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| | | | Checkliste der Unterfamilie Scarabaeinae Latreille, 1802 (Pillendreher)
für die Region Afrika (Kontinente)
Klasse
| Insecta
Linnaeus, 1758
Freikiefler, Insekten, Kerbtiere, Kerfe | 943
36180
241410 | Familien
Gattungen
Arten |
Taxom
|
Unterklasse
| Pterygota
Lang, 1888
Fluginsekten | 939
36100
241106 | Familien
Gattungen
Arten |
Taxom
|
Teilklasse
| Neoptera
van der Wulp, 1890
Neuflügler | 896
35668
239021 | Familien
Gattungen
Arten |
Taxom
|
Überordnung
| Holometabola
Heider, 1889
Holometabolen Insekten | 524
27533
203737 | Familien
Gattungen
Arten |
Taxom
|
Ordnung
| Coleoptera
Linnaeus, 1758
Käfer | 146
11745
86825 | Familien
Gattungen
Arten |
Taxom
|
Unterordnung
| Polyphaga
Emery, 1886
| 125
10691
74394 | Familien
Gattungen
Arten |
Taxom
|
Teilordnung
| Scarabaeiformia
Crowson, 1960
| 15
1621
14333 | Familien
Gattungen
Arten |
Taxom
|
Überfamilie
| Scarabaeoidea
Latreille, 1802
Blatthornkäfer | 15
1621
14333 | Familien
Gattungen
Arten |
Taxom
| | Familie | Scarabaeidae
Latreille, 1802
Blatthornkäfer | 1437
13254 | Gattungen
Arten |
Taxom
| | Unterfamilie | Scarabaeinae
Latreille, 1802
Pillendreher | 162
3178 | Gattungen
Arten |
Taxom
|
Gattungen für Afrika
162 Gattungen mit 3178 Arten #ABCDEFGHIJKLMNOPSTUVWX
| Acanthonitis Janssens, 2012 | | 4 | Arten | Taxom | | Afrodrepanus Krikken, 1921 | | 2 | Arten | Taxom | | Agamopus Bates, 1887 | | 1 | Art | Taxom | | Aleiantus Olsoufieff, 1898 | | 4 | Arten | Taxom | | Aliuscanthoniola Deschodt & S..., 2008 | | 1 | Art | Taxom | | Allogymnopleurus Janssens, 1941 | | 18 | Arten | Taxom | | Allonitis Janssens, 1907 | | 1 | Art | Taxom | | Alloscelus Boucomont, 1923 | | 4 | Arten | Taxom | | Amietina Cambefort, 1989 | | 4 | Arten | Taxom | | Anachalcos Hope, 1901 | | 6 | Arten | Taxom | | Anonychonitis Janssens, 1950 | | 1 | Art | Taxom | | Aphengoecus Péringuey, 1987 | | 2 | Arten | Taxom | | Apotolamprus Olsofieff, 2008 | | 32 | Arten | Taxom | | Apterepilissus Montreuil, 1958 | | 4 | Arten | Taxom | | Aptychonitis Janssens, 1868 | | 2 | Arten | Taxom | | Arachnodes Westwood, 2011 | | 63 | Arten | Taxom | | Ateuchus Weber, 1996 | | 7 | Arten | Taxom | | Bohepilissus Paulian, 1857 | | 2 | Arten | Taxom | | Bubas Mulsant, 1811 | | 3 | Arten | Taxom | | Byrrhidium Harold, 1869 | | 3 | Arten | Taxom | | Caccobius Thomson, 2014 | | 74 | Arten | Taxom | | Cambefortantus Paulian, 1975 | | 8 | Arten | Taxom | | Cambefortius Branco, 1936 | | 8 | Arten | Taxom | | Canthidium Erichson, 1981 | | 18 | Arten | Taxom | | Canthochilum Chapin, 2008 | | 1 | Art | Taxom | | Canthodimorpha Davis, Schol..., 1999 | | 1 | Art | Taxom | | Canthon Hoffmannsegg, 1964 | | 14 | Arten | Taxom | | Catharsiocopris | | 1 | Art | Taxom | | Catharsius Hope, 1857 | | 104 | Arten | Taxom | | Cheironitis van Lansberge, 1918 | | 18 | Arten | Taxom | | Chironitis Lansberge, 1849 | | 3 | Arten | Taxom | | Circellium Latreille, 1781 | | 1 | Art | Taxom | | Cleptocaccobius Cambefort, 1967 | | 17 | Arten | Taxom | | Clypeodrepanus Krikken, 1953 | | 3 | Arten | Taxom | | Copridaspidus Bloucomont, 1878 | | 1 | Art | Taxom | | Copris Geoffroy, 1967 | | 153 | Arten | Taxom | | Coprophanaeus Olsoufieff, 2008 | | 5 | Arten | Taxom | | Coptodactyla Burmeister, 1877 | | 1 | Art | Taxom | | Coptorhina Hope, 1857 | | 9 | Arten | Taxom | | Cyptochirus Lesne, 1851 | | 4 | Arten | Taxom | | Delopleurus Erichson, 1857 | | 9 | Arten | Taxom | | Deltochilum Eschscholtz, 1938 | | 5 | Arten | Taxom | | Diastellopalpus van Lansberge, 2000 | | 36 | Arten | Taxom | | Dichotomius Hope, 1867 | | 6 | Arten | Taxom | | Dicranocara | | 4 | Arten | Taxom | | Digitonthophagus Balthasar, 2017 | | 14 | Arten | Taxom | | Disphysema Harold, 1873 | | 1 | Art | Taxom | | Dorbignyolus Branco, 1913 | | 1 | Art | Taxom | | Drepanocerus Kirby, 1851 | | 16 | Arten | Taxom | | Drepanoplatynus Boucomont, 1921 | | 1 | Art | Taxom | | Drepanopodus Janssens, 1901 | | 2 | Arten | Taxom | | Dwesasilvasedis | | 1 | Art | Taxom | | Endroedyolus Scholtz & Ho..., 1987 | | 1 | Art | Taxom | | Eodrepanus Barbero, Pal..., 1877 | | 5 | Arten | Taxom | | Epactoides Olsoufieff, 1976 | | 39 | Arten | Taxom | | Epilissus Blanchard, 2022 | | 42 | Arten | Taxom | | Epirinus Reiche, 1901 | | 36 | Arten | Taxom | | Euoniticellus Janssens, 1953 | | 17 | Arten | Taxom | | Euonthophagus Balthasar, 1901 | | 16 | Arten | Taxom | | Eurysternus Dalman, 1880 | | 4 | Arten | Taxom | | Eusaproecius Branco, 1923 | | 7 | Arten | Taxom | | Eutrichillum | | 1 | Art | Taxom | | Frankenbergerius | | 9 | Arten | Taxom | | Garreta Janssens, 1967 | | 25 | Arten | Taxom | | Gilletellus Janssens, 1857 | | 2 | Arten | Taxom | | Gymnopleurus Illiger, 1843 | | 49 | Arten | Taxom | | Hammondantus Cambefort, 1978 | | 1 | Art | Taxom | | Haroldius Boucomont, 2010 | | 6 | Arten | Taxom | | Helictopleurus Orbigny, 1898 | | 79 | Arten | Taxom | | Heliocopris Hope, 2013 | | 62 | Arten | Taxom | | Heteroclitopus Péringuey, 1991 | | 10 | Arten | Taxom | | Heteronitis Gillet, 1840 | | 5 | Arten | Taxom | | Heterosyphus Paulian, 1975 | | 1 | Art | Taxom | | Hyalonthophagus Palestrini &..., 1910 | | 12 | Arten | Taxom | | Ixodina Roth, 0 | | 12 | Arten | Taxom | | Janssensantus Paulian, 1987 | | 3 | Arten | Taxom | | Janssensellus Cambefort, 1995 | | 2 | Arten | Taxom | | Kheper Janssens, 1986 | | 30 | Arten | Taxom | | Kolbeellus Jacobson, 1875 | | 1 | Art | Taxom | | Krikkenius Branco, 1982 | | 1 | Art | Taxom | | Latodrepanus Krikken, 2009 | | 6 | Arten | Taxom | | Lepanus Balthasar, 1888 | | 2 | Arten | Taxom | | Liatongus Reitter, 1914 | | 17 | Arten | Taxom | | Litocopris Waterhouse, 1869 | | 4 | Arten | Taxom | | Lophodonitis Janssens, 2011 | | 2 | Arten | Taxom | | Macroderes Westwood, 1880 | | 25 | Arten | Taxom | | Madateuchus Paulian, 1953 | | 1 | Art | Taxom | | Malagoniella Martínez, 1789 | | 1 | Art | Taxom | | Megalonitis Janssens, 1855 | | 3 | Arten | Taxom | | Megaponerophilus Janssens, 1914 | | 1 | Art | Taxom | | Metacatharsius Paulian, 1965 | | 66 | Arten | Taxom | | Milichus Péringuey, 1907 | | 17 | Arten | Taxom | | Mimonthophagus Balthasar, 1902 | | 8 | Arten | Taxom | | Mnematidium Ritsema, 1845 | | 2 | Arten | Taxom | | Mnematium MacLeay, 1821 | | 1 | Art | Taxom | | Namakwanus Scholtz & Ho..., 2005 | | 7 | Arten | Taxom | | Nanos Westwood, 2011 | | 44 | Arten | Taxom | Neateuchus Gillet, 1911 | | 0 | Arten | Taxom | | Nebulasilvius Deschodt & C..., 2008 | | 2 | Arten | Taxom | | Neonitis Péringuey, 1918 | | 2 | Arten | Taxom | | Neosaproecius Branco, 1958 | | 2 | Arten | Taxom | | Neosisyphus Müller, 1891 | | 25 | Arten | Taxom | | Nesosisyphus Vinson, 1939 | | 4 | Arten | Taxom | | Nesovinsonia Martinez & P..., 1939 | | 1 | Art | Taxom | | Nitiocellus Branco, 1921 | | 2 | Arten | Taxom | | Odontoloma Boheman, 1987 | | 20 | Arten | Taxom | | Oniticellus Serville, 1879 | | 12 | Arten | Taxom | | Onitis Fabricius, 2012 | | 158 | Arten | Taxom | | Ontherus Erichson, 1996 | | 3 | Arten | Taxom | | Onthophagus Latreille, 1954 | Kotfresser | 1043 | Arten | Taxom | | Outenikwanus Scholtz & Ho..., 1987 | | 1 | Art | Taxom | | Oxysternon Laporte, 1758 | | 1 | Art | Taxom | | Pachylomera Griffith & P..., 1874 | | 2 | Arten | Taxom | | Pachysoma MacLeay, 1953 | | 19 | Arten | Taxom | | Panelus Lewis, 1975 | | 2 | Arten | Taxom | | Paracanthon Balthasar, 1938 | | 1 | Art | Taxom | | Paraphytus Harold, 1984 | | 6 | Arten | Taxom | | Paronitis | | 1 | Art | Taxom | | Parvuhowdenius Deschodt & C..., 2008 | | 1 | Art | Taxom | | Peckolus Scholtz & Ho..., 2008 | | 3 | Arten | Taxom | | Pedaria Laporte, 1939 | | 73 | Arten | Taxom | | Phalops Erichson, 2003 | | 36 | Arten | Taxom | | Phanaeus MacLeay, 1863 | | 9 | Arten | Taxom | | Pinacopodius Branco, 2007 | | 2 | Arten | Taxom | | Pinacotarsus Harold, 1969 | | 2 | Arten | Taxom | | Platyonitis Janssens, 1974 | | 3 | Arten | Taxom | | Pleuronitis Lansberge, 2011 | | 4 | Arten | Taxom | | Proagoderus Lansberge, 2004 | | 122 | Arten | Taxom | | Pseudocanthon Bates, 1847 | | 1 | Art | Taxom | | Pseudochironitis Ferreira, 1897 | | 3 | Arten | Taxom | | Pseudocopris Ferreira, 1910 | | 2 | Arten | Taxom | | Pseudosaproecius Balthasar, 1884 | | 12 | Arten | Taxom | | Pycnopanelus Arrow, 1913 | | 2 | Arten | Taxom | | Sarophorus Erichson, 1840 | | 12 | Arten | Taxom | | Scaptocnemis Péringuey, 1901 | | 1 | Art | Taxom | | Scarabaeus Linnaeus, 1901 | | 137 | Arten | Taxom | | Scatimus Erichson, 1862 | | 2 | Arten | Taxom | | Sceliages Westwood, 1844 | | 8 | Arten | Taxom | Sebasteos Westwood, 1845 | | 0 | Arten | Taxom | | Silvaphilus | | 2 | Arten | Taxom | | Sisyphus Latreille, 1973 | | 97 | Arten | Taxom | Sphaerocanthon Olsoufieff, 1947 | | 0 | Arten | Taxom | | Stiptocnemis Branco, 1991 | | 2 | Arten | Taxom | | Stiptopodius Harold, 1901 | | 15 | Arten | Taxom | | Stiptotarsus Branco, 1963 | | 3 | Arten | Taxom | | Strandius Balthasar, 1980 | | 1 | Art | Taxom | | Sukelus Branco, 1991 | | 1 | Art | Taxom | | Sulcodrepanus Krikken, 1840 | | 1 | Art | Taxom | | Sulcophanaeus Olsoufieff, 1856 | | 2 | Arten | Taxom | | Tanzanolus Scholtz & Ho..., 1960 | | 1 | Art | Taxom | | Tesserodon Hope, 1974 | | 1 | Art | Taxom | | Tibiodrepanus Krikken, 2011 | | 2 | Arten | Taxom | | Tiniocellus Péringuey, 1851 | | 7 | Arten | Taxom | | Tomogonus d´Orbigny, 1971 | | 11 | Arten | Taxom | | Tragiscus Klug, 1855 | | 1 | Art | Taxom | | Trichillidium Vaz-de-Mello, 1948 | | 1 | Art | Taxom | | Tropidonitis Janssens, 1857 | | 1 | Art | Taxom | | Unidentis | | 1 | Art | Taxom | | Uroxys Westwood, 1940 | | 5 | Arten | Taxom | | Versicorpus Deschodt, Da..., 2005 | | 4 | Arten | Taxom | | Walterantus Cambefort, 1960 | | 1 | Art | Taxom | | Xinidium Harold, 1993 | | 4 | Arten | Taxom |
Artenliste für Afrika
3178 Arten und 121 Unterarten | Acanthonitis bouyeri Moretto, 2010 | | | Acanthonitis nicolasi Cambefort, 1995 | | | Acanthonitis oberthuri Janssens, 1937 | | | Acanthonitis spissus Josso, 2012 | | | Afrodrepanus impressicollis Boheman, 1857 | | | Afrodrepanus marshalli Boucomont, 1921 | | | Agamopus lampros Bates, 1887 | | | Aleiantus fiorii (Paulian, 1976) | ⇒ Epactoides fiorii Paulian, 1976 | | Aleiantus incertus (Lebis, 1953) | ⇒ Epactoides incertus Lebis, 1953 | | Aleiantus perinetanus (Paulian, 1975) | ⇒ Epactoides perinetanus Paulian, 1975 | | Aleiantus perrieri (Fairmaire, 1898) | ⇒ Epactoides perrieri Fairmaire, 1898 | | Aliuscanthoniola similaris Deschodt & S..., 2008 | | | Allogymnopleurus aeneus Harold, 1875 | | | Allogymnopleurus alluaudi Garreta, 1914 | | | Allogymnopleurus anthracinus Klug, 1845 | | | Allogymnopleurus chloris Klug, 1855 | | | Allogymnopleurus consocius Péringuey, 1901 | | | Allogymnopleurus histrio Balthasar, 1963 | | | Allogymnopleurus indigaceus Reiche, 1847 | | | Allogymnopleurus infranitens Fairmaire, 1887 | | | Allogymnopleurus jeanneli Garreta, 1914 | | | Allogymnopleurus olivieri Castelnau, 1840 | | | Allogymnopleurus sagnai Nicolas & Mo..., 2002 | | | Allogymnopleurus sericeicollis Frey, 1958 | | | Allogymnopleurus signaticollis Waterhouse, 1890 | | | Allogymnopleurus splendidus Bertoloni, 1849 | | | Allogymnopleurus thalassinus Klug, 1855 | | | Allogymnopleurus umbrinus Gerstaecker, 1871 | | | Allogymnopleurus youngai Endrödi, 1973 | | | Allogymnopleurus zavattarii Müller, 1941 | | | Allonitis nasutus Felsche, 1907 | | | Alloscelus combesi Paulian, 1949 | | | Alloscelus hirticomus d´Orbigny, 1913 | | | Alloscelus longipilis Frey, 1960 | | | Alloscelus paradoxus Boucomont, 1923 | | | Amietina cambeforti Branco, 1989 | | | Amietina eburnea Cambefort, 1981 | | | Amietina larrochei Cambefort, 1981 | | | Amietina nyassalandica Branco, 1989 | | | Anachalcos aurescens Bates, 1888 | | | Anachalcos convexus Boheman, 1857 | | | Anachalcos cupreus Fabricius, 1775 | | | Anachalcos mfwangani ♰ R. Paulian, 1976 | | | Anachalcos procerus Gerstaecker, 1871 | | | Anachalcos spectabilis (Péringuey, 1901) | ⇒ Chalconotus procerus Gerstaecker, 1871 | | Anonychonitis freyi Janssens, 1950 | | | Aphengoecus clypeatus Péringuey, 1901 | | | Aphengoecus multiserratus Scholtz & Ho..., 1987 | | | Apotolamprus ambohitsitondronensis Lebis, 1953 | | | Apotolamprus andranobeensis Montreuil & ..., 2012 | | | Apotolamprus antsingyensis Montreuil, 2008 | | | Apotolamprus bemarahaensis Montreuil, 2008 | | | Apotolamprus cyanescens Brancsik, 1892 | | | Apotolamprus darainaensis Montreuil, 2008 | | | Apotolamprus densepunctatus Montreuil, 2008 | | | Apotolamprus hagai Montreuil & ..., 2012 | | | Apotolamprus hanskii Montreuil, 2005 | | | Apotolamprus helenae Montreuil, 2004 | | | Apotolamprus latipennis Boucomont, 1937 | | | Apotolamprus mahajangaensis Montreuil, 2008 | | | Apotolamprus manjarivoloensis Montreuil, 2008 | | | Apotolamprus manomboensis Montreuil, 2008 | | | Apotolamprus marojejyensis Montreuil, 2005 | | | Apotolamprus metallicus Montreuil, 2008 | | | Apotolamprus milloti Lebis, 1953 | | | Apotolamprus nanoides Paulian, 1976 | | | Apotolamprus orangeaensis Montreuil, 2008 | | | Apotolamprus perinetensis Montreuil, 2008 | | | Apotolamprus peyrierasi Paulian, 1985 | | | Apotolamprus pseudomanomboensis Montreuil, 2008 | | | Apotolamprus pseudoquadrimaculatus Montreuil & ..., 2012 | | | Apotolamprus quadrimaculatus Lebis, 1953 | | | Apotolamprus quadrinotatus Boucomont, 1937 | | | Apotolamprus resplendens Montreuil, 2008 | | | Apotolamprus sahatezaensis Montreuil & ..., 2012 | | | Apotolamprus sericeus Paulian, 1976 | | | Apotolamprus sinuatotibialis Montreuil, 2008 | | | Apotolamprus sparsepunctatus Montreuil, 2008 | | | Apotolamprus vadoni Paulian, 1976 | | | Apotolamprus zombitsyensis Montreuil, 2008 | | | Apterepilissus analavelonae Paulian, 1975 | | | Apterepilissus centralis Paulian, 1976 | | | Apterepilissus ovalis Felsche, 1911 | | | Apterepilissus sericeus Lebis, 1958 | | | Aptychonitis anomalus Gestro, 1895 | | | Aptychonitis sulcicollis Harold, 1868 | | | Arachnodes abadiei Lebis, 1953 | | | Arachnodes aeneoviridis Boucomont, 1937 | | | Arachnodes ampasindavae Paulian, 1976 | | | Arachnodes andriai Paulian, 1976 | | | Arachnodes angulosus Boucomont, 1937 | | | Arachnodes bicolor Montreuil, 2011 | | | Arachnodes biimpressus Lebis, 1953 | | | Arachnodes brunnipes Castelnau, 1840 | | | Arachnodes delaunayi Lebis, 1953 | | | Arachnodes delphinensis Lebis, 1953 | | | Arachnodes dichrous Paulian, 1976 | | | Arachnodes didiensis Lebis, 1953 | | | Arachnodes dieganus Paulian, 1976 | | | Arachnodes fairmairei Boucomont, 1937 | | | Arachnodes fulgens Kunckel, 1887 | | | Arachnodes fusconitens Fairmaire, 1899 | | | Arachnodes fuscovirens Fairmaire, 1889 | | | Arachnodes globuloides Paulian, 1976 | | | Arachnodes globulosus (Kunckel, 1887) | ⇒ Arachnodes globuloides Paulian, 1976 | | Arachnodes grossepunctatus Paulian, 1986 | | | Arachnodes hanskii Montreuil, 2003 | | | Arachnodes histeroides Castelnau, 1840 | | | Arachnodes insularis Paulian, 1976 | | | Arachnodes jeanneli Lebis, 1953 | | | Arachnodes kelifelyi Paulian, 1976 | | | Arachnodes luctuosus Boucomont, 1937 | | | Arachnodes mahafalyensis Montreuil, 2006 | | | Arachnodes manaitrai Montreuil, 2006 | | | Arachnodes manaitrai kirindyensis Montreuil, 2006 | | | Arachnodes manaitrai manaitrai | | | Arachnodes manomboensis Montreuil, 2006 | | | Arachnodes mantillerii Montreuil, 2003 | | | Arachnodes micans Lebis, 1953 | | | Arachnodes minutus Paulian, 1976 | | | Arachnodes morati Paulian, 1975 | | | Arachnodes morio Harold, 1879 | | | Arachnodes neonitidus Montreuil, 2011 | | | Arachnodes nitidus Castelnau, 1840 | | | Arachnodes nosybeensis Lebis, 1961 | | | Arachnodes oberthuri Boucomont, 1937 | | | Arachnodes philippi Lebis, 1953 | | | Arachnodes pierrettae Lebis, 1953 | | | Arachnodes pillula Paulian, 1976 | | | Arachnodes pusillus Lebis, 1953 | | | Arachnodes refulgens Fairmaire, 1901 | | | Arachnodes renaudi Lebis, 1958 | | | Arachnodes robinsoni Boucomont, 1937 | | | Arachnodes rubrolimbatus Boucomont, 1937 | | | Arachnodes rubrotinctus Lebis, 1953 | | | Arachnodes sakarahae Paulian, 1976 | | | Arachnodes saprinoides Fairmaire, 1889 | | | Arachnodes semichalceus Lebis, 1953 | | | Arachnodes seminitidus Lebis, 1953 | | | Arachnodes seyrigi Lebis, 1953 | | | Arachnodes soganus Paulian, 1976 | | | Arachnodes splendidus Lebis, 1953 | | | Arachnodes tibialis Lebis, 1956 | | | Arachnodes variolosus Lebis, 1953 | | | Arachnodes vicinus Castelnau, 1840 | | | Arachnodes viettei Montreuil, 2011 | | | Arachnodes vieui Paulian, 1976 | | | Arachnodes vigilans Lebis, 1953 | | | Arachnodes zombitsyensis Montreuil, 2011 | | | Ateuchus earthorus Kohlmann & S..., 2009 | | | Ateuchus fetteri Kohlmann, 1996 | | | Ateuchus hendrichsi Kohlmann, 1996 | | | Ateuchus pius (Illiger, 1803) | ⇒ Scarabaeus pius Illiger, 1803 | | Ateuchus rodriguezi Preudhomme d..., 1886 | | | Ateuchus spinipes Thunberg, 1818 | | | Ateuchus zoebischi Kohlmann, 1996 | | | Bohepilissus nitidus Balthasar, 1965 | | | Bohepilissus subtilis Boheman, 1857 | | | Bubas bison