny, 1905
Proagoderus dargei Moretto, 2013
Proagoderus dives Harold, 1877
Proagoderus dudleyi Cambefort, 1980
Proagoderus edithae Josso, 2014
Proagoderus elgoni d´Orbigny, 1915
Proagoderus euchlorus d´Orbigny, 1902
Proagoderus extensus Harold, 1878
Proagoderus flexicollis d´Orbigny, 1904
Proagoderus fossidorsis d´Orbigny, 1905
Proagoderus furcifer Boheman, 1860
Proagoderus furciramus d´Orbigny, 1902
Proagoderus gemmatus Péringuey, 1901
Proagoderus gerstaeckeri Harold, 1867
Proagoderus gibbiramus d´Orbigny, 1902
Proagoderus harpax Fabricius, 1801
Proagoderus hemicyaneus
Proagoderus hemicyanus d´Orbigny, 1902
Proagoderus ignitus d´Orbigny, 1905
Proagoderus incostatus d´Orbigny, 1902
Proagoderus kachowskii Olsoufief, 1900
Proagoderus kafueensis Josso, 2014
Proagoderus kilimanus Kolbe, 1891
Proagoderus kolbei Marcus, 1921
Proagoderus kwaleensis Moretto & Ni..., 2002
Proagoderus lallieri d´Orbigny, 1902
Proagoderus lanista Castelnau, 1840
Proagoderus longefossus d´Orbigny, 1905
Proagoderus loricatus Klug, 1855
Proagoderus louisi Josso, 2014
Proagoderus lucasi d´Orbigny, 1902
Proagoderus lujendae Bates, 1888
Proagoderus marginidens d´Orbigny, 1904
Proagoderus maringouensis Palestrini, 1992
Proagoderus minettii Moretto & Ni..., 2002
Proagoderus moorei Moretto, 2013
Proagoderus multicornis d´Orbigny, 1908
Proagoderus nasidens d´Orbigny, 1905
Proagoderus negus Raffray, 1882
Proagoderus nigricornis Fairmaire, 1887
Proagoderus nuba d´Orbigny, 1902
Proagoderus palestriniae Moretto & Ni..., 2004
Proagoderus panoplus Bates, 1888
Proagoderus paolae Palestrini, 1992
Proagoderus pegesimallus Marcus, 1917
Proagoderus plato Bates, 1888
Proagoderus porrectus Reiche, 1847
Proagoderus prostans Reiche, 1847
Proagoderus pseudoflexicollis Moretto, 2004
Proagoderus pyramidalis Klug, 1855
Proagoderus quadriarmatus Fairmaire, 1892
Proagoderus quadricristatus d´Orbigny, 1908
Proagoderus quadrituber d´Orbigny, 1908
Proagoderus ramosicornis d´Orbigny, 1902
Proagoderus rangifer Klug, 1855
Proagoderus rarus Guérin-Mén..., 1829
Proagoderus rectefurcatus Fairmaire, 1891
Proagoderus revoili Lansberge, 1882
Proagoderus sapphirinus (Péringuey, 1901)⇒ Onthophagus aureiceps d´Orbigny, 1902
Proagoderus sexcornutus d´Orbigny, 1902
Proagoderus somalicus d´Orbigny, 1902
Proagoderus subextensus Kolbe, 1895
Proagoderus triarmatus d´Orbigny, 1904
Proagoderus tricornifrons d´Orbigny, 1908
Proagoderus uluguru Frey, 1975
Proagoderus versus d´Orbigny, 1913
Proagoderus werneri Moretto & Ni..., 2002
Proagoderus worsissa Roth, 1851
Pseudochironitis bennigseni Felsche, 1907
Pseudochironitis benningseni Felsche, 1907
Pseudochironitis stuhlmanni Kolbe, 1897
Pseudocopris lunaticeps Gillet, 1910
Pseudocopris tubericollis Felsche, 1910
Pseudosaproecius binasus d´Orbigny, 1907
Pseudosaproecius cylindroides d´Orbigny, 1908
Pseudosaproecius mirepunctatus d´Orbigny, 1905
Pseudosaproecius portentosus Felsche, 1907
Pseudosaproecius validicornis Quedenfeldt, 1884
Sarophorus cicatricosus Péringuey, 1901
Sarophorus costatus Fahraeus, 1857
Sarophorus nitidus Frolov & Sch..., 2003
Scaptocnemis segregis Péringuey, 1901
Scarabaeus (Scarabaeus) pius Illiger, 1803
Scarabaeus (Scarabaeus) sacer (Linnaeus, 1758)⇒ Scarabaeus pius Illiger, 1803
Scarabaeus aegaeon Drury, 1773
Scarabaeus aegyptiacus Stolfa, 1938
Scarabaeus aegyptiorum Latreille, 1827
Scarabaeus aegyptiorum purpurascens Gerstaecker, 1871