Linnaeus, 1767 | | | Bubas bubaloides Janssens, 1938 | | | Bubas bubalus Olivier, 1811 | | | Byrrhidium convexum Scholtz & Ho..., 1987 | | | Byrrhidium namaquensis Scholtz & Ho..., 1987 | | | Byrrhidium ovale Harold, 1869 | | | Caccobius anthracites d´Orbigny, 1904 | | | Caccobius atomarius (Balthasar, 1935) | ⇒ Caccobius brevis Waterhouse, 1875 | | Caccobius atratus Walter & Cam..., 1977 | | | Caccobius auberti d´Orbigny, 1902 | | | Caccobius bama Walter, 2014 | | | Caccobius brevis Waterhouse, 1875 | | | Caccobius cabellai Walter, 2019 | | | Caccobius callosifrons D Orbigny, 1905 | | | Caccobius castaneus (Klug, 1855) | ⇒ Caccocnemus castaneus Klug, 1855 | | Caccobius cavatus d´Orbigny, 1908 | | | Caccobius convexifrons (Raffray, 1877) | ⇒ Caccophilus convexifrons Raffray, 1877 | | Caccobius corniceps d´Orbigny, 1913 | | | Caccobius cribrarius Boucomont, 1928 | | | Caccobius croceocinctus d´Orbigny, 1913 | | | Caccobius curvicornis Walter & Cam..., 1977 | | | Caccobius cuspidiger d´Orbigny, 1913 | | | Caccobius cyclotis Cambefort, 1984 | | | Caccobius discrepans Péringuey, 1901 | | | Caccobius dorsalis (Harold, 1867) | ⇒ Caccophilus dorsalis Harold, 1867 | | Caccobius dybowskii d´Orbigny, 1902 | | | Caccobius elephantinus Balthasar, 1967 | | | Caccobius ferrugineus Fahraeus, 1857 | | | Caccobius foveolatus Frey, 1955 | | | Caccobius fuliginosus (Roth, 1851) | ⇒ Caccophilus reticuliger d´Orbigny, 1904 | | Caccobius gananensis d´Orbigny, 1904 | | | Caccobius genierorum Walter, 2014 | | | Caccobius gibbosulus d´Orbigny, 1915 | | | Caccobius globaticeps d´Orbigny, 1905 | | | Caccobius grossegranosus Balthasar, 1963 | | | Caccobius hirsutus Frey, 1958 | | | Caccobius histerinus Fahraeus, 1857 | | | Caccobius histeroides Ménétriés, 1832 | | | Caccobius histrio Balthasar, 1967 | | | Caccobius inconspicuus (Fahraeus, 1857) | ⇒ Caccophilus fuliginosus Roth, 1851 | | Caccobius inops Péringuey, 1901 | | | Caccobius ivorensis Cambefort, 1984 | | | Caccobius jossoi Walter, 2014 | | | Caccobius krikkeni Cambefort, 1980 | | | Caccobius lamottei (Cambefort, 1983) | ⇒ Cleptocaccobius lamottei Cambefort, 1983 | | Caccobius lateralis d´Orbigny, 1905 | | | Caccobius leleupi Balthasar, 1963 | | | Caccobius longipennis d´Orbigny, 1904 | | | Caccobius megaponerae (Brauns, 1914) | ⇒ Megaponerophilus megaponerae Brauns, 1914 | | Caccobius meridionalis Boucomont, 1914 | | | Caccobius minor (Walter & Cam..., 1977) | ⇒ Cleptocaccobius minor Walter & Cam..., 1977 | | Caccobius mirabilepunctatus Cambefort, 1971 | | | Caccobius montreuilli Walter, 2019 | | | Caccobius morettoi Walter, 2014 | | | Caccobius nigritulus Klug, 1855 | | | Caccobius obtusus Fahraeus, 1857 | | | Caccobius ocellipennis d´Orbigny, 1913 | | | Caccobius pentagonus d´Orbigny, 1908 | | | Caccobius pilosus Frey, 1958 | | | Caccobius polygonus d´Orbigny, 1911 | | | Caccobius postlulatus | | | Caccobius postlutatus d´Orbigny, 1905 | | | Caccobius pseudocorniceps Josso, 2018 | | | Caccobius pseudolaevis d´Orbigny, 1908 | | | Caccobius punctatissimus Harold, 1867 | | | Caccobius reticuliger d´Orbigny, 1904 | | | Caccobius schreberi Linnaeus, 1767 | | | Caccobius semiaeneus d´Orbigny, 1905 | | | Caccobius semiluteus d´Orbigny, 1905 | | | Caccobius sericeus Frey, 1958 | | | Caccobius sericoides Balthasar, 1966 | | | Caccobius setifer d´Orbigny, 1905 | | | Caccobius signatipennis Harold, 1867 | | | Caccobius tai Cambefort, 1984 | | | Caccobius tuberifrons d´Orbigny, 1902 | | | Caccobius ugandicus Frey, 1975 | | | Caccobius upembanus Frey, 1958 | | | Caccobius villiersi Cambefort, 1975 | | | Caccobius zairensis Walter & Cam..., 1977 | | | Caccobius zambiensis Walter, 2014 | | | Cambefortantus ankaratrae Paulian, 1975 | | | Cambefortantus blanci Paulian, 1975 | | | Cambefortantus helenae Montreuil, 2008 | | | Cambefortantus micros Paulian, 1986 | | | Cambefortantus myops Lebis, 1953 | | | Cambefortantus pauliani Montreuil, 2008 | | | Cambefortantus ranomafanensis Montreuil, 2008 | | | Cambefortantus ranomandryi Paulian, 1975 | | | Cambefortius bidentiger d´Orbigny, 1905 | | | Cambefortius cavipectus Branco, 1989 | | | Cambefortius dudleyi Branco, 1989 | | | Cambefortius eburneus Branco, 1989 | | | Cambefortius guineensis Branco, 1989 | | | Cambefortius lamizanai Cambefort, 1983 | | | Cambefortius teke Walter & Cam..., 1977 | | | Cambefortius turneri Boucomont, 1936 | | | Canthidium (Eucanthidium) laetum (Harold, 1867) | ⇒ Eucanthidium laetum Harold, 1867 | | Canthidium angusticeps Bates, 1887 | | | Canthidium annagabrielae Solis & Kohl..., 2004 | | | Canthidium ardens Bates, 1887 | | | Canthidium centrale Boucomont, 1928 | | | Canthidium discopygidiale Howden & Young, 1981 | | | Canthidium gerstaeckeri Harold, 1867 | | | Canthidium guanacaste Howden & Gill, 1987 | | | Canthidium haroldi Preudhomme d..., 1886 | | | Canthidium hespenheidei Howden & Young, 1981 | | | Canthidium manni Arrow, 1913 | | | Canthidium marianelae Solis & Kohl..., 2004 | | | Canthidium muticum Boheman, 1858 | | | Canthidium perceptibile Howden & Young, 1981 | | | Canthidium smaragdinum Harold, 1867 | | | Canthidium stofeli Carvalho de ..., 2019 | | | Canthidium tuberifrons Howden & Young, 1981 | | | Canthidium vespertinum Howden & Young, 1981 | | | Canthochilum magnum Philips & Ivie, 2008 | | | Canthodimorpha lawrencei Davis, Schol..., 1999 | | | Canthon (Boreocanthon) praticola Le Conte, 1859 | | | Canthon (Canthon) cyanellus LeConte, 1859 | | | Canthon (Canthon) indigaceus (LeConte, 1866) | ⇒ Canthon humectus Say, 1831 | | Canthon angustatus Harold, 1867 | | | Canthon deyrollei Harold, 1868 | | | Canthon dives Harold, 1868 | | | Canthon euryscelis Bates, 1887 | | | Canthon hartmanni Howden & Gill, 1987 | | | Canthon meridionalis Martinez, Ha..., 1964 | | | Canthon morsei Howden, 1966 | | | Canthon rutilans (Castelnau, 1840) | ⇒ Francmonrosia rutilans Castelnau, 1840 | | Canthon staigi Pereira, 1953 | | | Canthon subhyalinus Harold, 1867 | | | Canthon vazquezae Martinez, Ha..., 1964 | | | Catharsiocopris quadrituberculatus Balthasar, 1967 | | | Catharsius achates Olivier, 1789 | | | Catharsius adamastor Gillet, 1932 | | | Catharsius aegeus Génier, 2017 | | | Catharsius africanus Ferreira, 1960 | | | Catharsius alpheus Gillet, 1932 | | | Catharsius approximans Kolbe, 1893 | | | Catharsius balubanus Kolbe, 1893 | | | Catharsius bellus Gillet, 1918 | | | Catharsius belus Gillet, 1918 | | | Catharsius biconifer Felsche, 1907 | | | Catharsius bicornutus Kolbe, 1893 | | | Catharsius bradshawi Lansberge, 1887 | | | Catharsius brevicornis Felsche, 1907 | | | Catharsius brittoni Ferreira, 1963 | | | Catharsius brutus Harold, 1880 | | | Catharsius calaharicus Kolbe, 1893 | | | Catharsius cassius Kolbe, 1893 | | | Catharsius chinai Ferreira, 1960 | | | Catharsius congolensis Ferreira, 1962 | | | Catharsius coptorhinaformis Josso & Prév..., 2009 | | | Catharsius crassicornis Gillet, 1911 | | | Catharsius disseptus (Péringuey, 1901) | ⇒ Catharsius calaharicus Kolbe, 1893 | | Catharsius dubius (Péringuey, 1901) | ⇒ Catharsius molossus Linnaeus, 1758 | | Catharsius duciformis Ferreira, 1959 | | | Catharsius dux Harold, 1878 | | | Catharsius erechtheus Felsche, 1907 | | | Catharsius eteocles (Laporte, 1840) | ⇒ Catharsius alpheus Gillet, 1932 | | Catharsius fastidiosus Thomson, 1858 | | | Catharsius folicornis Josso & Prév..., 2019 | | | Catharsius furcillatus Guérin-Ménev..., 1847 | | | Catharsius furcillatus furcillatus Guérin, 1847 | | | Catharsius furcillatus laeviplaga G. Müller, 1941 | | | Catharsius garambae Ferreira, 1962 | | | Catharsius genieri Moretto, 2014 | | | Catharsius gorilla Thomson, 1858 | | | Catharsius gorilloides Felsche, 1907 | | | Catharsius guineensis Ferreira, 1963 | | | Catharsius haafi Ferreira, 1960 | | | Catharsius haroldi Takano, 2021 | | | Catharsius harpagus Harold, 1877 | | | Catharsius heros Boheman, 1860 | | | Catharsius hertli Ferreira, 1964 | | | Catharsius jacksoni Waterhouse, 1891 | | | Catharsius juheli Josso, 2011 | | | Catharsius karlwerneri Moretto, 2008 | | | Catharsius kolbei Felsche, 1907 | | | Catharsius laticeps Boheman, 1857 | | | Catharsius latifossa Müller, 1941 | | | Catharsius lepineyi Boucomont, 1934 | | | Catharsius longiceps Gillet, 1907 | | | Catharsius luluensis Ferreira, 1960 | | | Catharsius lycaon Kolbe, 1893 | | | Catharsius machadoi Ferreira, 1960 | | | Catharsius marcellus Kolbe, 1893 | | | Catharsius melancholicus (Boheman, 1860) | ⇒ Catharsius longiceps Gillet, 1907 | | Catharsius minutus (Lansberge, 1882) | ⇒ Metacatharsius minutus Lansberge, 1882 | | Catharsius mirabilis Felsche, 1901 | | | Catharsius molossus Linnaeus, 1758 | Dickkopf | | Catharsius morettoi Josso & Prév..., 2009 | | | Catharsius mossambicanus Ferreira, 1960 | | | Catharsius mulanjeensis Josso & Prév..., 2019 | | | Catharsius neptunus Kolbe, 1893 | | | Catharsius ninus Gillet, 1918 | | | Catharsius oedipus Felsche, 1907 | | | Catharsius omoensis Müller, 1941 | | | Catharsius oryx Frey, 1974 | | | Catharsius pallas Génier, 2017 | | | Catharsius pandion Harold, 1877 | | | Catharsius parafastidiosus Ferreira, 1971 | | | Catharsius peleus (Olivier, 1789) | ⇒ Metacatharsius peleus Olivier, 1789 | | Catharsius phidias (Olivier, 1789) | ⇒ Catharsius platicornis Ferreira, 1960 | | Catharsius philus Kolbe, 1893 | | | Catharsius pithecius Fabricius, 1775 | Indianische Schulteraale | | Catharsius platicornis Ferreira, 1960 | | | Catharsius platycerus Klug, 1855 | | | Catharsius platynotus Felsche, 1907 | | | Catharsius polynices Kolbe, 1893 | | | Catharsius pseudocongolensis Ferreira, 1963 | | | Catharsius pseudolycaon Ferreira, 1960 | | | Catharsius pseudooedipus Ferreira, 1960 | | | Catharsius pseudoplaticornis Josso & Prév..., 2019 | | | Catharsius pseudovitulus Müller, 1941 | | | Catharsius punctatus Josso & Prév..., 2019 | | | Catharsius rhinoceros Klug, 1855 | | | Catharsius saegeri Ferreira, 1962 | | | Catharsius satyrus Kolbe, 1893 | | | Catharsius scopas Felsche, 1907 | | | Catharsius semirubidus Fairmaire, 1891 | | | Catharsius sesostris Waterhouse, 1888 | | | Catharsius severini Gillet, 1907 | | | Catharsius simillimus Ferreira, 1960 | | | Catharsius somalicus Gillet, 1918 | | | Catharsius spectabilis Gillet, 1918 | | | Catharsius straeleni Ferreira, 1963 | | | Catharsius stricticornis Josso & Prév..., 2019 | | | Catharsius stuhlmanni Kolbe, 1893 | | | Catharsius takanoi Josso & Prév..., 2019 | | | Catharsius tricornutus DeGeer, 1778 | | | Catharsius tridens Josso & Prév..., 2009 | | | Catharsius ugandicus Gillet, 1918 | | | Catharsius ulysses Boheman, 1857 | | | Catharsius upembanus Ferreira, 1962 | | | Catharsius vansoni Ferreira, 1960 | | | Catharsius vitulus Boheman, 1857 | | | Cheironitis arrowi Janssens, 1937 | | | Cheironitis asbenicus Gillet, 1909 | | | Cheironitis audens Péringuey, 1901 | | | Cheironitis dentifemoralis Ferreira, 1976 | | | Cheironitis flabellatus Boucomont, 1923 | | | Cheironitis furcifer Rossi, 1792 | | | Cheironitis hoplosternus Harold, 1868 | | | Cheironitis hungaricus Herbst, 1789 | | | Cheironitis imitator Balthasar, 1963 | | | Cheironitis indicus (Lansberge, 1875) | ⇒ Cheironitis arrowi Janssens, 1937 | | Cheironitis irroratus Rossi, 1790 | | | Cheironitis muelleri Janssens, 1943 | | | Cheironitis osiridis Reiche, 1856 | | | Cheironitis scabrosus Fabricius, 1776 | | | Cheironitis socotranus Gahan, 1900 | | | Cheironitis ungaricus Herbst, 1789 | | | Cheironitis ungaricus ungaricus Herbst, 1789 | | | Cheironitis viridicans Gillet, 1918 | | | Chironitis furcifer Rossi, 1792 | | | Chironitis irroratus Rossi, 1790 | | | Chironitis pamphilus Ménétriése, 1849 | | | Circellium bacchus Fabricius, 1781 | Riesen-Pillendreher | | Cleptocaccobius balthasarianus Cambefort, 1979 | | | Cleptocaccobius biceps d´Orbigny, 1905 | | | Cleptocaccobius clementi Cambefort, 1971 | | | Cleptocaccobius convexifrons Raffray, 1877 | | | Cleptocaccobius dorbignyi Cambefort, 1984 | | | Cleptocaccobius humilis d´Orbigny, 1902 | | | Cleptocaccobius lamottei Cambefort, 1983 | | | Cleptocaccobius minor Walter & Cam..., 1977 | | | Cleptocaccobius morettoi Josso & Prév..., 2006 | | | Cleptocaccobius postlutatus d´Orbigny, 1905 | | | Cleptocaccobius schaedlei d´Orbigny, 1902 | | | Cleptocaccobius semiluteus d´Orbigny, 1905 | | | Cleptocaccobius seminulum Klug, 1855 | | | Cleptocaccobius signaticollis d´Orbigny, 1902 | | | Cleptocaccobius uniseries d´Orbigny, 1905 | | | Cleptocaccobius viridicollis Fahraeus, 1857 | | | Cleptocaccobius youngai Balthasar, 1967 | | | Clypeodrepanus digitatus Krikken, 2009 | | | Clypeodrepanus striatus Boucomont, 1921 | | | Clypeodrepanus strigatus Janssens, 1953 | | | Copridaspidus peregrinus Harold, 1878 | | | Copris achates (Péringuey, 1901) | ⇒ Copris cornifrons Boheman, 1860 | | Copris afgoi Cambefort, 1992 | | | Copris algol Nguyen-phung, 1989 | | | Copris amabilis Kolbe, 1914 | | | Copris amyntor (Klug, 1855) | ⇒ Copris harrisi Waterhouse, 1891 | | Copris anceus Olivier, 1789 | | | Copris angustus Nguyen-phung..., 1986 | | | Copris antares Bacescu & Ca..., 1947 | | | Copris arcturus Gillet, 1907 | | | Copris ares Zelenka, 1993 | | | Copris armiger Gillet, 1910 | | | Copris atropolitus Gillet, 1910 | | | Copris basilewskyi Ferreira, 1962 | | | Copris bidens (Pfeffer, 1888) | ⇒ Liatongus femoratus Illiger, 1800 | | Copris bihamatus Balthasar, 1965 | | | Copris binodis Thunberg, 1818 | | | Copris bootes Klug, 1855 | | | Copris bovinus Gillet, 1908 | | | Copris brittoni Ferreira, 1962 | | | Copris caelatus Fabricius, 1794 | | | Copris cambeforti Nguyen-phung, 1988 | | | Copris camerunus Felsche, 1904 | | | Copris capensis Waterhouse, 1891 | | | Copris carmelita Fabricius, 1801 | | | Copris cassius Péringuey, 1901 | | | Copris ciliatus | | | Copris colmanti Gillet, 1908 | | | Copris complexus Nguyen-phung, 1989 | | | Copris coriarius Gillet, 1907 | | | Copris cornifrons Boheman, 1860 | | | Copris corniger (Sahlberg, 1823) | ⇒ Onthophagus taurus Schreber, 1759 | | Copris costaricensis Gahan, 1894 | | | Copris crassus Deschodt & D..., 2015 | | | Copris cribricollis (Gillet, 1910) | ⇒ Catharsius pithecius Fabricius, 1775 | | Copris curlettii Moretto, 2011 | | | Copris davisi Nguyen-phung..., 1986 | | | Copris decellei Cambefort, 1992 | | | Copris denticulatus Nguyen-phung, 1988 | | | Copris diversus Waterhouse, 1891 | | | Copris draco Arrow, 1906 | | | Copris dracunculus Fereira, 1959 | | | Copris dudleyi Cambefort, 1992 | | | Copris eburneus Cambefort, 1992 | | | Copris elpheneroides Felsche, 1910 | | | Copris elphenor Klug, 1855 | | | Copris evanidus Klug, 1855 | | | Copris fallaciosus Gillet, 1907 | | | Copris fallax Felsche, 1910 | | | Copris fidius Olivier, 1789 | | | Copris freyi Ferreira, 1961 | | | Copris garambae Cambefort, 1992 | | | Copris gazellarum Gillet, 1918 | | | Copris gilleti Kolbe, 1907 | | | Copris gladiator Arrow, 1933 | | | Copris gracilis Waterhouse, 1891 | | | Copris graueri (Kolbe, 1914) | ⇒ Copris camerunus Felsche, 1904 | | Copris harrisi Waterhouse, 1891 | | | Copris hartli Frey, 1975 | | | Copris hispanus Linnaeus, 1764 | Schwarzglänzende Spanische Nashornkäfer | | Copris hispanus cavolinii Petagna, 1792 | | | Copris hispanus hispanus Linné, 1764 | | | Copris humilis Gillet, 1908 | | | Copris hybridus Gillet, 1910 | | | Copris incertus Say, 1835 | | | Copris inhalatus Quedenfeldt, 1884 | | | Copris inhalatus grandis Nguyen-phung..., 1986 | | | Copris inhalatus inhalatus | | | Copris inhalatus ngotto Moretto, 2011 | | | Copris inhalatus perturbator Péringuey, 1901 | | | Copris inhalatus sanctaeluciae Nguyen-phung..., 1986 | | | Copris insidiosus Péringuey, 1901 | | | Copris integer (Reiche, 1847) | ⇒ Copris fallaciosus Gillet, 1907 | | Copris interioris Kolbe, 1897 | | | Copris jacchoides Nguyen-phung..., 1987 | | | Copris jacchus Lang, 1971 | | | Copris jahi Nguyen-phung..., 1986 | | | Copris jucundus Gillet, 1932 | | | Copris kasanka Josso & Prév..., 2016 | | | Copris katangae Cambefort & ..., 1996 | | | Copris laiiformis Nguyen-phung..., 1987 | | | Copris laioides Boucomont, 1932 | | | Copris laius Harold, 1868 | | | Copris laius angusticornis Nguyen-phung..., 1987 | | | Copris laius laius | | | Copris leakeyorum ♰ R. Paulian, 1976 | | | Copris leruei Josso & Prév..., 2017 | | | Copris lugubris Boheman, 1858 | | | Copris macer Péringuey, 1901 | | | Copris martinae Nguyen-phung, 1988 | | | Copris martinae martinae | | | Copris martinae ntchisi Moretto, 2011 | | | Copris megaceratoides Waterhouse, 1891 | | | Copris megasoma Matthews & H..., 1959 | | | Copris mesacanthus Harold, 1878 | | | Copris mesacanthus mesacanthus | | | Copris mesacanthus transvaalensis Nguyen-phung, 1988 | | | Copris misellus Péringuey, 1901 | | | Copris moffartsi Gillet, 1907 | | | Copris montivagus (Gillet, 1908) | ⇒ Copris harrisi Waterhouse, 1891 | | Copris morettoi Josso & Prév..., 2020 | | | Copris morphaeus Gillet, 1932 | | | Copris mourgliai Zelenka, 1993 | | | Copris muticus (Boheman, 1857) | ⇒ Litocopris muticus Boheman, 1857 | | Copris neotridens Génier & Pré..., 2017 | | | Copris nepos Gillet, 1908 | | | Copris obesus Boheman, 1857 | | | Copris orion Klug, 1835 | | | Copris orion caffer Gillet, 1908 | | | Copris orion centralis Gillet, 1908 | | | Copris orion orion | | | Copris orion overlaeti Balthasar, 1961 | | | Copris orphanus (Guérin-Ménev..., 1847) | ⇒ Copris evanidus Klug, 1855 | | Copris pauliani Nguyen-phung..., 1986 | | | Copris perturbator (Péringuey, 1901) | ⇒ Copris inhalatus perturbator Péringuey, 1901 | | Copris phungae Cambefort, 1992 | | | Copris phungae gabonicus Cambefort, 1992 | | | Copris phungae phungae | | | Copris phylax Gillet, 1908 | | | Copris prevosti Josso, 2011 | | | Copris pseudobootes Ferreira, 1962 | | | Copris pseudomoffartsi Moretto, 2011 | | | Copris pueli Mollandin De..., 1905 | | | Copris puncticollis Boheman, 1857 | | | Copris punctiventris Waterhouse, 1891 | | | Copris renwarti Nguyen-phung..., 1986 | | | Copris ritsemae Harold, 1875 | | | Copris ruandanus Balthasar, 1961 | | | Copris rugosus (Gillet, 1908) | ⇒ Copris inhalatus Quedenfeldt, 1884 | | Copris sarpedon Harold, 1868 | | | Copris scorpio Zelenka, 1993 | | | Copris serius Nguyen-phung, 1987 | | | Copris sexdentatus (Thunberg, 1818) | ⇒ Copris sarpedon Harold, 1868 | | Copris simonettai Cambefort, 1992 | | | Copris simplex (Harold, 1869) | ⇒ Dichotomius simplex Taschenberg, 1870 | | Copris simulator Nguyen-phung..., 1986 | | | Copris singularis Nguyen-phung..., 1986 | | | Copris sinon Olivier, 1789 | | | Copris sphaeropterus Harold, 1877 | | | Copris subsidens Péringuey, 1901 | | | Copris tridens (Felsche, 1901) | ⇒ Copris hispanus hispanus Linné, 1764 | | Copris truncatus Felsche, 1901 | | | Copris typhoeus Gerstaecker, 1884 | | | Copris urus (Boheman, 1857) | ⇒ Catharsius molossus Linnaeus, 1758 | | Copris usambaricus Gillet, 1908 | | | Copris vankhaii Nguyen-phung, 1988 | | | Copris vankhaii genopunctatus Nguyen-phung, 1988 | | | Copris vankhaii vankhaii | | | Copris vankhalii Nguyen-phung, 1988 | | | Copris victorini (Boheman, 1857) | ⇒ Copris ritsemae Harold, 1875 | | Copris vilhenai Ferreira, 1962 | | | Copris vrydaghi Ferreira, 1962 | | | Copris wiesei Kolbe, 1914 | | | Copris youngai Balthasar, 1967 | | | Coprophanaeus (Megaphanaeus) bellicosus Olivier, 1789 | | | Coprophanaeus boucardi Nevinson, 1891 | | | Coprophanaeus kohlmanni (Arnaud, 2002) | ⇒ Coprophanaeus morenoi Arnaud, 1982 | | Coprophanaeus telamon Erichson, 1847 | | | Coprophanaeus uhleri (Maly & Pokorny, 2008) | ⇒ Coprophanaeus chiriquensis Olsufieff, 1924 | | Coptodactyla subaenea Harold, 1877 | | | Coptorhina angolensis Arrow, 1906 | | | Coptorhina auspicata Péringuey, 1901 | | | Coptorhina davidi Janssens, 1939 | | | Coptorhina excavata Frolov, Akhm..., 2008 | | | Coptorhina klugi Hope, 1833 | | | Coptorhina klugii Hope, 1833 | | | Coptorhina nanus Péringuey, 1888 | | | Coptorhina nitefacta Gillet, 1932 | | | Coptorhina nitidipennis Boheman, 1857 | | | Cyptochirus ambiguus (Kirby, 1828) | ⇒ Drepanocerus trogiformis Roth, 1851 | | Cyptochirus decellei Simonis & Zu..., 1980 | | | Cyptochirus distinctus Janssens, 1953 | | | Cyptochirus trogiformis Roth, 1851 | | | Delopleurus darrenmanni Frolov, 2014 | | | Delopleurus fossatus Frolov, 2014 | | | Delopleurus giletti Janssens, 1939 | | | Delopleurus gilletti Janssens, 1939 | | | Delopleurus krikkeni Frolov, 2014 | | | Delopleurus mencli Kral, 2014 | | | Delopleurus naviauxi Frolov & Cam..., 2014 | | | Delopleurus pubescens Frolov, 2014 | | | Delopleurus pullus Boheman, 1857 | | | Deltochilum enceladus Kolbe, 1893 | | | Deltochilum lobipes Bates, 1887 | | | Deltochilum mexicanum Burmeister, 1848 | | | Deltochilum parile Bates, 1887 | | | Deltochilum pseudoparile Paulian, 1938 | | | Diastellopalpus anthonyi Walter, 1989 | | | Diastellopalpus astorgisorum Walter, 2013 | | | Diastellopalpus balubanus Kolbe, 1893 | | | Diastellopalpus basilobatus d´Orbigny, 1902 | | | Diastellopalpus cambeforti Josso & Pré..., 2000 | | | Diastellopalpus camiadei Moretto, 1999 | | | Diastellopalpus conradti d´Orbigny, 1902 | | | Diastellopalpus drumonti Moretto, 1999 | | | Diastellopalpus dudleyi Cambefort, 1980 | | | Diastellopalpus fuelleborni Kolbe, 1901 | | | Diastellopalpus gilleti d´Orbigny, 1908 | | | Diastellopalpus hassoni Moretto, 1999 | | | Diastellopalpus ikelenge Josso & Pré..., 2014 | | | Diastellopalpus infernalis Lansberge, 1887 | | | Diastellopalpus jirouxi Josso & Pré..., 2000 | | | Diastellopalpus johnstoni Waterhouse, 1885 | | | Diastellopalpus laevibasis d´Orbigny, 1902 | | | Diastellopalpus lamellicollis Quedenfeldt, 1884 | | | Diastellopalpus legrandi Josso & Pré..., 2000 | | | Diastellopalpus metriogonus (d´Orbigny, 1914) | ⇒ Diastellopalpus neavei d´Orbigny, 1914 | | Diastellopalpus monapoides Bates, 1888 | | | Diastellopalpus murrayi Harold, 1868 | | | Diastellopalpus neavei d´Orbigny, 1914 | | | Diastellopalpus nigerrimus Kolbe, 1893 | | | Diastellopalpus noctis Thomson, 1858 | | | Diastellopalpus pluton d´Orbigny, 1902 | | | Diastellopalpus pseudoschultzei Josso & Pré..., 2000 | | | Diastellopalpus quinquedens Bates, 1888 | | | Diastellopalpus schultzei Gillet, 1927 | | | Diastellopalpus semirubidus d´Orbigny, 1913 | | | Diastellopalpus sulciger Kolbe, 1893 | | | Diastellopalpus thomsoni Bates, 1888 | | | Diastellopalpus thomsoni malawiensis Josso & Pré..., 2000 | | | Diastellopalpus thomsoni thomsoni | | | Diastellopalpus tridens (Fairmaire, 1891) | ⇒ Diastellopalpus thomsoni Bates, 1888 | | Diastellopalpus walteri Josso & Pré..., 2000 | | | Dichotomius (Luederwaldtinia) centralis Harold, 1869 | | | Dichotomius (Luederwaldtinia) guaribensis Valois, Vaz-..., 2017 | | | Dichotomius (Luederwaldtinia) yucatanus Bates, 1887 | | | Dichotomius (Selenocopris) simplex (Taschenberg, 1870) | ⇒ Selenocopris simplex Taschenberg, 1870 | | Dichotomius annae Linnaeus, 1767 | | | Dichotomius satanas Harold, 1867 | | | Dicranocara deschodti Frolov & Sch..., 2003 | | | Dicranocara inexpectata Deschodt & S..., 2007 | | | Dicranocara tatasensis Deschodt & S..., 2007 | | | Dicranocara vandersmisseni Moretto, 2016 | | | Digitonthophagus aksumensis Génier, 2017 | | | Digitonthophagus biflagellatus Génier, 2017 | | | Digitonthophagus catta Fabricius, 1787 | | | Digitonthophagus dilatatus Génier, 2017 | | | Digitonthophagus eucatta Génier, 2017 | | | Digitonthophagus falciger Génier, 2017 | | | Digitonthophagus fimator Génier, 2017 | | | Digitonthophagus gazella Fabricius, 1787 | | | Digitonthophagus lusinganus Orbigny, 1905 | | | Digitonthophagus namaquensis Génier, 2017 | | | Digitonthophagus petilus Génier, 2017 | | | Digitonthophagus sahelicus Moretto, 2017 | | | Digitonthophagus ulcerosus Génier, 2017 | | | Digitonthophagus viridicollis Génier, 2017 | | | Disphysema candezei Harold, 1873 | | | Dorbignyolus mirecribratus d´Orbigny, 1913 | | | Drepanocerus abyssinicus (Roth, 1851) | ⇒ Ixodina abyssinica Roth, 1851 | | Drepanocerus ambiguus Kirby, 1828 | | | Drepanocerus bechynei (Janssens, 1953) | ⇒ Eodrepanus bechynei Janssens, 1953 | | Drepanocerus endroedy Endrodi, 1976 | | | Drepanocerus fastiditus (Péringuey, 1901) | ⇒ Eodrepanus fastiditus Péringuey, 1901 | | Drepanocerus impressicollis (Boheman, 1857) | ⇒ Afrodrepanus impressicollis Boheman, 1857 | | Drepanocerus kirbyi Kirby, 1828 | | | Drepanocerus laticollis (Fahraeus, 1857) | ⇒ Latodrepanus laticollis Fahraeus, 1857 | | Drepanocerus marshalli (Boucomont, 1921) | ⇒ Afrodrepanus marshalli Boucomont, 1921 | | Drepanocerus orientalis Krikken, 2009 | | | Drepanocerus parallelus (Raffray, 1877) | ⇒ Eodrepanus parallelus Raffray, 1877 | | Drepanocerus patrizii Boucomont, 1923 | | | Drepanocerus striatus (Boucomont, 1921) | ⇒ Clypeodrepanus striatus Boucomont, 1921 | | Drepanocerus strigatus (Janssens, 1953) | ⇒ Clypeodrepanus strigatus Janssens, 1953 | | Drepanocerus sulcicollis (Castelnau, 1840) | ⇒ Sulcodrepanus sulcicollis Castelnau, 1840 | | Drepanocerus trogiformis Roth, 1851 | | | Drepanoplatynus gilleti Boucomont, 1921 | | | Drepanopodus costatus Wiedemann, 1823 | | | Drepanopodus proximus Péringuey, 1901 | | | Dwesasilvasedis medinae Deschodt & S..., 2008 | | | Endroedyolus paradoxus Scholtz & Ho..., 1987 | | | Eodrepanus bechynei Janssens, 1953 | | | Eodrepanus fastiditus Péringuey, 1901 | | | Eodrepanus morgani Barbero, Pal..., 2009 | | | Eodrepanus paolae Barbero, Pal..., 2009 | | | Eodrepanus parallelus Raffray, 1877 | | | Epactoides andringitrae Paulian, 1975 | | | Epactoides ankasokae Paulian, 1975 | | | Epactoides basilewskyi Balthasar, 1960 | | | Epactoides betschi Paulian, 1975 | | | Epactoides costatus Paulian, 1992 | | | Epactoides emkeni Montreuil, 2017 | | | Epactoides femoralis Montreuil, 2005 | | | Epactoides fiorii Paulian, 1976 | | | Epactoides frontalis Montreuil, 2003 | | | Epactoides giganteus Rossini, Vaz..., 2021 | | | Epactoides hanskii Montreuil, 2005 | | | Epactoides helenae Montreuil, 2005 | | | Epactoides humberti Paulian, 1975 | | | Epactoides humeralis Paulian, 1976 | | | Epactoides hyphydroides Lebis, 1953 | | | Epactoides incertus Lebis, 1953 | | | Epactoides jounii Wirta & Mont..., 2008 | | | Epactoides lacustris Paulian, 1976 | | | Epactoides lissus Lebis, 1953 | | | Epactoides madecassus Paulian, 1936 | | | Epactoides mahaboi Paulian, 1976 | | | Epactoides major Paulian, 1991 | | | Epactoides mananarae Paulian, 1976 | | | Epactoides mangabeensis Wirta & Mont..., 2008 | | | Epactoides masoalae Paulian, 1976 | | | Epactoides olsoufieffi Lebis, 1953 | | | Epactoides paradoxus Paulian, 1976 | | | Epactoides pauliani Lebis, 1953 | | | Epactoides perinetanus Paulian, 1975 | | | Epactoides perrieri Fairmaire, 1898 | | | Epactoides rahagai Montreuil, 2005 | | | Epactoides sambavae Paulian, 1976 | | | Epactoides semiaeneus Lebis, 1953 | | | Epactoides spinicollis Montreuil, 2003 | | | Epactoides tiinae Montreuil, 2005 | | | Epactoides vadoni Montreuil, 2005 | | | Epactoides vagecarinatus Lebis, 1953 | | | Epactoides viridicollis Montreuil, 2005 | | | Epactoides vondrozoi Paulian, 1976 | | | Epilissus alluaudi Lebis, 1953 | | | Epilissus andranobeensis Montreuil, 2011 | | | Epilissus antoetrae Paulian, 1975 | | | Epilissus apotolamproides Lebis, 1961 | | | Epilissus armipes Lebis, 1953 | | | Epilissus bemarahaensis Montreuil & ..., 2022 | | | Epilissus colasi Paulian, 1935 | | | Epilissus cuprarius Fairmaire, 1889 | | | Epilissus cuprarius discretus Montreuil, 2006 | | | Epilissus cuprarius serindicus Paulian, 1976 | | | Epilissus delphinensis Lebis, 1953 | | | Epilissus discretus Montreuil, 2006 | | | Epilissus emmae Lebis, 1953 | | | Epilissus emmae emmae Lebis, 1953 | | | Epilissus emmae obscuripennis Montreuil, 2006 | | | Epilissus fantamattii Montreuil, 2011 | | | Epilissus fuscovirens (Fairmaire, 1889) | ⇒ Arachnodes fuscovirens Fairmaire, 1889 | | Epilissus genieri Montreuil, 2006 | | | Epilissus guillaumeti Paulian, 1975 | | | Epilissus hova Künckel d’..., 1887 | | | Epilissus mantasoae Paulian, 1976 | | | Epilissus micheli Lebis, 1953 | | | Epilissus morio (Harold, 1879) | ⇒ Arachnodes morio Harold, 1879 | | Epilissus morio hova Künckel d’He..., 1887 | | | Epilissus morio morio Harold, 1879 | | | Epilissus morio serindicus Paulian, 1976 | | | Epilissus planatus Montreuil, 2006 | | | Epilissus prasinus Klug, 1832 | | | Epilissus robustus Montreuil & ..., 2022 | | | Epilissus ruteri Lebis, 1953 | | | Epilissus semicribrosus (Fairmaire, 1898) | ⇒ Nanos semicribrosus Fairmaire, 1898 | | Epilissus semilissus Montreuil & ..., 2022 | | | Epilissus sikorai Paulian, 1977 | | | Epilissus splendidus (Fairmaire, 1889) | ⇒ Arachnodes splendidus Lebis, 1953 | | Epilissus splendidus delphinensis Lebis, 1953 | | | Epilissus splendidus descarpentriesi Lebis, 1953 | | | Epilissus splendidus splendidus Fairmaire, 1889 | | | Epilissus striatoides Paulian, 1976 | | | Epilissus striatus Castelnau, 1840 | | | Epilissus superbus Montreuil & ..., 2022 | | | Epilissus viridis Klug, 1832 | | | Epilissus vohemarensis Montreuil & ..., 2022 | | | Epirinus aeneus Wiedemann, 1823 | | | Epirinus aquilus Medina & Sch..., 2005 | | | Epirinus asper Péringuey, 1901 | | | Epirinus bentoi Ferreira, 1964 | | | Epirinus comosus Péringuey, 1901 | | | Epirinus convexus Scholtz & Ho..., 1987 | | | Epirinus davisi Scholtz & Ho..., 1987 | | | Epirinus drakomontanus Scholtz & Ho..., 1987 | | | Epirinus flagellatus Fabricius, 1775 | | | Epirinus granulatus Scholtz & Ho..., 1987 | | | Epirinus gratus Péringuey, 1901 | | | Epirinus hilaris Péringuey, 1901 | | | Epirinus hluhluwensis Medina & Sch..., 2005 | | | Epirinus hopei Castelnau, 1840 | | | Epirinus inparrugosus Deschodt & D..., 2019 | | | Epirinus jacobsae Deschodt & D..., 2019 | | | Epirinus minimus Medina & Sch..., 2005 | | | Epirinus montanus Scholtz & Ho..., 1987 | | | Epirinus mucrodentatus Scholtz & Ho..., 1987 | | | Epirinus muellerae Deschodt & D..., 2019 | | | Epirinus ngomae Medina & Sch..., 2005 | | | Epirinus obtusus Boheman, 1857 | | | Epirinus pseudorelictus Deschodt & D..., 2019 | | | Epirinus pseudorugosus Medina & Sch..., 2005 | | | Epirinus punctatus Scholtz & Ho..., 1987 | | | Epirinus pygidialis Scholtz & Ho..., 1987 | | | Epirinus pygidialus Scholtz & Ho..., 1987 | | | Epirinus relictus Scholtz & Ho..., 1987 | | | Epirinus rugosus Scholtz & Ho..., 1987 | | | Epirinus schoolmeestersi Deschodt & D..., 2019 | | | Epirinus scrobiculatus Harold, 1880 | | | Epirinus sebastiani Medina & Sch..., 2005 | | | Epirinus silvestris Cambefort, 1978 | | | Epirinus striatus Scholtz & Ho..., 1987 | | | Epirinus sulcipennis Boheman, 1857 | | | Epirinus validus Péringuey, 1901 | | | Euoniticellus africanus Harold, 1873 | | | Euoniticellus capnus Cambefort, 1996 | | | Euoniticellus fulvus Goeze, 1777 | | | Euoniticellus fumigatus Boucomont, 1923 | | | Euoniticellus inaequalis Reiche, 1850 | | | Euoniticellus intermedius Reiche, 1848 | | | Euoniticellus kawanus Janssens, 1939 | | | Euoniticellus nasicornis Reiche, 1849 | | | Euoniticellus pallens Olivier, 1789 | | | Euoniticellus pallipes Fabricius, 1781 | | | Euoniticellus parvus Kraatz, 1895 | | | Euoniticellus perniger Cambefort & ..., 1980 | | | Euoniticellus tai Cambefort, 1983 | | | Euoniticellus tibatensis Kolbe, 1904 | | | Euoniticellus triangulatus Harold, 1873 | | | Euoniticellus wittei Janssens, 1939 | | | Euoniticellus zumpti Janssens, 1953 | | | Euonthophagus amieti Cambefort, 1972 | | | Euonthophagus amyntas Olivier, 1789 | | | Euonthophagus bedeli Reitter, 1891 | | | Euonthophagus carbonarius Klug, 1855 | | | Euonthophagus consanguineus Müller, 1944 | | | Euonthophagus consobrinus Péringuey, 1901 | | | Euonthophagus conterminus Petrovitz, 1971 | | | Euonthophagus crocatus Mulsant & Go..., 1870 | | | Euonthophagus crocatus clementi Baraud, 1960 | | | Euonthophagus crocatus crocatus | | | Euonthophagus flavimargo Orbigny, 1902 | | | Euonthophagus jeanneli diversiceps Frey, 1975 | | | Euonthophagus jeanneli jeanneli | | | Euonthophagus maindroni d´Orbigny, 1897 | | | Euonthophagus ustus d´Orbigny, 1905 | | | Euonthophagus vicarius Péringuey, 1901 | | | Eurysternus caribaeus Herbst, 1789 | | | Eurysternus magnus Castelnau, 1840 | | | Eurysternus mexicanus Harold, 1869 | | | Eurysternus plebejus Harold, 1880 | | | Eusaproecius congolanus Balthasar, 1963 | | | Eusaproecius dudleyi Branco, 1992 | | | Eusaproecius laticeps d´Orbigny, 1908 | | | Eusaproecius magnigena d´Orbigny, 1913 | | | Eusaproecius optatus Péringuey, 1901 | | | Eusaproecius punctulatus Boucomont, 1928 | | | Eusaproecius tinantae Boucomont, 1923 | | | Eutrichillum arcus Solis & Kohl..., 2003 | | | Frankenbergerius armatus Boheman, 1857 | | | Frankenbergerius armatus armatus | | | Frankenbergerius armatus tuberculatus Frolov & Sch..., 2005 | | | Frankenbergerius barratti Waterhouse, 1876 | | | Frankenbergerius forcipatus Harold, 1880 | | | Frankenbergerius gomesi Ferreira, 1954 | | | Frankenbergerius nanus Frolov & Sch..., 2005 | | | Frankenbergerius nitidus Frolov & Sch..., 2005 | | | Frankenbergerius opacus Frolov & Sch..., 2005 | | | Garreta annoyeri Moretto, 2021 | | | Garreta australugens Davis & Desc..., 2018 | | | Garreta azareus | | | Garreta azureus Fabricius, 1801 | | | Garreta basilewskyi Balthasar, 1961 | | | Garreta bechynei Pokorný & Zí..., 2018 | | | Garreta caffer Fahraeus, 1857 | | | Garreta crenulatus Kolbe, 1895 | | | Garreta dejeani Castelnau, 1840 | | | Garreta diffinis Waterhouse, 1890 | | | Garreta fastiditus Harold, 1867 | | | Garreta laetus Hope, 1842 | | | Garreta laetus laetus | | | Garreta laetus olivaceus Quedenfeldt, 1884 | | | Garreta lugens Fairmaire, 1891 | | | Garreta malleolus Kolbe, 1895 | | | Garreta matabelensis Janssens, 1938 | | | Garreta namalugens Davis & Desc..., 2018 | | | Garreta nitens Olivier, 1789 | | | Garreta nyassicus Kolbe, 1897 | | | Garreta rutilans Castelnau, 1840 | | | Garreta sumptuosus Castelnau, 1840 | | | Garreta unicolor Fahraeus, 1857 | | | Garreta wahlbergi Fahraeus, 1857 | | | Garreta zumpti Frey, 1967 | | | Gilletellus collarti Janssens, 