Scarabaeus anderseni Waterhouse, 1890
Scarabaeus andreaei zur Strassen, 1963
Scarabaeus basuto zur Strassen, 1962
Scarabaeus bohemani Harold, 1868
Scarabaeus bornemisszai zur Strassen, 1980
Scarabaeus brahminus Castelnau, 1840
Scarabaeus caffer Boheman, 1857
Scarabaeus catenatus Gerstaecker, 1871
Scarabaeus clanceyi Ferreira, 1954
Scarabaeus confusus Müller, 1942
Scarabaeus convexus Hausmann, 1807
Scarabaeus deludens zur Strassen, 1961
Scarabaeus ebenus Klug, 1855
Scarabaeus ermienae Deschodt & D..., 2015
Scarabaeus fraterculus Kolbe, 1895
Scarabaeus funebris Boheman, 1857
Scarabaeus galenus Westwood, 1847
Scarabaeus gangeticus Castelnau, 1840
Scarabaeus geminogalenus Davis & Desc..., 2017
Scarabaeus goryi Castelnau, 1840
Scarabaeus gracai Ferreira, 1952
Scarabaeus indus (Linnaeus, 1758)⇒ Euphoria inda Linnaeus, 1758
Scarabaeus interstitialis Boheman, 1857
Scarabaeus intricatus Fabricius, 1801
Scarabaeus isidis Castelnau, 1840
Scarabaeus janssensi Balthasar, 1940
Scarabaeus karlwerneri Nicolas & Mo..., 2002
Scarabaeus laevifrons Fairmaire, 1884
Scarabaeus licitus Ferreira, 1954
Scarabaeus montreuili meridionalis Moretto, 2023
Scarabaeus montreuili montreuili
Scarabaeus nepos (Fairmaire, 1884)⇒ Scarabaeus goryi Castelnau, 1840
Scarabaeus nitidicollis Lansberge, 1882
Scarabaeus opacipennis Fairmaire, 1884
Scarabaeus osiridis Moretto, 2023
Scarabaeus palemo Olivier, 1789
Scarabaeus planifrons Fairmaire, 1884
Scarabaeus planipennis Davis & Desc..., 2015
Scarabaeus platynotus Bates, 1888
Scarabaeus poggei Waterhouse, 1890
Scarabaeus prodigiosus Erichson, 1843
Scarabaeus profanus Fabricius, 1792
Scarabaeus radama Fairmaire, 1895
Scarabaeus rixosus Péringuey, 1901
Scarabaeus satyrus (Boheman, 1860)⇒ Xyloryctes jamaicensis Drury, 1773
Scarabaeus scholtzi Mostert & Holm, 1982
Scarabaeus sevoistra Alluaud, 1902
Scarabaeus sulcipennis Quedenfeldt, 1888
Scarabaeus tonckeri Boucomont, 1923
Scarabaeus venerabilis Harold, 1871
Scarabaeus vethi Lansberge, 1886
Scarabaeus viettei Paulian, 1953
Scarabaeus westwoodi (Harold, 1869)⇒ Scarabaeus galenus Westwood, 1847
Scarabaeus xavieri Ferreira, 1968
Scarabaeus zambesianus Péringuey, 1901
Sceliages adamastor Lepeletier &..., 1828
Sceliages augias Gillet, 1908
Sceliages difficilis zur Strassen, 1965
Sceliages gagates Shipp, 1895
Sisyphus (Sisyphus) struwei Schafer & Fi..., 1992
Sisyphus struwei kivuensis Schafer & Fi..., 2001
Sisyphus struwei struwei
Sisyphus africanus orientalis Montreuil, 2017
Sisyphus alveatus Boucomont, 1935
Sisyphus angulicollis (Felsche, 1909)⇒ Neosisyphus angulicollis Felsche, 1909
Sisyphus astriatus Schafer & Fi..., 2001
Sisyphus auricomus Daniel & Davis, 2018
Sisyphus barbarossa (Wiedemann, 1823)⇒ Neosisyphus spinipes Thunberg, 1818
Sisyphus basilewskyi (Haaf, 1956)⇒ Neosisyphus basilewskyi Haaf, 1956
Sisyphus biarmatus Felsche, 1909
Sisyphus bicuariensis Daniel & Davis, 2018
Sisyphus caffer Boheman, 1857
Sisyphus caffer goryi Harold, 1859
Sisyphus calcaratus (Boucomont, 1937)⇒ Neosisyphus rugosus Gory, 1833
Sisyphus costatus Thunberg, 1818
Sisyphus crispatus Gory, 1833
Sisyphus crispatus crispatus
Sisyphus dierkensi Montreuil, 2015
Sisyphus ethiopicus Montreuil, 2017
Sisyphus fortuitus Péringuey, 1900
Sisyphus gazanus Arrow, 1909
Sisyphus genierorum Montreuil, 2015
Sisyphus goryi (Harold, 1859)⇒ Sisyphus caffer goryi Harold, 1859
Sisyphus granulifer Montreuil, 2014
Sisyphus ibaya Montreuil, 2017