1937 | | | Gilletellus porculus Boheman, 1857 | | | Gymnopleurus aenescens Wiedemann, 1821 | | | Gymnopleurus aeruginosus Harold, 1867 | | | Gymnopleurus andreaei Ferreira, 1954 | | | Gymnopleurus asperrimus Felsche, 1909 | | | Gymnopleurus bicolor (Latreille, 1823) | ⇒ Gymnopleurus leei Fabricius, 1792 | | Gymnopleurus coerulescens Olivier, 1789 | | | Gymnopleurus colmanti Janssens, 1938 | | | Gymnopleurus coriarius (Herbst, 1789) | ⇒ Gymnopleurus flagellatus Fabricius, 1787 | | Gymnopleurus cyaneus (Fabricius, 1798) | ⇒ Gymnopleurus sericeifrons Fairmaire, 1887 | | Gymnopleurus flagellatus Fabricius, 1787 | | | Gymnopleurus flagellatus flagellatus Fabricius, 1787 | | | Gymnopleurus foricarius Garreta, 1914 | | | Gymnopleurus fulgidus Olivier, 1789 | | | Gymnopleurus geoffroyi Fuessly, 1775 | | | Gymnopleurus geoffroyi geoffroyi | | | Gymnopleurus humanus MacLeay, 1821 | | | Gymnopleurus humeralis Klug, 1855 | | | Gymnopleurus ignitus Klug, 1855 | | | Gymnopleurus imitator Balthasar, 1963 | | | Gymnopleurus jacksoni Waterhouse, 1890 | | | Gymnopleurus koenigi Fabricius, 1775 | | | Gymnopleurus laetus Hope, 1842 | | | Gymnopleurus laevicollis (Castelnau, 1840) | ⇒ Gymnopleurus purpureus Garreta, 1914 | | Gymnopleurus latreillei Castelnau, 1840 | | | Gymnopleurus leei Fabricius, 1792 | | | Gymnopleurus moerens Kolbe, 1895 | | | Gymnopleurus mopsus Pallas, 1781 | Breiter Pillendreher | | Gymnopleurus mopsus mopsus | | | Gymnopleurus nyankpalaensis Endrödi, 1976 | | | Gymnopleurus particolor Davis, 1986 | | | Gymnopleurus plicatulus Fairmaire, 1890 | | | Gymnopleurus profanus (Fabricius, 1792) | ⇒ Garreta nitens Olivier, 1789 | | Gymnopleurus pumilus (Reiche, 1847) | ⇒ Gymnopleurus virens Erichson, 1843 | | Gymnopleurus puncticollis (Gillet, 1909) | ⇒ Gymnopleurus virens Erichson, 1843 | | Gymnopleurus purpureus Garreta, 1914 | | | Gymnopleurus pustulatus Kolbe, 1895 | | | Gymnopleurus reichei Waterhouse, 1890 | | | Gymnopleurus rhodesianus Balthasar, 1963 | | | Gymnopleurus ruandensis Janssens, 1938 | | | Gymnopleurus sericatus (Erichson, 1843) | ⇒ Gymnopleurus humanus MacLeay, 1821 | | Gymnopleurus sericeifrons Fairmaire, 1887 | | | Gymnopleurus smaragdinus (Fahraeus, 1857) | ⇒ Gymnopleurus ignitus Klug, 1855 | | Gymnopleurus somaliensis Lansberge, 1882 | | | Gymnopleurus sturmi (MacLeay, 1821) | ⇒ Gymnopleurus sturmii MacLeay, 1821 | | Gymnopleurus sturmii MacLeay, 1821 | | | Gymnopleurus thelwalli Waterhouse, 1890 | | | Gymnopleurus thoracicus Harold, 1868 | | | Gymnopleurus tristis Castelnau, 1840 | | | Gymnopleurus virens Erichson, 1843 | | | Hammondantus psammophilus Cambefort, 1978 | | | Haroldius convexus Philips & Sc..., 2000 | | | Haroldius ennearthrus Janssens, 1949 | | | Haroldius leleupi Janssens, 1953 | | | Haroldius lyleae Daniel, Strü..., 2021 | | | Haroldius modestus Janssens, 1953 | | | Haroldius stefanii Montreuil, 2010 | | | Helictopleurus aeneoniger Lebis, 1960 | | | Helictopleurus ambiguus Paulian & Ca..., 1991 | | | Helictopleurus aurocupreus Fairmaire, 1901 | | | Helictopleurus basicornis Fairmaire, 1904 | | | Helictopleurus bivittatus d´Orbigny, 1915 | | | Helictopleurus cambeforti Montreuil, 2005 | | | Helictopleurus carbonarius Lebis, 1960 | | | Helictopleurus clouei Harold, 1869 | | | Helictopleurus coruscus d´Orbigny, 1915 | | | Helictopleurus cribricollis Lebis, 1960 | | | Helictopleurus dorbignyi Montreuil, 2005 | | | Helictopleurus fasciolatus Fairmaire, 1898 | | | Helictopleurus fasciolatus fasciolatus | | | Helictopleurus fasciolatus obscurus Lebis, 1960 | | | Helictopleurus fasciolatus pseudofasciolatus Montreuil, 2007 | | | Helictopleurus fasciolatus suturalis Montreuil, 2012 | | | Helictopleurus fissicollis Fairmaire, 1895 | | | Helictopleurus fissicollis australis Montreuil, 2007 | | | Helictopleurus fissicollis fissicollis | | | Helictopleurus fissicollis morettoi Montreuil, 2005 | | | Helictopleurus fulgens Lebis, 1960 | | | Helictopleurus fulgens fulgens | | | Helictopleurus fulgens purpuricollis Lebis, 1960 | | | Helictopleurus fungicola Fairmaire, 1899 | | | Helictopleurus fungicola peyrierasi Paulian & Ca..., 1991 | | | Helictopleurus furcicornis Lebis, 1960 | | | Helictopleurus gibbicollis Fairmaire, 1895 | | | Helictopleurus giganteus Harold, 1869 | | | Helictopleurus halffteri Balthasar, 1964 | | | Helictopleurus hanskii Montreuil, 2005 | | | Helictopleurus heidiae Montreuil, 2007 | | | Helictopleurus hypocrita Balthasar, 1941 | | | Helictopleurus infimus Fairmaire, 1901 | | | Helictopleurus littoralis Montreuil, 2005 | | | Helictopleurus marsyas Olivier, 1789 | | | Helictopleurus minutus Montreuil, 2007 | | | Helictopleurus neoamplicollis Krell, 2000 | | | Helictopleurus neuter Fairmaire, 1898 | | | Helictopleurus nicollei Lebis, 1960 | | | Helictopleurus niger d´Orbigny, 1915 | | | Helictopleurus nigricans d´Orbigny, 1915 | | | Helictopleurus nigritulus Lebis, 1960 | | | Helictopleurus nitidicollis Lebis, 1960 | | | Helictopleurus patricii Paulian, 1937 | | | Helictopleurus perpunctatus Balthasar, 1963 | | | Helictopleurus perrieri Fairmaire, 1898 | | | Helictopleurus peyrierasi (Paulian & Ca..., 1991) | ⇒ Helictopleurus fungicola peyrierasi Paulian & Ca..., 1991 | | Helictopleurus pluristriatus d´Orbigny, 1915 | | | Helictopleurus politicollis Fairmaire, 1902 | | | Helictopleurus praerugosus Fairmaire, 1898 | | | Helictopleurus pseudoniger Montreuil, 2005 | | | Helictopleurus quadripunctatus Olivier, 1789 | | | Helictopleurus rubricollis Lebis, 1960 | | | Helictopleurus rudicollis Fairmaire, 1898 | | | Helictopleurus semimetallicus Fairmaire, 1888 | | | Helictopleurus seminiger d´Orbigny, 1915 | | | Helictopleurus semivirens d´Orbigny, 1915 | | | Helictopleurus sericeus Lebis, 1960 | | | Helictopleurus sicardi Paulian, 1975 | | | Helictopleurus sinuaticornis Fairmaire, 1898 | | | Helictopleurus sinuatocornis Fairmaire, 1898 | | | Helictopleurus splendidicollis Fairmaire, 1893 | | | Helictopleurus splendidicollis similis Montreuil, 2007 | | | Helictopleurus splendidicollis splendidicollis | | | Helictopleurus splendidus d´Orbigny, 1915 | | | Helictopleurus steineri Paulian & Ca..., 1991 | | | Helictopleurus subretusus Fairmaire, 1895 | | | Helictopleurus sumptuosus Harold, 1873 | | | Helictopleurus tamatavensis Montreuil, 2005 | | | Helictopleurus tristis (d´Orbigny, 1915) | ⇒ Helictopleurus quadripunctatus Olivier, 1789 | | Helictopleurus undatus Olivier, 1789 | | | Helictopleurus unifasciatus Fairmaire, 1901 | | | Helictopleurus vadoni Lebis, 1960 | | | Helictopleurus viettei Paulian & Ca..., 1984 | | | Helictopleurus villiersi Paulian & Ca..., 1984 | | | Helictopleurus viridans Fairmaire, 1901 | | | Helictopleurus viridans pseudoviridans Montreuil, 2007 | | | Helictopleurus viridans viridans | | | Helictopleurus viridiflavus Fairmaire, 1898 | | | Heliocopris alatus Felsche, 1910 | | | Heliocopris anadematus Gillet, 1908 | | | Heliocopris andersoni Bates, 1868 | | | Heliocopris anguliceps Janssens, 1943 | | | Heliocopris antenor (Olivier, 1789) | ⇒ Heliocopris colossus Bates, 1868 | | Heliocopris atropos Boheman, 1860 | | | Heliocopris beccarii Harold, 1871 | | | Heliocopris biimpressus Kolbe, 1893 | | | Heliocopris bucephalus Fabricius, 1775 | Indianische Ochsenkopf | | Heliocopris camerunus Pokorny & Zidek, 2009 | | | Heliocopris colossus Bates, 1868 | | | Heliocopris corniculatus Janssens, 1939 | | | Heliocopris coronatus Felsche, 1901 | | | Heliocopris cuneifer Lesne, 1906 | | | Heliocopris densissa Roth, 1851 | | | Heliocopris dianae (Hope, 1842) | ⇒ Heliocopris mutabilis Kolbe, 1893 | | Heliocopris dilloni Guérin-Ménev..., 1847 | | | Heliocopris dolosus Janssens, 1939 | | | Heliocopris erycoides Felsche, 1907 | | | Heliocopris eryx (Fabricius, 1801) | ⇒ Heliocopris neptunus Boheman, 1857 | | Heliocopris eryx eryx | | | Heliocopris eryx ikelenge Moretto, 2017 | | | Heliocopris eryx mutabilis (Kolbe, 1895) | ⇒ Heliocopris mutabilis Kolbe, 1893 | | Heliocopris eryx tridentatus Moretto, 2017 | | | Heliocopris faunus Boheman, 1857 | | | Heliocopris felschei Kolbe, 1904 | | | Heliocopris fischeri Schäfer, 2019 | | | Heliocopris fonsecai Ferreira, 1967 | | | Heliocopris furcithorax Müller, 1941 | | | Heliocopris gigas Linnaeus, 1758 | Ägyptische Riese | | Heliocopris gigas erythraeus Müller, 1941 | | | Heliocopris gigas gigas | | | Heliocopris hamadryas Fabricius, 1775 | | | Heliocopris hamifer Harold, 1878 | | | Heliocopris haroldi Kolbe, 1893 | | | Heliocopris helleri Felsche, 1907 | | | Heliocopris hermes Gillet, 1911 | | | Heliocopris hunteri Waterhouse, 1891 | | | Heliocopris japetus Klug, 1855 | | | Heliocopris kolbei Felsche, 1901 | | | Heliocopris letiranti Moretto, 2021 | | | Heliocopris maou Moretto, 2014 | | | Heliocopris marshalli Péringuey, 1901 | | | Heliocopris mimus Janssens, 1939 | | | Heliocopris minos Gillet, 1907 | | | Heliocopris mutabilis Kolbe, 1893 | | | Heliocopris myrmidon Kolbe, 1893 | | | Heliocopris neptuniformis Felsche, 1907 | | | Heliocopris neptunoides Janssens, 1939 | | | Heliocopris neptunus Boheman, 1857 | | | Heliocopris pauliani Janssens, 1939 | | | Heliocopris pirmal Fabricius, 1798 | | | Heliocopris quinqueangulatus Janssens, 1939 | | | Heliocopris samson Harold, 1878 | | | Heliocopris sirius (Gillet, 1925) | ⇒ Heliocopris hamifer Harold, 1878 | | Heliocopris smithi Moretto, 2017 | | | Heliocopris solitarius Kolbe, 1893 | | | Heliocopris staudingeri Kolbe, 1893 | | | Heliocopris stroehlei Moretto, 2014 | | | Heliocopris sylvanus Gillet, 1925 | | | Heliocopris tyrannus Thomson, 1859 | | | Heliocopris werneri Moretto & Mi..., 2013 | | | Heteroclitopus annamariae Branco, 1991 | | | Heteroclitopus collarti Janssens, 1939 | | | Heteroclitopus ferreirae Branco, 1991 | | | Heteroclitopus foveatus Boucomont, 1923 | | | Heteroclitopus foveolatus Branco & Josso, 2007 | | | Heteroclitopus freyi Ferreira, 1969 | | | Heteroclitopus herteli Ferreira, 1969 | | | Heteroclitopus remipes Péringuey, 1901 | | | Heteroclitopus schulzei Branco, 1991 | | | Heteroclitopus simonettai Branco, 1991 | | | Heteronitis castelnaui Harold, 1862 | | | Heteronitis pauliani Janssens, 1942 | | | Heteronitis ragazzii Fairmaire, 1893 | | | Heteronitis tricornutus Fairmaire, 1891 | | | Heteronitis tridens Castelnau, 1840 | | | Heterosyphus sicardi Paulian, 1975 | | | Hyalonthophagus alcedo Orbigny, 1913 | | | Hyalonthophagus alcyon Klug, 1855 | | | Hyalonthophagus alcyonides Orbigny, 1913 | | | Hyalonthophagus hyalcyon Palestrini &..., 1989 | | | Hyalonthophagus mixtifrons Orbigny, 1910 | | | Hyalonthophagus nigroviolaceus Orbigny, 1902 | | | Hyalonthophagus novalcedo Josso, 2021 | | | Hyalonthophagus pseudoalcyon Orbigny, 1913 | | | Hyalonthophagus pseudovirens Orbigny, 1913 | | | Hyalonthophagus pulcher Deschodt & D..., 2019 | | | Hyalonthophagus virens d´Orbigny, 1913 | | | Hyalonthophagus viridiceps Orbigny, 1910 | | | Ixodina abyssinica Roth, 1851 | | | Ixodina abyssinica abyssinica | | | Ixodina abyssinica tangana Endrödi, 1971 | | | Ixodina abyssinica tanganus Endrodi, 1971 | | | Ixodina bos Endrödi, 1976 | | | Ixodina endroedyi Endrödi, 1976 | | | Ixodina freyi Janssens, 1953 | | | Ixodina kovacsi Endrödi, 1976 | | | Ixodina saegeri Balthasar, 1963 | | | Ixodina szunyoghyi Endrödi, 1971 | | | Ixodina szunyoghyi occidentalis Endrödi, 1976 | | | Ixodina szunyoghyi szunyoghyi | | | Janssensantus alatus Josso, 2022 | | | Janssensantus inexpectatus Paulian, 1976 | | | Janssensantus pauliani Scholtz & Ho..., 1987 | | | Janssensellus bicaviventris Cambefort, 1975 | | | Janssensellus rasplusi Cambefort, 1995 | | | Kheper aegyptiorum Latreille, 1823 | | | Kheper aegyptiorum aegyptiorum | | | Kheper aegyptiorum purpurascens Gerstaecker, 1871 | | | Kheper aeratus Gerstaecker, 1871 | | | Kheper asceticus Gillet, 1907 | | | Kheper bonellii MacLeay, 1821 | | | Kheper clericus Boheman, 1857 | | | Kheper cupreus Castelnau, 1840 | | | Kheper cuvieri MacLeay, 1821 | | | Kheper festivus Harold, 1868 | | | Kheper festivus festivus | | | Kheper festivus nicolasi Moretto, 1998 | | | Kheper festivus simoni Moretto, 1998 | | | Kheper festivus wellsi Moretto, 1998 | | | Kheper intermedius Gillet, 1909 | | | Kheper kalaharicus Davis, Desch..., 2011 | | | Kheper laevistriatus Fairmaire, 1893 | | | Kheper lamarcki MacLeay, 1821 | | | Kheper nigroaeneus Boheman, 1857 | | | Kheper paganus Harold, 1878 | | | Kheper platynotus Bates, 1888 | | | Kheper porosus Bates, 1888 | | | Kheper prodigiosus Erichson, 1843 | | | Kheper pustulosus Gerstaecker, 1871 | | | Kheper rolciki Pokorný & Zí..., 2015 | | | Kheper subaeneus Harold, 1869 | | | Kheper vanderkelleni Lansberge, 1886 | | | Kheper venerabilis Harold, 1871 | | | Kheper vethi Lansberge, 1886 | | | Kheper zurstrasseni Davis, 1986 | | | Kolbeellus ateuchoides Lansberge, 1875 | | | Krikkenius smeenki Krikken, 1982 | | | Latodrepanus caelatus Gerstaecker, 1871 | | | Latodrepanus laticollis Fahraeus, 1857 | | | Latodrepanus nicolasi Barbero, Pal..., 2009 | | | Latodrepanus pulvinarius Balthasar, 1963 | | | Latodrepanus schimperi Janssens, 1953 | | | Latodrepanus simonisi Barbero, Pal..., 2009 | | | Lepanus monteithi Matthews, 1974 | | | Lepanus pygmaeus MacLeay, 1888 | | | Liatongus amitinus Boucomont, 1921 | | | Liatongus arrowi Boucomont, 1921 | | | Liatongus aterrimus Boucomont, 1921 | | | Liatongus femoratus Illiger, 1800 | | | Liatongus ferreirae Balthasar, 1964 | | | Liatongus fulvostriatus d´Orbigny, 1916 | | | Liatongus interruptus (Quedenfeldt, 1884) | ⇒ Liatongus fulvostriatus d´Orbigny, 1916 | | Liatongus militaris Castelnau, 1840 | | | Liatongus raffrayi Lansberge, 1886 | | | Liatongus schoutedeni Boucomont, 1920 | | | Liatongus sjostedti Felsche, 1904 | | | Liatongus spathulatus Roth, 1851 | | | Liatongus taurus Boucomont, 1920 | | | Liatongus testudo Lansberge, 1886 | | | Liatongus tuberculicollis Felsche, 1909 | | | Liatongus upembanus Janssens, 1953 | | | Liatongus urus Kolbe, 1914 | | | Litocopris muticus Boheman, 1857 | | | Litocopris punctiventris Waterhouse, 1891 | | | Litocopris ruandanus Balthasar, 1961 | | | Litocopris simplex Harold, 1869 | | | Lophodonitis carinatus Felsche, 1907 | | | Lophodonitis minettii Moretto, 2011 | | | Macroderes amplior Frolov & Sch..., 2004 | | | Macroderes arrowi Janssens, 1939 | | | Macroderes bias Olivier, 1789 | | | Macroderes cederbergensis Abdalla & De..., 2018 | | | Macroderes cornutus Frolov & Sch..., 2004 | | | Macroderes dubius Péringuey, 1901 | | | Macroderes endroedyi Frolov & Sch..., 2004 | | | Macroderes fornicatus Sharp, 1880 | | | Macroderes foveatus Frolov & Sch..., 2004 | | | Macroderes gifboomi Abdalla & De..., 2018 | | | Macroderes greeni Kirby, 1818 | | | Macroderes leipoldti Abdalla & De..., 2018 | | | Macroderes minutus Frolov & Sch..., 2004 | | | Macroderes mutilans Kolbe, 1908 | | | Macroderes mutilatus Kolbe, 1908 | | | Macroderes namakwanus Frolov & Sch..., 2004 | | | Macroderes nitidus Harold, 1877 | | | Macroderes oreatus Abdalla & De..., 2018 | | | Macroderes politulus (Preudhomme d..., 1880) | ⇒ Macroderes bias Olivier, 1789 | | Macroderes porselinus Abdalla & De..., 2018 | | | Macroderes pristinus Sharp, 1880 | | | Macroderes soleiana Abdalla & De..., 2018 | | | Macroderes spectabilis Péringuey, 1901 | | | Macroderes tortuosus Abdalla & De..., 2018 | | | Macroderes undulatus Preudhomme d..., 1880 | | | Madateuchus viettei (Paulian, 1953) | ⇒ Scarabaeus viettei Paulian, 1953 | | Malagoniella astyanax Olivier, 1789 | | | Megalonitis bohemani Lansberge, 1875 | | | Megalonitis bohemanni Lansberge, 1875 | | | Megalonitis gigas Bertoloni, 1855 | | | Megaponerophilus megaponerae Brauns, 1914 | | | Metacatharsius abortivus Fairmaire, 1891 | | | Metacatharsius anderseni Waterhouse, 1891 | | | Metacatharsius anomalus Felsche, 1910 | | | Metacatharsius auberti Fairmaire, 1891 | | | Metacatharsius basilewskyi Balthasar, 1961 | | | Metacatharsius bateke Walter, 1976 | | | Metacatharsius benadirensis Müller, 1947 | | | Metacatharsius bidentatus Ferreira, 1964 | | | Metacatharsius brevicosta Müller, 1947 | | | Metacatharsius clypeolatus Balthasar, 1940 | | | Metacatharsius convexiusculus Shipp, 1897 | | | Metacatharsius darfurensis Balthasar, 1965 | | | Metacatharsius dentinum Ferreira, 1964 | | | Metacatharsius duplicatus Müller, 1941 | | | Metacatharsius exiguiformis Ferreira, 1964 | | | Metacatharsius exiguus Boheman, 1860 | | | Metacatharsius ferreirae Balthasar, 1965 | | | Metacatharsius ferrugineus Olivier, 1789 | | | Metacatharsius freyi Ferreira, 1964 | | | Metacatharsius gardoensis Balthasar, 1941 | | | Metacatharsius holyi Balthasar, 1965 | | | Metacatharsius imitator Balthasar, 1965 | | | Metacatharsius inermis Castelnau, 1840 | | | Metacatharsius janssensi Frey, 1963 | | | Metacatharsius kindiai Paulian, 1939 | | | Metacatharsius latifrons Harold, 1868 | | | Metacatharsius lomii Müller, 1941 | | | Metacatharsius marani Balthasar, 1940 | | | Metacatharsius minutus Lansberge, 1882 | | | Metacatharsius modestus Balthasar, 1965 | | | Metacatharsius