Sisyphus impressipennis Lansberge, 1886
Sisyphus inconspicuus Daniel & Davis, 2018
Sisyphus infuscatus (Klug, 1855)⇒ Neosisyphus spinipes Thunberg, 1818
Sisyphus jossoi Montreuil, 2015
Sisyphus juheli Montreuil, 2015
Sisyphus kibale Montreuil, 2016
Sisyphus manni Montreuil, 2015
Sisyphus minettii Montreuil, 2014
Sisyphus mirabilis Arrow, 1927
Sisyphus mkomazi Montreuil, 2017
Sisyphus moorei Montreuil, 2015
Sisyphus morenoi Montreuil, 2015
Sisyphus morettoi Montreuil, 2015
Sisyphus muricatus Olivier, 1789
Sisyphus nanniscus Péringuey, 1901
Sisyphus negus Montreuil, 2015
Sisyphus neobornemisszanus Daniel & Davis, 2016
Sisyphus nodifer Gerstaecker, 1871
Sisyphus ocellatus Reiche, 1847
Sisyphus ocellatus nanniscus Péringuey, 1901
Sisyphus ongo Montreuil, 2017
Sisyphus oralensis Daniel & Davis, 2016
Sisyphus penicillatus Harold, 1880
Sisyphus quadricollis (Péringuey, 1901)⇒ Neosisyphus spinipes Thunberg, 1818
Sisyphus rubrus (Paschalidis, 1974)⇒ Neosisyphus rubrus Paschalidis, 1974
Sisyphus rugosus (Roth, 1851)⇒ Sisyphus costatus Thunberg, 1818
Sisyphus seminulum (Gerstaecker, 1871)⇒ Sisyphus costatus Thunberg, 1818
Sisyphus setiger (Roth, 1851)⇒ Neosisyphus confrater Kolbe, 1914
Sisyphus sordidus (Boheman, 1857)⇒ Sisyphus caffer Boheman, 1857
Sisyphus spinipes (Thunberg, 1818)⇒ Neosisyphus spinipes Thunberg, 1818
Sisyphus tibialis Raffray, 1877
Sisyphus transvaalensis (Péringuey, 1908)⇒ Sisyphus impressipennis Lansberge, 1886
Sisyphus trichodichromicus Montreuil, 2015
Sisyphus trochantericus (Fairmaire, 1891)⇒ Neosisyphus spinipes Thunberg, 1818
Sisyphus usambaricus Montreuil, 2015
Sisyphus werneri Montreuil, 2017
Stiptocnemis gilleti Boucomont, 1923
Stiptocnemis hiekei Branco, 1991
Stiptopodius doriae Harold, 1871
Stiptopodius glabricollis Müller, 1942
Stiptopodius granulosus Branco, 2009
Stiptopodius krausei Branco, 1991
Stiptopodius longipedis Branco, 1991
Stiptopodius nitidus Boucomont, 1923
Stiptopodius patrizii Boucomont, 1923
Stiptopodius singularis Péringuey, 1901
Sukelus jessopi Branco, 1991
Sulcodrepanus sulcicollis Castelnau, 1840
Tanzanolus leleupi Balthasar, 1960
Tibiodrepanus sulcicollis Castelnau, 1840
Tiniocellus dolosus Branco, 2010
Tiniocellus setifer Kraatz, 1895
Tiniocellus spinipes Roth, 1851
Tomogonus cambeforti Nicolas, 2002
Tomogonus chauvetorum Walter, 2012
Tomogonus crassoides Boucomont, 1923
Tomogonus crassus d´Orbigny, 1902
Tomogonus samburu Walter, 2012
Tragiscus dimidiatus Klug, 1855
Tropidonitis paradoxus Boheman, 1857
Unidentis vwaza Josso & Prév..., 2008
Xinidium davisi Cambefort, 1985
Xinidium dewitzi Harold, 1878

Unterknoten von Scarabaeinae mit Vorkommen in Ost-Afrika
TribusAteuchini
Montreuil, 1998
4
68
Gattungen
Arten

Taxom

TribusCanthonini
Péringuey, 1900
19
241
Gattungen
Arten

Taxom

TribusCoprini
Kolbe, 1805
10
257
Gattungen
Arten

Taxom

TribusGymnopleurini
Lacordaire, 1856
3
53
Gattungen
Arten

Taxom

TribusOniticellini
d´Orbigny, 1916
19
147
Gattungen
Arten

Taxom

TribusOnitini
Hope, 1837
14
136
Gattungen
Arten

Taxom

TribusOnthophagini
Lacordaire, 1856
Kotkäfer
27
821
Gattungen
Arten

Taxom

TribusPhanaeini
MacLeay, 1819
1
1
Gattung
Art

Taxom

TribusScarabaeini
Latreille, 1802
7
85
Gattungen
Arten

Taxom

TribusSisyphini
Mulsant, 1842
3
77
Gattungen
Arten

Taxom



Links, Quellen und weitere Informationen
Scarabaeinae bei artfakta.se
Scarabaeinae bei dyntaxa.se
Scarabaeinae bei itis.gov