murati Paulian, 1939 | | | Metacatharsius nubiensis Harold, 1878 | | | Metacatharsius omoensis Müller, 1941 | | | Metacatharsius opacus Waterhouse, 1891 | | | Metacatharsius ovulum Balthasar, 1940 | | | Metacatharsius parcepunctatus Balthasar, 1965 | | | Metacatharsius patrizii Müller, 1947 | | | Metacatharsius peleus Olivier, 1789 | | | Metacatharsius pernitidus Müller, 1941 | | | Metacatharsius perpunctatus Balthasar, 1965 | | | Metacatharsius petrovitzi Balthasar, 1965 | | | Metacatharsius politus Paulian, 1934 | | | Metacatharsius pollicatus Harold, 1878 | | | Metacatharsius pseudoopacus Ferreira, 1965 | | | Metacatharsius pumilioniformis Ferreira, 1964 | | | Metacatharsius pumilionis Wallengren, 1881 | | | Metacatharsius pusio Kolbe, 1908 | | | Metacatharsius renaudi Frey, 1960 | | | Metacatharsius rochai Ferreira, 1964 | | | Metacatharsius rosinae Balthasar, 1970 | | | Metacatharsius rubidus Fairmaire, 1891 | | | Metacatharsius rugosipennis Frey, 1975 | | | Metacatharsius rusingae ♰ R. Paulian, 1976 | | | Metacatharsius seminulum Balthasar, 1940 | | | Metacatharsius simulator Balthasar, 1939 | | | Metacatharsius tchadensis Balthasar, 1961 | | | Metacatharsius tenellus Müller, 1947 | | | Metacatharsius togoensis Kolbe, 1893 | | | Metacatharsius transvaalensis Balthasar, 1968 | | | Metacatharsius trianguliceps Balthasar, 1965 | | | Metacatharsius tricarenatus Ferreira, 1964 | | | Metacatharsius troglodytes (Boheman, 1857) | ⇒ Metacatharsius anderseni Waterhouse, 1891 | | Metacatharsius tuberifrons Fairmaire, 1884 | | | Metacatharsius useramus Kolbe, 1893 | | | Metacatharsius wittei Ferreira, 1964 | | | Metacatharsius zuluanus Balthasar, 1965 | | | Milichus annae Nicolas & Ca..., 1994 | | | Milichus apicalis (Fahraeus, 1857) | ⇒ Milichus rhodesianus Péringuey, 1904 | | Milichus boucomonti Cambefort, 1996 | | | Milichus dudleyae Cambefort, 1996 | | | Milichus dudleyi Cambefort, 1996 | | | Milichus dudleyorum Cambefort, 1996 | | | Milichus durandi Cambefort, 1996 | | | Milichus inaequalis Boucomont, 1928 | | | Milichus inaequalis inaequalis | | | Milichus inaequalis lamottei Cambefort, 1996 | | | Milichus incisiceps Janssens, 1954 | | | Milichus jugatus d´Orbigny, 1907 | | | Milichus lecourti Nicolas & Ca..., 1994 | | | Milichus merzi Cambefort, 1983 | | | Milichus picticollis Gerstaecker, 1871 | | | Milichus rhodesianus Péringuey, 1904 | | | Milichus serratus d´Orbigny, 1907 | | | Mimonthophagus ambiguus Péringuey, 1901 | | | Mimonthophagus anomalus Klug, 1855 | | | Mimonthophagus apicehirtus d´Orbigny, 1915 | | | Mimonthophagus hinnulus Klug, 1832 | | | Mimonthophagus limbibasis d´Orbigny, 1905 | | | Mimonthophagus pictisternum d´Orbigny, 1902 | | | Mimonthophagus semicirculifer d´Orbigny, 1902 | | | Mimonthophagus semisetosus d´Orbigny, 1902 | | | Mnematidium multidentatum (Klug, 1845) | ⇒ Scarabaeus multidentatum Klug, 1845 | | Mnematidium multidentatus Klug, 1845 | | | Mnematium ritchiei (MacLeay, 1821) | ⇒ Scarabaeus ritchiei MacLeay, 1821 | | Namakwanus davisi Deschodt & S..., 2007 | | | Namakwanus endroedyi Deschodt, Da..., 2011 | | | Namakwanus irishi Scholtz & Ho..., 1987 | | | Namakwanus kamfferi Deschodt & D..., 2018 | | | Namakwanus minutus Deschodt & D..., 2018 | | | Namakwanus scholtzi Deschodt & D..., 2017 | | | Namakwanus streyi Frolov, 2005 | | | Nanos agaboides Boucomont, 1937 | | | Nanos andreiae Montreuil & ..., 2014 | | | Nanos ankaranae Paulian, 1976 | | | Nanos antsalovaensis Montreuil & ..., 2011 | | | Nanos antsihanakensis Lebis, 1953 | | | Nanos ater Castelnau, 1840 | | | Nanos bemarahaensis Montreuil & ..., 2014 | | | Nanos bicoloratus Lebis, 1953 | | | Nanos bimaculatus Kunckel, 1887 | | | Nanos binotatus Lebis, 1953 | | | Nanos clypeatus Castelnau, 1840 | | | Nanos constricticollis Montreuil & ..., 2014 | | | Nanos dubitatus Lebis, 1953 | | | Nanos fusconitens Fairmaire, 1899 | | | Nanos hanskii Montreuil & ..., 2007 | | | Nanos humbloti Lebis, 1953 | | | Nanos humeralis Paulian, 1975 | | | Nanos incertus Paulian, 1976 | | | Nanos magnus Montreuil & ..., 2014 | | | Nanos manomboensis Montreuil & ..., 2007 | | | Nanos manongorivoensis Montreuil & ..., 2014 | | | Nanos marojejyensis Montreuil & ..., 2014 | | | Nanos minutus Castelnau, 1840 | | | Nanos mirjae Montreuil & ..., 2014 | | | Nanos mixtus Montreuil & ..., 2014 | | | Nanos nitens Lebis, 1953 | | | Nanos nitens nitens | | | Nanos nitens pseudonitens Lebis, 1953 | | | Nanos occidentalis Paulian, 1976 | | | Nanos peyrierasi (Paulian, 1976) | ⇒ Apotolamprus peyrierasi Paulian, 1985 | | Nanos pseudofusconitens Montreuil & ..., 2014 | | | Nanos pseudominutus Montreuil & ..., 2014 | | | Nanos pseudorubromaculatus Montreuil & ..., 2014 | | | Nanos pseudoviettei Montreuil & ..., 2014 | | | Nanos punctatus Boucomont, 1937 | | | Nanos pygmaeus Castelnau, 1840 | | | Nanos ranomafanaensis Montreuil & ..., 2014 | | | Nanos rubromaculatus Kunckel d´He..., 1887 | | | Nanos rubrosignatus Lebis, 1953 | | | Nanos semicribrosus Fairmaire, 1898 | | | Nanos sinuatipes Boucomont, 1937 | | | Nanos vadoni Lebis, 1953 | | | Nanos viettei Paulian, 1976 | | | Nanos viridissimus Montreuil & ..., 2011 | | | Nebulasilvius insularis Deschodt & C..., 2008 | | | Nebulasilvius johani Deschodt & S..., 2008 | | | Neonitis nigritiae Gillet, 1918 | | | Neonitis rhodesiae Gillet, 1918 | | | Neosaproecius nyassalandicus Branco, 1990 | | | Neosaproecius trituberculatus Frey, 1958 | | | Neosisyphus angulicollis Felsche, 1909 | | | Neosisyphus armatus Gory, 1833 | | | Neosisyphus astriatus Schafer & Fi..., 2001 | | | Neosisyphus baoule Cambefort, 1984 | | | Neosisyphus barbarossa Wiedemann, 1823 | | | Neosisyphus basilewskyi Haaf, 1956 | | | Neosisyphus calcaratus Klug, 1855 | | | Neosisyphus confrater Kolbe, 1914 | | | Neosisyphus fortuitus Péringuey, 1901 | | | Neosisyphus gladiator Arrow, 1927 | | | Neosisyphus infuscatus Klug, 1855 | | | Neosisyphus kuehni Haaf, 1955 | | | Neosisyphus macrorubrus Paschalidis, 1974 | | | Neosisyphus mirabilis Arrow, 1927 | | | Neosisyphus paschalidisae Cambefort, 1984 | | | Neosisyphus penicillatus Harold, 1880 | | | Neosisyphus quadricollis Gory, 1833 | | | Neosisyphus rubripes Péringuey, 1901 | | | Neosisyphus rubrus Paschalidis, 1974 | | | Neosisyphus rugosus Gory, 1833 | | | Neosisyphus setiger Roth, 1851 | | | Neosisyphus spinipes Thunberg, 1818 | | | Neosisyphus tai Cambefort, 1984 | | | Neosisyphus tibialis Raffray, 1877 | | | Neosisyphus trochantericus Fairmaire, 1891 | | | Nesosisyphus pygmaeus Vinson, 1946 | | | Nesosisyphus regnardi Alluaud, 1898 | | | Nesosisyphus rotundatus Vinson, 1946 | | | Nesosisyphus vicinus Vinson, 1939 | | | Nesovinsonia vinsoni Paulian, 1939 | | | Nitiocellus collarti Janssens, 1939 | | | Nitiocellus panthera Boucomont, 1921 | | | Odontoloma apiculum Howden & Sch..., 1987 | | | Odontoloma cambeforti Howden & Sch..., 1987 | | | Odontoloma dentinum Harold, 1868 | | | Odontoloma disalatum Howden & Sch..., 1987 | | | Odontoloma doubei Howden & Sch..., 1987 | | | Odontoloma endroedyi Howden & Sch..., 1987 | | | Odontoloma louwi Howden & Sch..., 1987 | | | Odontoloma metasternale d´Orbigny, 1902 | | | Odontoloma multipunctatum Howden & Sch..., 1987 | | | Odontoloma obscurum Howden & Sch..., 1987 | | | Odontoloma pauliani Walter, 1976 | | | Odontoloma pauxillum Boheman, 1857 | | | Odontoloma peckorum Howden & Sch..., 1987 | | | Odontoloma planatum Howden & Sch..., 1987 | | | Odontoloma pusillum Howden & Sch..., 1987 | | | Odontoloma pygidiale Péringuey, 1901 | | | Odontoloma quadridens d´Orbigny, 1908 | | | Odontoloma relictum Cambefort, 1983 | | | Odontoloma sculpturatum Harold, 1868 | | | Odontoloma spinicaudum Howden & Sch..., 1987 | | | Oniticellus ambiguus (Lacordaire, 1856) | ⇒ Cyptochirus trogiformis Roth, 1851 | | Oniticellus egregius Klug, 1855 | | | Oniticellus formosus (Chevrolat, 1830) | ⇒ Oniticellus pictus Hausmann, 1807 | | Oniticellus inaequalis (Reiche, 1847) | ⇒ Euoniticellus inaequalis Reiche, 1850 | | Oniticellus intermedius (Reiche, 1848) | ⇒ Euoniticellus intermedius Reiche, 1848 | | Oniticellus militaris (Castelnau, 1840) | ⇒ Liatongus militaris Castelnau, 1840 | | Oniticellus pallens (Castelnau, 1840) | ⇒ Euoniticellus intermedius Reiche, 1848 | | Oniticellus pictus Hausmann, 1807 | | | Oniticellus planatus Castelnau, 1840 | | | Oniticellus pseudoplanatus Balthasar, 1964 | | | Oniticellus spinipes (Roth, 1851) | ⇒ Tiniocellus spinipes Roth, 1851 | | Oniticellus tessellatus Harold, 1879 | | | Onitis abyssinicus (Reiche, 1847) | ⇒ Onitis lycophron Klug, 1855 | | Onitis adelphus Kolbe, 1914 | | | Onitis adriani Ferreira, 1976 | | | Onitis aeneomicans Gillet, 1918 | | | Onitis aeneus Lansberge, 1875 | | | Onitis aerarius Harold, 1878 | | | Onitis aeruginosus Klug, 1855 | | | Onitis aethiops Lesne, 1906 | | | Onitis affinis Felsche, 1911 | | | Onitis albertcollarti Génier & Mor..., 2019 | | | Onitis alexis (Klug, 1835) | ⇒ Cheironitis ungaricus ungaricus Herbst, 1789 | | Onitis alexis alexis | | | Onitis alexis septentrionalis Balthasar, 1942 | | | Onitis androcles Janssens, 1937 | | | Onitis anthracinus Felsche, 1907 | | | Onitis archettii Janssens, 1943 | | | Onitis artuosus Gillet, 1909 | | | Onitis autumnalis Davis, 1986 | | | Onitis aygulus Fabricius, 1781 | | | Onitis bayanga Moretto, 2007 | | | Onitis belial Fabricius, 1798 | | | Onitis bilobatus Ferreira, 1976 | | | Onitis bispicticollis Krikken, 1977 | | | Onitis bocandei Gillet, 1911 | | | Onitis bredoi Janssens, 1949 | | | Onitis brucemillerorum Josso & Mont..., 2017 | | | Onitis caffer Boheman, 1857 | | | Onitis cambeforti Moretto, 2007 | | | Onitis camerunus (Gillet, 1933) | ⇒ Onitis cryptodus Gillet, 1918 | | Onitis camiadei Josso, 2012 | | | Onitis camiadeianus Josso & Mont..., 2017 | | | Onitis cerrutii Müller, 1949 | | | Onitis chalceus Lansberge, 1875 | | | Onitis confusus Boheman, 1860 | | | Onitis coxalis Gillet, 1918 | | | Onitis crenatus Reiche, 1847 | | | Onitis cribratus Lansberge, 1875 | | | Onitis cryptodus Gillet, 1918 | | | Onitis cupreus Castelnau, 1840 | | | Onitis curvipes Lansberge, 1875 | | | Onitis deceptor Péringuey, 1901 | | | Onitis denticoxa Müller, 1941 | | | Onitis dimidiatus Felsche, 1907 | | | Onitis dispar Péringuey, 1896 | | | Onitis endroedii Ferreira, 1976 | | | Onitis excavaticollis Balthasar, 1942 | | | Onitis fabricii Roth, 1851 | | | Onitis femoralis Josso, 2012 | | | Onitis fractipes Boucomont, 1923 | | | Onitis fulgidus (Klug, 1855) | ⇒ Pleuronitis fulgidus Castelnau, 1840 | | Onitis fulmineus Janssens, 1943 | | | Onitis gazanus Arrow, 1909 | | | Onitis gilleti Boucomont, 1933 | | | Onitis granicollis Felsche, 1907 | | | Onitis guineensis Gillet, 1918 | | | Onitis humerosus Pallas, 1771 | | | Onitis inflaticollis Felsche, 1907 | | | Onitis insuetus (Kolbe, 1914) | ⇒ Onitis sulcipennis Felsche, 1907 | | Onitis inversidens Lansberge, 1875 | | | Onitis ion Olivier, 1789 | | | Onitis janssenii Gomes Alves, 1954 | | | Onitis jeanneli Janssens, 1942 | | | Onitis jossoi Moretto, 2010 | | | Onitis kasanka Josso, 2012 | | | Onitis keniensis Gillet, 1933 | | | Onitis kingstoni Krikken, 1977 | | | Onitis laminosus Ferreira, 1976 | | | Onitis lamnifer Gillet, 1909 | | | Onitis laticollis Lansberge, 1875 | | | Onitis licitus Péringuey, 1901 | | | Onitis lobi Cambefort, 1984 | | | Onitis lobipes Felsche, 1907 | | | Onitis lognia Moretto, 2007 | | | Onitis longitibialis Ferreira, 1977 | | | Onitis lycophron Klug, 1855 | | | Onitis malleatus Janssens, 1937 | | | Onitis marshalli Janssens, 1939 | | | Onitis mendax Gillet, 1918 | | | Onitis menieri Cambefort, 1981 | | | Onitis meruensis Ferreira, 1977 | | | Onitis meyeri Kolbe, 1891 | | | Onitis miesseni Gusakov & Kl..., 2009 | | | Onitis minutus Lansberge, 1875 | | | Onitis mniszechi Lansberge, 1875 | | | Onitis mniszechianus Janssens, 1951 | | | Onitis mnizechianus | | | Onitis monstrosus Felsche, 1910 | | | Onitis montreuili Josso, 2012 | | | Onitis mossambicensis Ferreira, 1976 | | | Onitis mraci Moretto, 2021 | | | Onitis multidentatus Gillet, 1918 | | | Onitis nemoralis Gillet, 1932 | | | Onitis nigeriensis Ferreira, 1977 | | | Onitis nubiensis Lansberge, 1875 | | | Onitis numida Castelnau, 1840 | | | Onitis numida numida | | | Onitis obenbergeri Balthasar, 1942 | | | Onitis obscurus Lansberge, 1886 | | | Onitis occidentalis Gillet, 1933 | | | Onitis orthopus Lansberge, 1875 | | | Onitis overlaeti Janssens, 1938 | | | Onitis paraconfusus Ferreira, 1976 | | | Onitis paradoxus (Boheman, 1857) | ⇒ Tropidonitis paradoxus Boheman, 1857 | | Onitis parainflaticollis Ferreira, 1977 | | | Onitis paramniszechi Ferreira, 1976 | | | Onitis parvulus Balthasar, 1963 | | | Onitis pecuarius Lansberge, 1875 | | | Onitis perbrincki Ferreira, 1958 | | | Onitis perpunctatus Balthasar, 1963 | | | Onitis perturbator Péringuey, 1901 | | | Onitis picticollis Boheman, 1857 | | | Onitis pinheyi Ferreira, 1977 | | | Onitis podicinus Boucomont, 1932 | | | Onitis politus Lansberge, 1875 | | | Onitis popei Ferreira, 1977 | | | Onitis proximus Ferreira, 1977 | | | Onitis pseudojansenii Krikken, 1977 | | | Onitis pseudoorthopus Ferreira, 1976 | | | Onitis pseudosetosus Ferreira, 1976 | | | Onitis pumilio Balthasar, 1963 | | | Onitis pyramus Gillet, 1933 | | | Onitis reichei Lansberge, 1875 | | | Onitis retrodentatus Gillet, 1932 | | | Onitis ringenbachi Keith, 2007 | | | Onitis robustus Boheman, 1857 | | | Onitis rothi Lansberge, 1875 | | | Onitis senegalensis Lansberge, 1875 | | | Onitis setosus Lansberge, 1875 | | | Onitis shoensis (Reiche, 1847) | ⇒ Onitis vicinus Lansberge, 1875 | | Onitis sibutensis Janssens, 1943 | | | Onitis similis Felsche, 1911 | | | Onitis sphinx Fabricius, 1775 | | | Onitis spinicrus Fairmaire, 1893 | | | Onitis spinipes Drury, 1770 | | | Onitis subcrenatus Kolbe, 1897 | | | Onitis sulcipennis Felsche, 1907 | | | Onitis sulcipennis naromorus Krikken, 1977 | | | Onitis sulcipennis sulcipennis | | | Onitis tanzaniensis Ferreira, 1977 | | | Onitis thalassinus Lansberge, 1875 | | | Onitis thoracicus Ferreira, 1976 | | | Onitis tortuosus Houston, 1983 | | | Onitis trochantericus Ferreira, 1976 | | | Onitis tuberculifrons Josso, 2012 | | | Onitis tumidus Gillet, 1918 | | | Onitis uncinatoides Janssens, 1951 | | | Onitis uncinatus Klug, 1855 | | | Onitis unguiculatus Olivier, 1789 | | | Onitis upembanus Janssens, 1951 | | | Onitis vanderkelleni Lansberge, 1886 | | | Onitis vicinus Lansberge, 1875 | | | Onitis violaceus (Lansberge, 1875) | ⇒ Onitis humerosus Pallas, 1771 | | Onitis virens Lansberge, 1875 | | | Onitis viridulus (Boheman, 1857) | ⇒ Onitis pecuarius Lansberge, 1875 | | Onitis vrydaghi Janssens, 1937 | | | Onitis westermanni Lansberge, 1886 | | | Onitis wittei Janssens, 1938 | | | Onitis zambiensis Josso, 2012 | | | Ontherus azteca Harold, 1869 | | | Ontherus lunicollis Génier, 1996 | | | Ontherus pseudodidymus Génier, 1996 | | | Onthophagus (Afrostrandius) adspersus d´Orbigny, 1908 | | | Onthophagus (Afrostrandius) bimetallicus d´Orbigny, 1907 | | | Onthophagus (Afrostrandius) demeyeri Moretto, 2008 | | | Onthophagus (Afrostrandius) granulipennis (Lansberge, 1886) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus (Afrostrandius) loxodontaphilus Moretto, 2008 | | | Onthophagus (Afrostrandius) plebejus Klug, 1855 | | | Onthophagus (Afrostrandius) pseudoplebejus Moretto, 2008 | | | Onthophagus (Afrostrandius) tigrinus (d´Orbigny, 1908) | ⇒ Onthophagus pygmaeus Schaller, 1783 | | Onthophagus tigrinus kageraensis Moretto, 2009 | | | Onthophagus tigrinus tigrinus | | | Onthophagus (Amphionthophagus) melitaeus Fabricius, 1798 | | | Onthophagus (Digitonthophagus) gazella (Fabricius, 1787) | ⇒ Digitonthophagus gazella Fabricius, 1787 | | Onthophagus (Euonthophagus) carbonarius Klug, 1855 | | | Onthophagus (Euonthophagus) consobrinus Péringuey, 1901 | | | Onthophagus (Euonthophagus) crocatus Mulsant & Go..., 1870 | | | Onthophagus (Furconthophagus) furcatus (Fabricius, 1781) | ⇒ Onthophagus furcaticeps Masters, 1886 | | Onthophagus (Furconthophagus) monforti Josso & Prév..., 2006 | | | Onthophagus (Gibbonthophagus) privus Kabakov, 1998 | | | Onthophagus (Hyalonthophagus) hyalcyon Palestrini &..., 1989 | | | Onthophagus (Macronthophagus) uenoi Ochi, 1995 | | | Onthophagus (Onthophagus) aciculatulus Blatchley, 1928 | | | Onthophagus (Onthophagus) alternans (Raffray, 1877) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus (Onthophagus) batesi Howden and C..., 1963 | | | Onthophagus (Onthophagus) cyanellus Bates, 1887 | | | Onthophagus (Onthophagus) depressus Harold, 1871 | | | Onthophagus (Onthophagus) illyricus Scopoli, 1763 | | | Onthophagus (Onthophagus) incensus Say, 1835 | | | Onthophagus (Onthophagus) landolti (Harold, 1880) | ⇒ Onthophagus schaefferi Howden and C..., 1963 | | Onthophagus (Onthophagus) maculatus Fabricius, 1801 | | | Onthophagus (Onthophagus) margaritifer d´Orbigny, 1898 | | | Onthophagus (Onthophagus) nuchicornis Linnaeus, 1758 | | | Onthophagus (Onthophagus) orientalis (Harold, 1868) | ⇒ Onthophagus fissinasus Fairmaire, 1895 | | Onthophagus (Onthophagus) seabrai Gomes Alves, 1944 | | | Onthophagus (Onthophagus) subalternans d´Orbigny, 1902 | | | Onthophagus (Onthophagus) taurus Schreber, 1759 | Oechslein | | Onthophagus (Onthophagus) trituber Wiedemann, 1823 | | | Onthophagus (Palaeonthophagus) eulaminicornis Pittino, 2006 | | | Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790) | ⇒ Onthophagus similis Scriba, 1790 | | Onthophagus (Palaeonthophagus) latigena d´Orbigny, 1897 | | | Onthophagus (Palaeonthophagus) lemur Fabricius, 1781 | | | Onthophagus marginalis przewalski Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) opacicollis Reitter, 1893 | | | Onthophagus (Palaeonthophagus) petrovitzi (Frey, 1975) | ⇒ Onthophagus petrovitzianus Pittino, 2006 | | Onthophagus (Palaeonthophagus) petrovitzianus Pittino, 2006 | | | Onthophagus (Palaeonthophagus) ponticus Harold, 1883 | | | Onthophagus (Palaeonthophagus) semicornis Panzer, 1798 | Halbhörniger Kotkäfer | | Onthophagus (Palaeonthophagus) similis (Scriba, 1790) | ⇒ Onthophagus fracticornis Preyssler, 1790 | | Onthophagus (Palaeonthophagus) suturellus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) trigibber Reitter, 1893 | | | Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767 | Kühlein | | Onthophagus (Palaeonthophagus) viridis Ménétriés, 1832 | | | Onthophagus (Paleonthophagus) andalusicus Waltl, 1835 | | | Onthophagus andalusicus andalusicus | | | Onthophagus (Paleonthophagus) fracticornis Preyssler, 1790 | | | Onthophagus (Paleonthophagus) similis Scriba, 1790 | | | Onthophagus (Parentius) atricapillus d´Orbigny, 1908 | | | Onthophagus (Parentius) emarginatus Mulsant, 1842 | | | Onthophagus (Phanaeomorphus) ater Waterhouse, 1875 | | | Onthophagus (Proagoderus) alcedo d´Orbigny, 1913 | | | Onthophagus (Proagoderus) alcyon (Klug, 1855) | ⇒ Onthophagus alcedo d´Orbigny, 1913 | | Onthophagus (Proagoderus) alcyonides d´Orbigny, 1913 | | | Onthophagus (Proagoderus) aureiceps d´Orbigny, 1902 | | | Onthophagus (Proagoderus) brucei Reiche, 1847 | | | Onthophagus brucei cyanopterus (d´Orbigny, 1905) | ⇒ Proagoderus cyanopterus d´Orbigny, 1905 | | Onthophagus (Proagoderus) dives Harold, 1877 | | | Onthophagus (Proagoderus) fossidorsis d´Orbigny, 1905 | | | Onthophagus (Proagoderus) mixtifrons d´Orbigny, 1910 | | | Onthophagus (Proagoderus) pegesimallus Marcus, 1917 | | | Onthophagus (Proagoderus) pseudoalcyon d´Orbigny, 1913 | | | Onthophagus (Proagoderus) pseudovirens d´Orbigny, 1913 | | | Onthophagus (Proagoderus) viridiceps d´Orbigny, 1910 | | | Onthophagus (Relictonthophagus) nigellus (Illiger, 1803) | ⇒ Caccobius histeroides Ménétriés, 1832 | | Onthophagus (Relictonthophagus) punctatus (Illiger, 1803) | ⇒ Onthophagus emarginatus Mulsant, 1842 | | Onthophagus (Serrophorus) sagittarius Fabricius, 1775 | | | Onthophagus (Strandius) obliquus Olivier, 1789 | | | Onthophagus obliquus shimba (Cambefort, 1980) | ⇒ Onthophagus shimba Cambefort, 1980 | | Onthophagus (Trichonthophagus) hirtus Illiger, 1803 | | | Onthophagus (Trichonthophagus) maki Illiger, 1803 | | | Onthophagus abas Balthasar, 1946 | | | Onthophagus abeillei d´Orbigny, 1897 | | | Onthophagus abruptus d´Orbigny, 1913 | | | Onthophagus absyrtus Balthasar, 1946 | | | Onthophagus abyssinicus Gillet, 1925 | | | Onthophagus aceroides Krikken & Hu..., 2013 | | | Onthophagus aciculatus (Fahraeus, 1857) | ⇒ Onthophagus aciculatulus Blatchley, 1928 | | Onthophagus acuminatus (Harold, 1880) | ⇒ Onthophagus embrikianus Paulian, 1936 | | Onthophagus acutus d´Orbigny, 1908 | | | Onthophagus admetus Balthasar, 1946 | | | Onthophagus adornatus d´Orbigny, 1904 | | | Onthophagus aegrotus Balthasar, 1967 | | | Onthophagus aeneoniger d´Orbigny, 1913 | | | Onthophagus aeneopiceus d´Orbigny, 1902 | | | Onthophagus aequatus Péringuey, 1901 | | | Onthophagus aequepubens d´Orbigny, 1905 | | | Onthophagus aerarius Reitter, 1892 | | | Onthophagus aereidorsis d´Orbigny, 1902 | | | Onthophagus aeremicans d´Orbigny, 1904 | | | Onthophagus aerestriatus d´Orbigny, 1913 | | | Onthophagus aeruginosus Roth, 1851 | | | Onthophagus aesopus Lansberge, 1882 | | | Onthophagus aethiopicus d´Orbigny, 1902 | | | Onthophagus africanus Lansberge, 1886 | | | Onthophagus aguatus | | | Onthophagus ahenicollis d´Orbigny, 1902 | | | Onthophagus ahenomicans d´Orbigny, 1902 | | | Onthophagus akamineae Moretto & Gé..., 2020 | | | Onthophagus alainmonforti Josso, 2022 | | | Onthophagus albicomus d´Orbigny, 1908 | | | Onthophagus albipennis Péringuey, 1908 | | | Onthophagus albipodex d´Orbigny, 1902 | | | Onthophagus aliceae Josso, 2019 | | | Onthophagus alienus Frey, 1958 | | | Onthophagus alluaudi d´Orbigny, 1902 | | | Onthophagus altidorsis d´Orbigny, 1905 | | | Onthophagus altilamina d´Orbigny, 1905 | | | Onthophagus ambiguus (Péringuey, 1901) | ⇒ Mimonthophagus ambiguus Péringuey, 1901 | | Onthophagus ambiquus | | | Onthophagus amoenus d´Orbigny, 1908 | | | Onthophagus amphicoma Boucomont, 1914 | | | Onthophagus amplipennis d´Orbigny, 1905 | | | Onthophagus andersoni Howden & Gill, 1987 | | | Onthophagus androgynus d´Orbigny, 1905 | | | Onthophagus angolensis Paulian, 1937 | | | Onthophagus angularis d´Orbigny, 1908 | | | Onthophagus angulicornis (Mulsant, 1842) | ⇒ Onthophagus semicornis Panzer, 1798 | | Onthophagus annoyeri Moretto, 2010 | | | Onthophagus anogeissii Cambefort, 1984 | | | Onthophagus anomalicollis Frey, 1973 | | | Onthophagus anomalipes d´Orbigny, 1902 | | | Onthophagus antecornutus Josso, 2019 | | | Onthophagus antennalis Frey, 1961 | | | Onthophagus antilocapra Balthasar, 1937 | | | Onthophagus antoinei Walter, 1990 | | | Onthophagus antoineianus Josso, 2022 | | | Onthophagus apiciosus d´Orbigny, 1902 | | | Onthophagus apicosus | | | Onthophagus arcifer d´Orbigny, 1904 | | | Onthophagus areolatus d´Orbigny, 1905 | | | Onthophagus aries Kirby, 1825 | | | Onthophagus aschenborni Frey, 1975 | | | Onthophagus ashanticola Frey, 1973 | | | Onthophagus asimilis Péringuey, 1901 | | | Onthophagus asperatus d´Orbigny, 1905 | | | Onthophagus aspericeps d´Orbigny, 1908 | | | Onthophagus asperipennis d´Orbigny, 1902 | | | Onthophagus aspernatus d´Orbigny, 1907 | | | Onthophagus asperrimus d´Orbigny, 1902 | | | Onthophagus asperulus d´Orbigny, 1905 | | | Onthophagus astigma d´Orbigny, 1904 | | | Onthophagus athiensis d´Orbigny, 1913 | | | Onthophagus atricolor d´Orbigny, 1907 | | | Onthophagus atridorsis d´Orbigny, 1902 | | | Onthophagus atriglabrus Howden & Gill, 1987 | | | Onthophagus atroaereus d´Orbigny, 1908 | | | Onthophagus atrofasciatus d´Orbigny, 1905 | | | Onthophagus atronitidus d´Orbigny, 1902 | | | Onthophagus atrosplendens Frey, 1961 | | | Onthophagus atrostriatus d´Orbigny, 1913 | | | Onthophagus atrovirens d´Orbigny, 1905 | | | Onthophagus atrovittatus d´Orbigny, 1908 | | | Onthophagus auriculatus (Klug, 1855) | ⇒ Onthophagus latestriatus d´Orbigny, 1908 | | Onthophagus auritus Erichson, 1842 | | | Onthophagus axillaris (Boheman, 1860) | ⇒ Onthophagus criniger d´Orbigny, 1904 | | Onthophagus babaulti d´Orbigny, 1915 | | | Onthophagus bakweri Moretto, 2014 | | | Onthophagus baloghi Balthasar, 1967 | | | Onthophagus bandamai Cambefort, 1984 | | | Onthophagus baoule Cambefort, 1984 | | | Onthophagus barriorum Walter, 1990 | | | Onthophagus bartosi Balthasar, 1966 | | | Onthophagus basakata Walter & Cam..., 1977 | | | Onthophagus basilewskyi Frey, 1961 | | | Onthophagus bateke Walter & Cam..., 1977 | | | Onthophagus bayeri Balthasar, 1942 | | | Onthophagus bechynei Frey, 1953 | | | Onthophagus beiranus Péringuey, 1908 | | | Onthophagus belinga Walter, 1989 | | | Onthophagus bellus d´Orbigny, 1905 | | | Onthophagus benedictorum Walter & Cam..., 1977 | | | Onthophagus benguellianus Paulian, 1937 | | | Onthophagus bequaerti d´Orbigny, 1915 | | | Onthophagus bergeri Frey, 1975 | | | Onthophagus bernaudi Cambefort & ..., 1991 | | | Onthophagus betschuanus Frey, 1975 | | | Onthophagus bicallifrons d´Orbigny, 1902 | | | Onthophagus bicallosus Klug, 1855 | | | Onthophagus bicarinatus Gomes Alves, 1944 | | | Onthophagus bicavifrons d´Orbigny, 1902 | | | Onthophagus bicolor (Raffray, 1877) | ⇒ Onthophagus flavolimbatus Klug, 1855 | | Onthophagus biconifer d´Orbigny, 1905 | | | Onthophagus bicristatus d´Orbigny, 1905 | | | Onthophagus bicristiger d´Orbigny, 1913 | | | Onthophagus bidens Olivier, 1789 | | | Onthophagus bidentatus (Drapiez, 1819) | ⇒ Caccobius schreberi Linnaeus, 1767 | | Onthophagus bidentifrons d´Orbigny, 1902 | | | Onthophagus bifidicornis d´Orbigny, 1902 | | | Onthophagus bifidus Reiche, 1849 | | | Onthophagus bifrons d´Orbigny, 1905 | | | Onthophagus bilingula Balthasar, 1965 | | | Onthophagus bimarginatus d´Orbigny, 1902 | | | Onthophagus binodis Thunberg, 1818 | | | Onthophagus binodosus d´Orbigny, 1908 | | | Onthophagus binodulus d´Orbigny, 1913 | | | Onthophagus biplagiatus Thomson, 1858 | | | Onthophagus birugatus d´Orbigny, 1902 | | | Onthophagus birugifer d´Orbigny, 1908 | | | Onthophagus bisbicornis d´Orbigny, 1909 | | | Onthophagus bisignatus d´Orbigny, 1913 | | | Onthophagus bituber d´Orbigny, 1904 | | | Onthophagus bituberans d´Orbigny, 1905 | | | Onthophagus bituberculatus (Olivier, 1789) | ⇒ Onthophagus acutus d´Orbigny, 1908 | | Onthophagus bituberifrons Josso, 2018 | | | Onthophagus blanchardi Harold, 1869 | | | Onthophagus bocandei d´Orbigny, 1904 | | | Onthophagus boops d´Orbigny, 1905 | | | Onthophagus borassi Cambefort, 1984 | | | Onthophagus bordati Moretto, 2014 | | | Onthophagus bovimus | | | Onthophagus bovinus Péringuey, 1892 | | | Onthophagus brazzavillianus Balthasar, 1974 | | | Onthophagus breviceps d´Orbigny, 1902 | | | Onthophagus brevigena d´Orbigny, 1902 | | | Onthophagus brevipennis d´Orbigny, 1902 | | | Onthophagus brevisetis d´Orbigny, 1915 | | | Onthophagus brittoni Paulian, 1948 | | | Onthophagus brunellii Müller, 1942 | | | Onthophagus brutus Arrow, 1931 | | | Onthophagus bubalus Harold, 1867 | | | Onthophagus bufonidius Josso, 2019 | | | Onthophagus burchelli d´Orbigny, 1908 | | | Onthophagus caelator Balthasar, 1967 | | | Onthophagus caffrarius d´Orbigny, 1902 | | | Onthophagus calchas Balthasar, 1967 | | | Onthophagus calliger d´Orbigny, 1913 | | | Onthophagus callosipennis Boucomont, 1930 | | | Onthophagus cameloides d´Orbigny, 1900 | | | Onthophagus camerunicus d´Orbigny, 1905 | | | Onthophagus camilleae Josso, 2022 | | | Onthophagus cancer Lansberge, 1886 | | | Onthophagus carayoni Cambefort, 1986 | | | Onthophagus carcharias Harold, 1875 | | | Onthophagus carinicollis Raffray, 1877 | | | Onthophagus carinidorsis d´Orbigny, 1908 | | | Onthophagus carinifer d´Orbigny, 1904 | | | Onthophagus carpophilus Pereira & Ha..., 1961 | | | Onthophagus catherineae Josso, 2022 | | | Onthophagus catta Fabricius, 1787 | | | Onthophagus cavifrons Harold, 1886 | | | Onthophagus cavivertex d´Orbigny, 1913 | | | Onthophagus centricornis Fabricius, 1798 | | | Onthophagus cephalophi Cambefort, 1984 | | | Onthophagus cervus (Fabricius, 1798) | ⇒ Onthophagus falsus Gillet, 1925 | | Onthophagus chalcostolus d´Orbigny, 1902 | | | Onthophagus championi Bates, 1887 | | | Onthophagus chirindaensis Josso, 2022 | | | Onthophagus chirindanus d´Orbigny, 1913 | | | Onthophagus chitipaensis Josso, 2019 | | | Onthophagus chloroderus d´Orbigny, 1905 | | | Onthophagus chlorophanus d´Orbigny, 1902 | | | Onthophagus choanicus d´Orbigny, 1904 | | | Onthophagus chryses Bates, 1887 | | | Onthophagus chrysoderus d´Orbigny, 1905 | | | Onthophagus cicer Balthasar, 1964 | | | Onthophagus cincticollis d´Orbigny, 1902 | | | Onthophagus cinctifrons d´Orbigny, 1902 | | | Onthophagus cinctipennis Quedenfeldt, 1884 | | | Onthophagus cineraceus d´Orbigny, 1902 | | | Onthophagus circumdatus d´Orbigny, 1913 | | | Onthophagus civettae Cambefort, 1984 | | | Onthophagus clavisetis d´Orbigny, 1905 | | | Onthophagus clitellarius d´Orbigny, 1908 | | | Onthophagus clusifrons d´Orbigny, 1905 | | | Onthophagus coiffaiti Frey, 1970 | | | Onthophagus colasi Paulian, 1937 | | | Onthophagus comatulus d´Orbigny, 1905 | | | Onthophagus concavifrons d´Orbigny, 1904 | | | Onthophagus confertus Péringuey, 1908 | | | Onthophagus confluens (d´Orbigny, 1902) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus confusus Boucomont, 1932 | | | Onthophagus conradsi Balthasar, 1937 | | | Onthophagus contiguicornis d´Orbigny, 1913 | | | Onthophagus convexipennis Dierkens, 2022 | | | Onthophagus convexus d´Orbigny, 1908 | | | Onthophagus coptorhinodes Péringuey, 1901 | | | Onthophagus coptorhinoides Péringuey, 1901 | | | Onthophagus corniculatus (Reiche, 1847) | ⇒ Onthophagus senegalensis d´Orbigny, 1902 | | Onthophagus corniculiger d´Orbigny, 1913 | | | Onthophagus cornifrons Thomson, 1858 | | | Onthophagus cornutus Ferreira, 1971 | | | Onthophagus coronatus d´Orbigny, 1902 | | | Onthophagus coscineus Bates, 1887 | | | Onthophagus costatus d´Orbigny, 1913 | | | Onthophagus costiger d´Orbigny, 1915 | | | Onthophagus costilatus d´Orbigny, 1908 | | | Onthophagus costulicollis Frey, 1975 | | | Onthophagus crantor Balthasar, 1967 | | | Onthophagus crassicollis Boucomont, 1913 | | | Onthophagus creber d´Orbigny, 1905 | | | Onthophagus cretus (Péringuey, 1901) | ⇒ Onthophagus setosus Fahraeus, 1857 | | Onthophagus cribellum d´Orbigny, 1902 | | | Onthophagus cribripennis d´Orbigny, 1902 | | | Onthophagus criniger d´Orbigny, 1904 | | | Onthophagus crinitus (Harold, 1869) | ⇒ Onthophagus criniger d´Orbigny, 1904 | | Onthophagus cristatus d´Orbigny, 1905 | | | Onthophagus croesulus Bates, 1888 | | | Onthophagus crucenotatus d´Orbigny, 1905 | | | Onthophagus cruentatus Klug, 1855 | | | Onthophagus cuniculus MacLeay, 1864 | | | Onthophagus cupreopinguis Josso, 2022 | | | Onthophagus cupreovirens d´Orbigny, 1904 | | | Onthophagus cupreus Harold, 1880 | | | Onthophagus cupricollis Péringuey, 1888 | | | Onthophagus curtipilis d´Orbigny, 1905 | | | Onthophagus curtulus Frey, 1961 | | | Onthophagus curvifrons d´Orbigny, 1906 | | | Onthophagus curvilamina Müller, 1942 | | | Onthophagus cyaneiceps d´Orbigny, 1913 | | | Onthophagus cyaneoniger d´Orbigny, 1902 | | | Onthophagus cyanochlorus d´Orbigny, 1902 | | | Onthophagus davisi Frey, 1975 | | | Onthophagus debilis d´Orbigny, 1905 | | | Onthophagus decedens Péringuey, 1904 | | | Onthophagus decedeus | | | Onthophagus decens Balthasar, 1964 | | | Onthophagus declivicollis d´Orbigny, 1902 | | | Onthophagus declivis Harold, 1869 | | | Onthophagus decolor d´Orbigny, 1902 | | | Onthophagus decoratus d´Orbigny, 1905 | | | Onthophagus decorsei d´Orbigny, 1908 | | | Onthophagus dedecor Wallengren, 1881 | | | Onthophagus deflexus d´Orbigny, 1908 | | | Onthophagus delahayei Josso, 2011 | | | Onthophagus delicatulus Raffray, 1877 | | | Onthophagus delicatus d´Orbigny, 1915 | | | Onthophagus delphinensis d´Orbigny, 1913 | | | Onthophagus densatus Frey, 1963 | | | Onthophagus densepunctatus Frey, 1963 | | | Onthophagus densipilis d´Orbigny, 1902 | | | Onthophagus denticulatus d´Orbigny, 1902 | | | Onthophagus denudatus d´Orbigny, 1902 | | | Onthophagus denudatus denudatus | | | Onthophagus denudatus endroedii Frey, 1973 | | | Onthophagus depilatus d´Orbigny, 1913 | | | Onthophagus depilis d´Orbigny, 1902 | | | Onthophagus deplanatus Lansberge, 1883 | | | Onthophagus depressifrons d´Orbigny, 1915 | | | Onthophagus depressipennis Cambefort, 1975 | | | Onthophagus derasus d´Orbigny, 1902 | | | Onthophagus desaegeri Frey, 1961 | | | Onthophagus deterrens Péringuey, 1901 | | | Onthophagus diadematus d´Orbigny, 1902 | | | Onthophagus dicax Balthasar, 1966 | | | Onthophagus dicella Bates, 1888 | | | Onthophagus dilutus d´Orbigny, 1905 | | | Onthophagus dinoderus d´Orbigny, 1913 | | | Onthophagus discovirens d´Orbigny, 1902 | | | Onthophagus discretus Péringuey, 1901 | | | Onthophagus dispar Péringuey, 1901 | | | Onthophagus distichus Roth, 1851 | | | Onthophagus ditus (Péringuey, 1901) | ⇒ Onthophagus fungicola d´Orbigny, 1908 | | Onthophagus diversus Reiche, 1847 | | | Onthophagus dlabolai Balthasar, 1964 | | | Onthophagus dohertyi d´Orbigny, 1902 | | | Onthophagus dorsuosus d´Orbigny, 1902 | | | Onthophagus drumonti Moretto, 1999 | | | Onthophagus ducorpsi d´Orbigny, 1913 | | | Onthophagus dudleyianus Josso, 2019 | | | Onthophagus dunningi Harold, 1869 | | | Onthophagus duvivieri d´Orbigny, 1904 | | | Onthophagus ebenicolor d´Orbigny, 1902 | | | Onthophagus ebenus Péringuey, 1888 | | | Onthophagus eberus | | | Onthophagus eburneus d´Orbigny, 1913 | | | Onthophagus echinus Boucomont, 1914 | | | Onthophagus ecopas Josso & Prév..., 2006 | | | Onthophagus elegans Klug, 1832 | | | Onthophagus emeritus Péringuey, 1901 | | | Onthophagus endroedianus Walter & Cam..., 1977 | | | Onthophagus enuguensis Frey, 1975 | | | Onthophagus epilamprus Bates, 1888 | | | Onthophagus erectinasus d´Orbigny, 1902 | | | Onthophagus escalerai d´Orbigny, 1902 | | | Onthophagus estelleae Josso, 2022 | | | Onthophagus euzeti Walter & Cam..., 1977 | | | Onthophagus evae Balthasar, 1944 | | | Onthophagus exasperatus (Gerstaecker, 1871) | ⇒ Onthophagus latissimus d´Orbigny, 1902 | | Onthophagus excisiceps d´Orbigny, 1902 | | | Onthophagus exiguus (Raffray, 1877) | ⇒ Onthophagus rubens d´Orbigny, 1902 | | Onthophagus exilis d´Orbigny, 1913 | | | Onthophagus extensicollis d´Orbigny, 1907 | | | Onthophagus extensus Harold, 1878 | | | Onthophagus extraneus Josso, 2021 | | | Onthophagus falcarius Balthasar, 1966 | | | Onthophagus fallaciosus Raffray, 1877 | | | Onthophagus fallax d´Orbigny, 1913 | | | Onthophagus fasciculiger d´Orbigny, 1902 | | | Onthophagus fasciolatus Boucomont, 1924 | | | Onthophagus feai d´Orbigny, 1905 | | | Onthophagus filicornis Harold, 1873 | | | Onthophagus fimetarius Roth, 1851 | | | Onthophagus fissinasus Fairmaire, 1895 | | | Onthophagus fitiniensis Cambefort, 1984 | | | Onthophagus flavibasis d´Orbigny, 1904 | | | Onthophagus flaviclava d´Orbigny, 1902 | | | Onthophagus flavimargo d´Orbigny, 1902 | | | Onthophagus flavipennis d´Orbigny, 1905 | | | Onthophagus flavipleurus Dierkens, 2022 | | | Onthophagus flavocinctus Frey, 1975 | | | Onthophagus flavolimbatus Klug, 1855 | | | Onthophagus flavorufus d´Orbigny, 1904 | | | Onthophagus flexicornis d´Orbigny, 1902 | | | Onthophagus flexifrons d´Orbigny, 1908 | | | Onthophagus foliaceus Lansberge, 1886 | | | Onthophagus foliiceps Quedenfeldt, 1884 | | | Onthophagus foraminosus d´Orbigny, 1902 | | | Onthophagus fossibasis d´Orbigny, 1908 | | | Onthophagus fossifrons d´Orbigny, 1902 | | | Onthophagus fossulatus d´Orbigny, 1908 | | | Onthophagus foulliouxi Cambefort, 1971 | | | Onthophagus foveatus Frey, 1957 | | | Onthophagus foveicollis Boucomont, 1930 | | | Onthophagus fradei Gomes Alves, 1956 | | | Onthophagus francoisi Josso, 2013 | | | Onthophagus fritschi d´Orbigny, 1902 | | | Onthophagus frontalis Raffray, 1877 | | | Onthophagus fugitivus Péringuey, 1901 | | | Onthophagus fulvocinctus d´Orbigny, 1902 | | | Onthophagus fumatus d´Orbigny, 1913 | | | Onthophagus funestus Moretto & Gé..., 2010 | | | Onthophagus fungicola d´Orbigny, 1908 | | | Onthophagus fungus Josso, 2022 | | | Onthophagus furcaticeps Masters, 1886 | | | Onthophagus furcatoides (Lansberge, 1886) | ⇒ Onthophagus neofurcatus Goidanich, 1926 | | Onthophagus furculifer d´Orbigny, 1905 | | | Onthophagus fuscatus d´Orbigny, 1908 | | | Onthophagus fuscidorsis (d´Orbigny, 1902) | ⇒ Onthophagus fuscatus d´Orbigny, 1908 | | Onthophagus fuscivestis d´Orbigny, 1902 | | | Onthophagus gabonensis Walter, 1989 | | | Onthophagus gaillardi d´Orbigny, 1911 | | | Onthophagus galeatus Boucomont, 1919 | | | Onthophagus ganalensis Gestro, 1895 | | | Onthophagus garambae Frey, 1961 | | | Onthophagus gemellatus Raffray, 1877 | | | Onthophagus geminatus d´Orbigny, 1907 | | | Onthophagus geminifrons d´Orbigny, 1905 | | | Onthophagus georgwerneri Josso, 2022 | | | Onthophagus ghanensis Balthasar, 1966 | | | Onthophagus gibber d´Orbigny, 1905 | | | Onthophagus gibberosus d´Orbigny, 1905 | | | Onthophagus gibbidorsis d´Orbigny, 1902 | | | Onthophagus gibbifrons d´Orbigny, 1902 | | | Onthophagus gibbus d´Orbigny, 1913 | | | Onthophagus giraffa Hausmann, 1807 | | | Onthophagus girardinae Walter, 1989 | | | Onthophagus giuseppecarpanetoi Tagliaferri ..., 2012 | | | Onthophagus gnu Frey, 1955 | | | Onthophagus gonopygus d´Orbigny, 1902 | | | Onthophagus gradivus Balthasar, 1966 | | | Onthophagus grandidorsis d´Orbigny, 1913 | | | Onthophagus grandifrons d´Orbigny, 1907 | | | Onthophagus graniceps d´Orbigny, 1908 | | | Onthophagus granosus d´Orbigny, 1913 | | | Onthophagus granulatus (Boheman, 1858) | ⇒ Onthophagus consentaneus Harold, 1867 | | Onthophagus granulifer (Harold, 1886) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus granulifrons Frey, 1961 | | | Onthophagus granulopygus Dierkens, 2022 | | | Onthophagus granulum d´Orbigny, 1904 | | | Onthophagus graphicus Wallengren, 1881 | | | Onthophagus grassei Walter, 1989 | | | Onthophagus gravearmatus Balthasar, 1944 | | | Onthophagus gravoti d´Orbigny, 1908 | | | Onthophagus guttatus (Boheman, 1860) | ⇒ Onthophagus quadrilunatus d´Orbigny, 1902 | | Onthophagus guttiger d´Orbigny, 1905 | | | Onthophagus haafi Frey, 1963 | | | Onthophagus helciatus Harold, 1871 | | | Onthophagus hemichalceus d´Orbigny, 1915 | | | Onthophagus hemichlorus d´Orbigny, 1915 | | | Onthophagus henus | | | Onthophagus hericiniformis Moretto, 2010 | | | Onthophagus hericius d´Orbigny, 1902 | | | Onthophagus herus Péringuey, 1901 | | | Onthophagus heteroclitus d´Orbigny, 1911 | | | Onthophagus hilaridis Cambefort, 1984 | | | Onthophagus hilaris d´Orbigny, 1905 | | | Onthophagus hildebrandti Harold, 1878 | | | Onthophagus hinnulus (Klug, 1832) | ⇒ Mimonthophagus hinnulus Klug, 1832 | | Onthophagus hirsutus d´Orbigny, 1902 | | | Onthophagus hirtellus Frey, 1957 | | | Onthophagus hirticulus d´Orbigny, 1915 | | | Onthophagus hirtipodex d´Orbigny, 1904 | | | Onthophagus hopfneri | | | Onthophagus horrens d´Orbigny, 1908 | | | Onthophagus horridus d´Orbigny, 1908 | | | Onthophagus hulstaerti Boucomont, 1932 | | | Onthophagus humpatensis Lansberge, 1886 | | | Onthophagus hyaena Fabricius, 1801 | | | Onthophagus ieti Cambefort, 1984 | | | Onthophagus ikelengeensis Josso, 2018 | | | Onthophagus illotus Péringuey, 1901 | | | Onthophagus imbellis d´Orbigny, 1905 | | | Onthophagus imberbis d´Orbigny, 1902 | | | Onthophagus immundus Boheman, 1858 | | | Onthophagus importunus Péringuey, 1901 | | | Onthophagus impressicollis Boheman, 1860 | | | Onthophagus impunctatus d´Orbigny, 1904 | | | Onthophagus impurus Harold, 1868 | | | Onthophagus inaequalis d´Orbigny, 1902 | | | Onthophagus incantatus Balthasar, 1967 | | | Onthophagus incisus Harold, 1877 | | | Onthophagus includens d´Orbigny, 1905 | | | Onthophagus inclusus d´Orbigny, 1905 | | | Onthophagus indigus Péringuey, 1901 | | | Onthophagus indutus d´Orbigny, 1902 | | | Onthophagus ineptus Harold, 1871 | | | Onthophagus inermiceps d´Orbigny, 1905 | | | Onthophagus inermicollis d´Orbigny, 1908 | | | Onthophagus infaustus Cambefort, 1984 | | | Onthophagus inflatus Walter & Cam..., 1977 | | | Onthophagus infuscatus Klug, 1845 | | | Onthophagus insignis (Péringuey, 1896) | ⇒ Proagoderus fossidorsis d´Orbigny, 1905 | | Onthophagus insulsus Péringuey, 1901 | | | Onthophagus intermixtus d´Orbigny, 1902 | | | Onthophagus interruptus (Raffray, 1877) | ⇒ Onthophagus leanus Goidanich, 1926 | | Onthophagus interstitialis (Fahraeus, 1857) | ⇒ Euonthophagus carbonarius Klug, 1855 | | Onthophagus intonsus d´Orbigny, 1913 | | | Onthophagus intricatus Moretto, 2010 | | | Onthophagus investis d´Orbigny, 1904 | | | Onthophagus iringaensis Dierkens, 2022 | | | Onthophagus iulicola Cambefort, 1984 | | | Onthophagus jacksoni d´Orbigny, 1913 | | | Onthophagus janssensi Frey, 1958 | | | Onthophagus jeanneli d´Orbigny, 1913 | | | Onthophagus jeanneli diversiceps Frey, 1975 | | | Onthophagus jocquei Josso, 2019 | | | Onthophagus jonathani Moretto & Gé..., 2020 | | | Onthophagus jugicola d´Orbigny, 1902 | | | Onthophagus juheli Josso, 2022 | | | Onthophagus juncticornis d´Orbigny, 1908 | | | Onthophagus juniperensis Josso, 2019 | | | Onthophagus justei Walter, 1989 | | | Onthophagus juvencus Klug, 1835 | | | Onthophagus kaomaensis Josso, 2022 | | | Onthophagus kapangaensis Dierkens, 2022 | | | Onthophagus karlwerneri Josso, 2022 | | | Onthophagus kassaicus d´Orbigny, 1908 | | | Onthophagus katangensis Frey, 1961 | | | Onthophagus kavirondi (d´Orbigny, 1913) | ⇒ Onthophagus kavirondus d´Orbigny, 1915 | | Onthophagus kavirondus d´Orbigny, 1915 | | | Onthophagus kindianus Frey, 1953 | | | Onthophagus kingstoni Cambefort, 1980 | | | Onthophagus kirki Frey, 1975 | | | Onthophagus kirokanus Frey, 1975 | | | Onthophagus kivuensis Walter, 2018 | | | Onthophagus kochi Frey, 1957 | | | Onthophagus koma Matsumura, 1937 | | | Onthophagus kouassii Cambefort, 1984 | | | Onthophagus kudrnaianus Josso, 2022 | | | Onthophagus kulzeri Frey, 1958 | | | Onthophagus kyleensis Frey, 1975 | | | Onthophagus laceratus (Gerstaecker, 1871) | ⇒ Onthophagus depressus Harold, 1871 | | Onthophagus lacustris Harold, 1877 | | | Onthophagus laetus d´Orbigny, 1913 | | | Onthophagus laevatus d´Orbigny, 1902 | | | Onthophagus laeviceps d´Orbigny, 1902 | | | Onthophagus laevidorsis d´Orbigny, 1907 | | | Onthophagus laevissimus d´Orbigny, 1905 | | | Onthophagus lagnyi Paulian, 1937 | | | Onthophagus lamellicornis Frey, 1953 | | | Onthophagus lamelliger (Gerstaecker, 1871) | ⇒ Onthophagus ochropygus d´Orbigny, 1902 | | Onthophagus laminicornis (Lansberge, 1886) | ⇒ Onthophagus eulaminicornis Pittino, 2006 | | Onthophagus laminidorsis d´Orbigny, 1902 | | | Onthophagus laminosus d´Orbigny, 1905 | | | Onthophagus lamnifer d´Orbigny, 1902 | | | Onthophagus lamotellus Cambefort, 1980 | | | Onthophagus lamottei Frey, 1962 | | | Onthophagus lamtoensis Cambefort, 1984 | | | Onthophagus lamtoi Cambefort, 1980 | | | Onthophagus lanista (Castelnau, 1840) | ⇒ Proagoderus extensus Harold, 1878 | | Onthophagus lassulus Balthasar, 1964 | | | Onthophagus latepunctatus d´Orbigny, 1913 | | | Onthophagus latestriatus d´Orbigny, 1908 | | | Onthophagus latevittatus d´Orbigny, 1905 | | | Onthophagus latigibber d´Orbigny, 1902 | | | Onthophagus latipennis d´Orbigny, 1897 | | | Onthophagus latissimus d´Orbigny, 1902 | | | Onthophagus leanus Goidanich, 1926 | | | Onthophagus lefebvrei d´Orbigny, 1913 | | | Onthophagus lefiniensis Balthasar, 1967 | | | Onthophagus legendrei Walter & Cam..., 1977 | | | Onthophagus lemagneni d´Orbigny, 1902 | | | Onthophagus lemekensis d´Orbigny, 1915 | | | Onthophagus leroyi d´Orbigny, 1902 | | | Onthophagus leucopygus (Harold, 1867) | ⇒ Onthophagus albipodex d´Orbigny, 1902 | | Onthophagus liberianus Lansberge, 1883 | | | Onthophagus lilongweensis Dierkens, 2022 | | | Onthophagus liodermus d´Orbigny, 1913 | | | Onthophagus lioides d´Orbigny, 1902 | | | Onthophagus liopterus Harold, 1880 | | | Onthophagus lituratus Roth, 1851 | | | Onthophagus liwondeensis Josso, 2019 | | | Onthophagus lobi Cambefort, 1984 | | | Onthophagus lomii Müller, 1942 | | | Onthophagus longegranus Frey, 1957 | | | Onthophagus longiceps d´Orbigny, 1904 | | | Onthophagus longipilis d´Orbigny, 1905 | | | Onthophagus lopeensis Josso, 2022 | | | Onthophagus loroi Balthasar, 1941 | | | Onthophagus loudetiae Cambefort, 1984 | | | Onthophagus loveni Gillet, 1928 | | | Onthophagus loxodontae Cambefort, 1984 | | | Onthophagus lugens (Fahraeus, 1857) | ⇒ Onthophagus fimetarius Roth, 1851 | | Onthophagus lugubris (Fahraeus, 1857) | ⇒ Onthophagus lituratus Roth, 1851 | | Onthophagus lumareti Walter & Cam..., 1977 | | | Onthophagus lunatus Harold, 1868 | | | Onthophagus lutaticollis d´Orbigny, 1907 | | | Onthophagus mabuensis Daniel, Strü..., 2023 | | | Onthophagus macroliberianus Moretto, 2010 | | | Onthophagus macrothorax d´Orbigny, 1902 | | | Onthophagus maculosus d´Orbigny, 1908 | | | Onthophagus makokou Walter, 1989 | | | Onthophagus malevolus Balthasar, 1964 | | | Onthophagus malthinus Gillet, 1930 | | | Onthophagus mankonoensis (Balthasar, 1966) | ⇒ Onthophagus mocquerysi d´Orbigny, 1902 | | Onthophagus marahouensis Cambefort, 1984 | | | Onthophagus marginicollis Harold, 1880 | | | Onthophagus marginifer Frey, 1953 | | | Onthophagus marginipennis Josso, 2018 | | | Onthophagus marshalli d´Orbigny, 1908 | | | Onthophagus martaponsae | | | Onthophagus martelli Frey, 1972 | | | Onthophagus martellii Frey, 1972 | | | Onthophagus masaicus d´Orbigny, 1905 | | | Onthophagus matae Cambefort, 1984 | | | Onthophagus maxwellianus Moretto, 2013 | | | Onthophagus mediofuscatus d´Orbigny, 1908 | | | Onthophagus melanocephalus Klug, 1845 | | | Onthophagus menkaoensis Walter & Cam..., 1977 | | | Onthophagus merdrignaci Walter & Cam..., 1977 | | | Onthophagus meruanus d´Orbigny, 1908 | | | Onthophagus merus Péringuey, 1901 | | | Onthophagus metalliger d´Orbigny, 1913 | | | Onthophagus micros d´Orbigny, 1902 | | | Onthophagus miles d´Orbigny, 1905 | | | Onthophagus millingeni d´Orbigny, 1900 | | | Onthophagus mimus d´Orbigny, 1913 | | | Onthophagus minans d´Orbigny, 1913 | | | Onthophagus minax d´Orbigny, 1904 | | | Onthophagus minettianus Josso, 2019 | | | Onthophagus minutissimus d´Orbigny, 1908 | | | Onthophagus minutulus Harold, 1875 | | | Onthophagus minutus (Hausmann, 1807) | ⇒ Onthophagus centricornis Fabricius, 1798 | | Onthophagus miricollis Frey, 1962 | | | Onthophagus miricornis d´Orbigny, 1902 | | | Onthophagus mirifrons d´Orbigny, 1905 | | | Onthophagus miscellaneus d´Orbigny, 1908 | | | Onthophagus miscellus d´Orbigny, 1905 | | | Onthophagus misellus d´Orbigny, 1907 | | | Onthophagus mixticeps d´Orbigny, 1905 | | | Onthophagus mixtidorsis d´Orbigny, 1905 | | | Onthophagus mocquerysi d´Orbigny, 1902 | | | Onthophagus modestus Harold, 1862 | | | Onthophagus monardi (Boucomont, 1936) | ⇒ Onthophagus monardiellus Frey, 1963 | | Onthophagus monardiellus Frey, 1963 | | | Onthophagus monodon Fahraeus, 1857 | | | Onthophagus montivagus d´Orbigny, 1902 | | | Onthophagus montreuili Moretto & Gé..., 2010 | | | Onthophagus morenoi Moretto, 2013 | | | Onthophagus morettoi Josso, 2022 | | | Onthophagus morosus Gerstaecker, 1871 | | | Onthophagus mpassa Walter, 1982 | | | Onthophagus mucronatus (Thomson, 1858) | ⇒ Pseudosaproecius validicornis Quedenfeldt, 1884 | | Onthophagus mucronifer d´Orbigny, 1905 | | | Onthophagus mugheseensis Josso, 2019 | | | Onthophagus murphyi Josso, 2019 | | | Onthophagus musculus Frey, 1963 | | | Onthophagus musculus ganganensis Frey, 1963 | | | Onthophagus musculus musculus | | | Onthophagus mwenseensis Dierkens, 2022 | | | Onthophagus naevius d´Orbigny, 1913 | | | Onthophagus naevuliger d´Orbigny, 1908 | | | Onthophagus nanus (Harold, 1878) | ⇒ Onthophagus waterhousei Gillet, 1927 | | Onthophagus naso Fahraeus, 1857 | | | Onthophagus natronis Cambefort, 1984 | | | Onthophagus nchilaensis Josso, 2022 | | | Onthophagus neavei d´Orbigny, 1914 | | | Onthophagus nebulosus Reiche, 1864 | | | Onthophagus nemorivagus Kohlmann & S..., 2001 | | | Onthophagus neofurcatus Goidanich, 1926 | | | Onthophagus neostenocerus Goidanich, 1926 | | | Onthophagus neyo Moretto, 2013 | | | Onthophagus nigerianus d´Orbigny, 1913 | | | Onthophagus nigriceps Raffray, 1877 | | | Onthophagus nigricornis Fairmaire, 1887 | | | Onthophagus nigrinus Paulian, 1937 | | | Onthophagus nigripennis d´Orbigny, 1908 | | | Onthophagus nigriventris d´Orbigny, 1902 | | | Onthophagus nigrivestis (d´Orbigny, 1902) | ⇒ Onthophagus laeviceps d´Orbigny, 1902 | | Onthophagus nigroflavus Josso, 2022 | | | Onthophagus nigroviolaceus d´Orbigny, 1902 | | | Onthophagus nilicola d´Orbigny, 1902 | | | Onthophagus niloticus Harold, 1879 | | | Onthophagus nimbatus d´Orbigny, 1905 | | | Onthophagus niokolokoba Moretto, 2007 | | | Onthophagus nitidifrons d´Orbigny, 1905 | | | Onthophagus nitidulus Klug, 1845 | | | Onthophagus nodieri d´Orbigny, 1908 | | | Onthophagus nonstriatus d´Orbigny, 1905 | | | Onthophagus notatus d´Orbigny, 1902 | | | Onthophagus nudatus d´Orbigny, 1913 | | | Onthophagus nudus d´Orbigny, 1908 | | | Onthophagus numidicus d´Orbigny, 1908 | | | Onthophagus nummelini Moretto & Gé..., 2020 | | | Onthophagus nyctopus Bates, 1887 | | | Onthophagus nyikaensis Josso, 2018 | | | Onthophagus obenbergeri Balthasar, 1965 | | | Onthophagus obliguus | | | Onthophagus obtusicornis (Fahraeus, 1857) | ⇒ Onthophagus marginalis przewalski Kabakov, 2006 | | Onthophagus obtusus Wiedemann, 1819 | | | Onthophagus obtutus Péringuey, 1901 | | | Onthophagus ocellidorsis d´Orbigny, 1915 | | | Onthophagus ocellifer d´Orbigny, 1902 | | | Onthophagus ochreatus d´Orbigny, 1897 | | | Onthophagus ochropygus d´Orbigny, 1902 | | | Onthophagus okahandjanus Balthasar, 1974 | | | Onthophagus olidus Balthasar, 1964 | | | Onthophagus omostigma d´Orbigny, 1902 | | | Onthophagus ophtalmicus Frey, 1958 | | | Onthophagus ornatulus d´Orbigny, 1908 | | | Onthophagus orphnoides (Bates, 1887) | ⇒ Onthophagus mirabilis Bates, 1887 | | Onthophagus orthocerus Thomson, 1858 | | | Onthophagus otjivarongus Balthasar, 1967 | | | Onthophagus ovigranosus Frey, 1971 | | | Onthophagus ovulum Gerstaecker, 1871 | | | Onthophagus pallens d´Orbigny, 1908 | | | Onthophagus pallidipennis (Fahraeus, 1857) | ⇒ Onthophagus rasipennis d´Orbigny, 1908 | | Onthophagus pallidus d´Orbigny, 1905 | | | Onthophagus paravinctus Tagliaferri ..., 2012 | | | Onthophagus parceguttatus d´Orbigny, 1902 | | | Onthophagus parcenotatus d´Orbigny, 1913 | | | Onthophagus parcepictus d´Orbigny, 1908 | | | Onthophagus parcepilosus Balthasar, 1966 | | | Onthophagus parenthesis Boucomont, 1913 | | | Onthophagus parisii (Balthasar, 1941) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus parumnotatus Fahraeus, 1857 | | | Onthophagus parvifrons (d´Orbigny, 1907) | ⇒ Onthophagus semiasper d´Orbigny, 1902 | | Onthophagus paucigranosus Lansberge, 1886 | | | Onthophagus paulianellus Frey, 1958 | | | Onthophagus pauxillus d´Orbigny, 1902 | | | Onthophagus pendjarius Josso & Prév..., 2006 | | | Onthophagus peringueyi Shipp, 1895 | | | Onthophagus perniger Boucomont, 1930 | | | Onthophagus peyrierasi Paulian, 1985 | | | Onthophagus phanaeomorphus Janssens, 1954 | | | Onthophagus philippemorettoi Josso, 2022 | | | Onthophagus picatus d´Orbigny, 1902 | | | Onthophagus piceiceps d´Orbigny, 1904 | | | Onthophagus pictipennis d´Orbigny, 1913 | | | Onthophagus pictipodex d´Orbigny, 1902 | | | Onthophagus picturatus d´Orbigny, 1908 | | | Onthophagus pictus (Reitter, 1893) | ⇒ Oniticellus tessellatus Harold, 1879 | | Onthophagus pierrei Josso, 2022 | | | Onthophagus pilicollis d´Orbigny, 1902 | | | Onthophagus pilipodex d´Orbigny, 1913 | | | Onthophagus pilosus (Fahraeus, 1857) | ⇒ Onthophagus echinus Boucomont, 1914 | | Onthophagus pinguis Gerstaecker, 1871 | | | Onthophagus pipitzi Ancey, 1883 | | | Onthophagus pisciphagus Walter & Cam..., 1977 | | | Onthophagus placens Péringuey, 1904 | | | Onthophagus planaticeps d´Orbigny, 1913 | | | Onthophagus planiceps (MacLeay, 1886) | ⇒ Onthophagus pinguis Gerstaecker, 1871 | | Onthophagus planidorsis Dierkens, 2022 | | | Onthophagus pleurogonus d´Orbigny, 1913 | | | Onthophagus plicatifrons d´Orbigny, 1908 | | | Onthophagus politissimus d´Orbigny, 1908 | | | Onthophagus pollicatus Harold, 1879 | | | Onthophagus polyedrus d´Orbigny, 1905 | | | Onthophagus polyodon d´Orbigny, 1913 | | | Onthophagus polystigma d´Orbigny, 1902 | | | Onthophagus pooensis Cambefort & ..., 1991 | | | Onthophagus possoi Walter, 1982 | | | Onthophagus posticicornis d´Orbigny, 1913 | | | Onthophagus praecellens Bates, 1887 | | | Onthophagus praedatus Harold, 1862 | | | Onthophagus praelaminatus Frey, 1957 | | | Onthophagus praestans Péringuey, 1901 | | | Onthophagus praetortus Péringuey, 1904 | | | Onthophagus priscus | | | Onthophagus probus Péringuey, 1901 | | | Onthophagus producticollis d´Orbigny, 1908 | | | Onthophagus profanus Frey, 1958 | | | Onthophagus pronocarinatus Josso, 2019 | | | Onthophagus proteus d´Orbigny, 1902 | | | Onthophagus protuberans Frey, 1955 | | | Onthophagus proximus d´Orbigny, 1908 | | | Onthophagus pseudoaeneus d´Orbigny, 1908 | | | Onthophagus pseudobidens d´Orbigny, 1915 | | | Onthophagus pseudobufonidius Josso, 2019 | | | Onthophagus pseudocribripennis Josso, 2019 | | | Onthophagus pseudofimetarius Balthasar, 1946 | | | Onthophagus pseudoliberianus Moretto, 2010 | | | Onthophagus pseudopilosus Frey, 1958 | | | Onthophagus pseudosanguineus Walter & Cam..., 1977 | | | Onthophagus pseudosellatus Balthasar, 1964 | | | Onthophagus pseudovinctus Tagliaferri ..., 2012 | | | Onthophagus puberulus d´Orbigny, 1902 | | | Onthophagus pugionatus Fahraeus, 1857 | | | Onthophagus pulchellus d´Orbigny, 1905 | | | Onthophagus pullatus d´Orbigny, 1905 | | | Onthophagus pullus (Roth, 1851) | ⇒ Onthophagus parenthesis Boucomont, 1913 | | Onthophagus purifrons d´Orbigny, 1908 | | | Onthophagus pusio (Fahraeus, 1857) | ⇒ Onthophagus confertus Péringuey, 1908 | | Onthophagus pygmaeus Schaller, 1783 | | | Onthophagus pyramidalis Klug, 1855 | | | Onthophagus quadraticeps Harold, 1867 | | | Onthophagus quadricallosus d´Orbigny, 1902 | | | Onthophagus quadricuspis d´Orbigny, 1908 | | | Onthophagus quadrilunatus d´Orbigny, 1902 | | | Onthophagus quadrimaculatus Raffray, 1877 | | | Onthophagus quadrinodosus Fahraeus, 1857 | | | Onthophagus quadrinotatus d´Orbigny, 1905 | | | Onthophagus quetzalis Howden & Gill, 1993 | | | Onthophagus quiproquo Moretto & Gé..., 2010 | | | Onthophagus ragazzii d´Orbigny, 1904 | | | Onthophagus rasipennis d´Orbigny, 1908 | | | Onthophagus rectestriatus d´Orbigny, 1915 | | | Onthophagus rectilamina d´Orbigny, 1902 | | | Onthophagus rectorispauliani Cambefort, 1980 | | | Onthophagus renejeanneli Dierkens, 2022 | | | Onthophagus reticulatus d´Orbigny, 1902 | | | Onthophagus reticuliger Frey, 1960 | | | Onthophagus reticulundaticeps Josso, 2022 | | | Onthophagus rhinocerus (Gomes Alves, 1944) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus rhodesianus Frey, 1975 | | | Onthophagus rhynchophorus Péringuey, 1904 | | | Onthophagus rhyncophorus | | | Onthophagus robertminettii Josso, 2022 | | | Onthophagus rohwedderi Moretto, 2017 | | | Onthophagus rojkoffianus Josso, 2018 | | | Onthophagus rosemarieae Josso, 2022 | | | Onthophagus rosettae Frey, 1958 | | | Onthophagus rostrifer (Reitter, 1893) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus rotundatus d´Orbigny, 1905 | | | Onthophagus rotundibasis d´Orbigny, 1902 | | | Onthophagus rougonorum Cambefort, 1984 | | | Onthophagus ruandanus Frey, 1971 | | | Onthophagus rubefactus (d´Orbigny, 1913) | ⇒ Onthophagus tessulatus Harold, 1871 | | Onthophagus rubellus d´Orbigny, 1908 | | | Onthophagus rubens d´Orbigny, 1902 | | | Onthophagus rubenticollis d´Orbigny, 1902 | | | Onthophagus rubidus d´Orbigny, 1913 | | | Onthophagus rubricatus d´Orbigny, 1902 | | | Onthophagus rufipodex d´Orbigny, 1913 | | | Onthophagus rufobasalis Fairmaire, 1887 | | | Onthophagus rufocastaneus Fairmaire, 1887 | | | Onthophagus rufolimbatus d´Orbigny, 1913 | | | Onthophagus rufonotatus d´Orbigny, 1902 | | | Onthophagus rufopygus Frey, 1957 | | | Onthophagus rufostillans d´Orbigny, 1907 | | | Onthophagus rufovirens d´Orbigny, 1904 | | | Onthophagus rugidorsis d´Orbigny, 1913 | | | Onthophagus rugosipennis Frey, 1973 | | | Onthophagus rugosissimus d´Orbigny, 1913 | | | Onthophagus rugulipennis Fairmaire, 1887 | | | Onthophagus sanguineus d´Orbigny, 1902 | | | Onthophagus sanguinolentus d´Orbigny, 1908 | | | Onthophagus sansibaricus Harold, 1878 | | | Onthophagus sapphirinus (Fahraeus, 1857) | ⇒ Proagoderus aureiceps d´Orbigny, 1902 | | Onthophagus savanicola Cambefort, 1984 | | | Onthophagus scaber Roth, 1851 | | | Onthophagus scaberrimus d´Orbigny, 1904 | | | Onthophagus scapularis d´Orbigny, 1902 | | | Onthophagus schaufussi Harold, 1867 | | | Onthophagus sculptilis Gerstaecker, 1871 | | | Onthophagus secundarius Roth, 1851 | | | Onthophagus sellatulus d´Orbigny, 1902 | | | Onthophagus sellatus Klug, 1845 | | | Onthophagus semiaratus d´Orbigny, 1902 | | | Onthophagus semiasper d´Orbigny, 1902 | | | Onthophagus semichalcites d´Orbigny, 1902 | | | Onthophagus semicroceus d´Orbigny, 1915 | | | Onthophagus semiflavus (Boheman, 1860) | ⇒ Onthophagus flavolimbatus Klug, 1855 | | Onthophagus semigraniger d´Orbigny, 1905 | | | Onthophagus semilaevis d´Orbigny, 1913 | | | Onthophagus seminitens d´Orbigny, 1902 | | | Onthophagus seminitidus d´Orbigny, 1908 | | | Onthophagus semivestitus Paulian, 1937 | | | Onthophagus semivirescens d´Orbigny, 1902 | | | Onthophagus semiviridis d´Orbigny, 1904 | | | Onthophagus senegalensis d´Orbigny, 1902 | | | Onthophagus senescens Péringuey, 1908 | | | Onthophagus sericans Frey, 1975 | | | Onthophagus sericeicollis Boucomont, 1928 | | | Onthophagus serienotatus d´Orbigny, 1902 | | | Onthophagus setosus Fahraeus, 1857 | | | Onthophagus shimba Cambefort, 1980 | | | Onthophagus sidama Gestro, 1895 | | | Onthophagus signatus Fahraeus, 1857 | | | Onthophagus simillimus d´Orbigny, 1913 | | | Onthophagus simoni d´Orbigny, 1902 | | | Onthophagus simplex Raffray, 1877 | | | Onthophagus simpliciceps d´Orbigny, 1905 | | | Onthophagus simulator d´Orbigny, 1905 | | | Onthophagus sinuosicollis d´Orbigny, 1905 | | | Onthophagus sinuosus d´Orbigny, 1913 | | | Onthophagus sipilouensis Cambefort, 1984 | | | Onthophagus sjoestethi d´Orbigny, 1904 | | | Onthophagus smeenki Cambefort, 1980 | | | Onthophagus snoflaki Balthasar, 1944 | | | Onthophagus solisi Howden & Gill, 1993 | | | Onthophagus somalicola Balthasar, 1941 | | | Onthophagus sophiae Nicolas, 2006 | | | Onthophagus splendidoides Josso, 2013 | | | Onthophagus splendidus Boucomont, 1932 | | | Onthophagus sprecherae Moretto, 2013 | | | Onthophagus spurcatus d´Orbigny, 1905 | | | Onthophagus stanleyi Moretto & Gé..., 2010 | | | Onthophagus stehliki Balthasar, 1966 | | | Onthophagus stellio Erichson, 1843 | | | Onthophagus stenocerus (Harold, 1867) | ⇒ Onthophagus neostenocerus Goidanich, 1926 | | Onthophagus sticticus Harold, 1867 | | | Onthophagus stictus d´Orbigny, 1913 | | | Onthophagus stigmosus d´Orbigny, 1902 | | | Onthophagus stillatus d´Orbigny, 1904 | | | Onthophagus stomachosus Krell, 2009 | | | Onthophagus strictestriatus d´Orbigny, 1913 | | | Onthophagus stuhlmanni d´Orbigny, 1908 | | | Onthophagus subcinctus d´Orbigny, 1913 | | | Onthophagus subdivisus d´Orbigny, 1908 | | | Onthophagus subhumeralis d´Orbigny, 1902 | | | Onthophagus subnudus d´Orbigny, 1902 | | | Onthophagus subocellatus d´Orbigny, 1905 | | | Onthophagus subplanus d´Orbigny, 1902 | | | Onthophagus subrugosus d´Orbigny, 1905 | | | Onthophagus subsulcatus d´Orbigny, 1908 | | | Onthophagus subulifer d´Orbigny, 1908 | | | Onthophagus suffusus Klug, 1855 | | | Onthophagus sugillatus Klug, 1855 | | | Onthophagus sulcatulus d´Orbigny, 1907 | | | Onthophagus sulcipennis d´Orbigny, 1902 | | | Onthophagus sumptuosus Paulian, 1937 | | | Onthophagus sutiliceps d´Orbigny, 1902 | | | Onthophagus suturalis (Péringuey, 1888) | ⇒ Onthophagus trituber Wiedemann, 1823 | | Onthophagus sylvestris Walter & Cam..., 1977 | | | Onthophagus synceri Cambefort, 1984 | | | Onthophagus taboranus d´Orbigny, 1902 | | | Onthophagus taiensis Cambefort, 1984 | | | Onthophagus talpa Fahraeus, 1857 | | | Onthophagus tanganus d´Orbigny, 1913 | | | Onthophagus tapirus Sharp, 1877 | | | Onthophagus teitanicus d´Orbigny, 1902 | | | Onthophagus temporalis d´Orbigny, 1902 | | | Onthophagus tenax Balthasar, 1964 | | | Onthophagus tenuigraniger d´Orbigny, 1913 | | | Onthophagus tenuistriatus d´Orbigny, 1905 | | | Onthophagus tersicollis Müller, 1947 | | | Onthophagus tersipennis d´Orbigny, 1902 | | | Onthophagus tersus d´Orbigny, 1913 | | | Onthophagus tesseratus d´Orbigny, 1908 | | | Onthophagus tessulatus Harold, 1871 | | | Onthophagus testaceoviolaceus Paulian, 1937 | | | Onthophagus thomsoni Bates, 1888 | | | Onthophagus tonsus d´Orbigny, 1902 | | | Onthophagus transcaspicus Koenig, 1888 | | | Onthophagus trapezicornis d´Orbigny, 1902 | | | Onthophagus traversii d´Orbigny, 1904 | | | Onthophagus triacanthus Castelnau, 1840 | | | Onthophagus tricariniger d´Orbigny, 1902 | | | Onthophagus trichopygus d´Orbigny, 1905 | | | Onthophagus tricorniger Boheman, 1860 | | | Onthophagus tridenticeps d´Orbigny, 1902 | | | Onthophagus trifidisetis d´Orbigny, 1905 | | | Onthophagus triimpressus d´Orbigny, 1905 | | | Onthophagus trinodosus Fahraeus, 1857 | | | Onthophagus trinominatus Goidanich, 1926 | | | Onthophagus tripartitus d´Orbigny, 1902 | | | Onthophagus tripolitanus Heyden, 1890 | | | Onthophagus triundulatus Balthasar, 1964 | | | Onthophagus tschadensis Balthasar, 1963 | | | Onthophagus tschoffeni d´Orbigny, 1904 | | | Onthophagus tshuapae Balthasar, 1964 | | | Onthophagus tubericollis (Raffray, 1877) | ⇒ Onthophagus bidens Olivier, 1789 | | Onthophagus tumidulus Gerstaecker, 1871 | | | Onthophagus turneri Boucomont, 1936 | | | Onthophagus tuzetae Walter & Cam..., 1977 | | | Onthophagus uelensis Frey, 1960 | | | Onthophagus ugandaensis Dierkens, 2022 | | | Onthophagus ugandicus Frey, 1975 | | | Onthophagus ugoi Moretto & Gé..., 2020 | | | Onthophagus ukerewensis Balthasar, 1937 | | | Onthophagus ulula Balthasar, 1966 | | | Onthophagus umbilicatus d´Orbigny, 1908 | | | Onthophagus umbratus d´Orbigny, 1902 | | | Onthophagus undaticeps d´Orbigny, 1902 | | | Onthophagus unicarina d´Orbigny, 1904 | | | Onthophagus ursinus d´Orbigny, 1902 | | | Onthophagus ursus Boucomont, 1926 | | | Onthophagus usambaricus Paulian, 1937 | | | Onthophagus valentineae Josso, 2019 | | | Onthophagus vaneyeni Frey, 1960 | | | Onthophagus variatus d´Orbigny, 1902 | | | Onthophagus variegatus (Fabricius, 1798) | ⇒ Onthophagus filicornis Harold, 1873 | | Onthophagus variegranosus d´Orbigny, 1905 | | | Onthophagus variolosus d´Orbigny, 1902 | | | Onthophagus varius d´Orbigny, 1913 | | | Onthophagus vassei d´Orbigny, 1908 | | | Onthophagus vatovai Müller, 1942 | | | Onthophagus velliger d´Orbigny, 1907 | | | Onthophagus ventrosus d´Orbigny, 1905 | | | Onthophagus venustulus Erichson, 1843 | | | Onthophagus venzoi Balthasar, 1941 | | | Onthophagus verae Josso, 2013 | | | Onthophagus verrucosus d´Orbigny, 1902 | | | Onthophagus versutus Péringuey, 1901 | | | Onthophagus verticalis (Fahraeus, 1857) | ⇒ Onthophagus praedatus Harold, 1862 | | Onthophagus vesanus Balthasar, 1967 | | | Onthophagus vestitus d´Orbigny, 1902 | | | Onthophagus vigens Péringuey, 1901 | | | Onthophagus vinctoides Frey, 1957 | | | Onthophagus vinctus Erichson, 1843 | | | Onthophagus violaceoviridis Paulian, 1937 | | | Onthophagus virens (d´Orbigny, 1913) | ⇒ Onthophagus alcyonides d´Orbigny, 1913 | | Onthophagus virescens Harold, 1867 | | | Onthophagus viridiaereus d´Orbigny, 1908 | | | Onthophagus viridicatus d´Orbigny, 1902 | | | Onthophagus viviensis d´Orbigny, 1905 | | | Onthophagus volsellatus (Boucomont, 1920) | ⇒ Onthophagus minax d´Orbigny, 1904 | | Onthophagus vuattouxi Cambefort, 1984 | | | Onthophagus vultuosus d´Orbigny, 1902 | | | Onthophagus vylderi d´Orbigny, 1913 | | | Onthophagus waterhousei Gillet, 1927 | | | Onthophagus waterloti d´Orbigny, 1908 | | | Onthophagus wensis Josso & Prév..., 2006 | | | Onthophagus willameorum Walter & Cam..., 1977 | | | Onthophagus wittei (Frey, 1958) | ⇒ Onthophagus witteianus Krell, 2009 | | Onthophagus witteianus Krell, 2009 | | | Onthophagus xanthochlorus Walter & Cam..., 1977 | | | Onthophagus xanthopterus d´Orbigny, 1908 | | | Onthophagus xanthopygus d´Orbigny, 1908 | | | Onthophagus zairensis Walter & Cam..., 1977 | | | Onthophagus zambundaticeps Josso, 2022 | | | Onthophagus zavattarii Müller, 1947 | | | Onthophagus zicsii Balthasar, 1967 | | | Onthophagus zumpti Frey, 1954 | | | Outenikwanus tomentosus Scholtz & Ho..., 1987 | | | Oxysternon festivum Linnaeus, 1758 | Staatskäfer | | Pachylomera femoralis Kirby, 1828 | | | Pachylomera opaca Lansberge, 1874 | | | Pachysoma aesculapius Olivier, 1789 | | | Pachysoma barbatus | | | Pachysoma bennigseni Felsche, 1907 | | | Pachysoma denticolle Péringuey, 1888 | | | Pachysoma denticollis Péringuey, 1888 | | | Pachysoma endroedyi Harrison, Sc..., 2003 | | | Pachysoma fitzsimonsi Ferreira, 1953 | | | Pachysoma gariepinum Ferreira, 1953 | | | Pachysoma gariepinus Ferreira, 1953 | | | Pachysoma glentoni Harrison, Sc..., 2003 | | | Pachysoma hippocrates MacLeay, 1821 | | | Pachysoma rodriguesi Ferreira, 1953 | | | Pachysoma rotundigena (Felsche, 1907) | ⇒ Scarabaeus rotundigenus Felsche, 1907 | | Pachysoma rotundigenus Felsche, 1907 | | | Pachysoma schinzi Fairmaire, 1888 | | | Pachysoma striata | | | Pachysoma striatum (Castelnau, 1840) | ⇒ Scarabaeus striatus Castelnau, 1840 | | Pachysoma striatus Castelnau, 1840 | | | Pachysoma valeflorae Ferreira, 1953 | | | Panelus pernitidus Balthasar, 1972 | | | Panelus rhodesiensis Paulian, 1975 | | | Paracanthon trichonotulus Balthasar, 1938 | | | Paraphytus africanus Boucomont, 1923 | | | Paraphytus aphodioides Boucomont, 1928 | | | Paraphytus bechynei Balthasar, 1958 | | | Paraphytus ritsemae Harold, 1880 | | | Paraphytus sancyi Paulian, 1949 | | | Paraphytus tai Cambefort, 1984 | | | Paronitis excavaticollis Balthasar, 1942 | | | Parvuhowdenius harrisoni Deschodt & C..., 2008 | | | Peckolus alpinus Howden & Sch..., 1988 | | | Peckolus parvus Scholtz & Ho..., 1987 | | | Peckolus poenskopius Deschodt & C..., 2008 | | | Pedaria acuta Josso & Prév..., 2015 | | | Pedaria aethiopica Josso & Prév..., 2003 | | | Pedaria alternans Waterhouse, 1890 | | | Pedaria angolana Boucomont, 1922 | | | Pedaria antoinei Josso & Prév..., 2009 | | | Pedaria armata Raffray, 1877 | | | Pedaria aspera Péringuey, 1901 | | | Pedaria barrei Josso & Prév..., 2006 | | | Pedaria biseria Josso & Prév..., 2003 | | | Pedaria brancoi Josso & Prév..., 2003 | | | Pedaria cambeforti Josso & Prév..., 2003 | | | Pedaria catherinae Josso & Prév..., 2009 | | | Pedaria cicatricosus (Péringuey, 1901) | ⇒ Sarophorus cicatricosus Péringuey, 1901 | | Pedaria conformis Péringuey, 1901 | | | Pedaria coprinarum Cambefort, 1982 | | | Pedaria criberrima Waterhouse, 1890 | | | Pedaria cylindrica Fahraeus, 1857 | | | Pedaria decorsei (Boucomont, 1922) | ⇒ Pedaria taylori Waterhouse, 1890 | | Pedaria dellacasai Josso & Prév..., 2009 | | | Pedaria demeyeri Josso & Prév..., 2003 | | | Pedaria dentata Raffray, 1877 | | | Pedaria dudleyi Josso & Prév..., 2003 | | | Pedaria durandi Paulian, Cam..., 1982 | | | Pedaria estellae Josso & Prév..., 2006 | | | Pedaria fernandezi Cambefort, 1982 | | | Pedaria genieri Josso & Prév..., 2003 | | | Pedaria genierorum Josso & Prév..., 2015 | | | Pedaria gigantea Josso & Prév..., 2003 | | | Pedaria granulosa Ferreira, 1962 | | | Pedaria hanae Balthasar, 1963 | | | Pedaria humana Cambefort, 1982 | | | Pedaria insularis Kolbe, 1914 | | | Pedaria intermedia Boucomont, 1922 | | | Pedaria jacksoni Waterhouse, 1890 | | | Pedaria jossoi Génier, 2017 | | | Pedaria juhellegrandi Josso & Prév..., 2003 | | | Pedaria manni Josso & Prév..., 2015 | | | Pedaria minetti Josso & Prév..., 2009 | | | Pedaria minima Josso & Prév..., 2003 | | | Pedaria montreuili Josso & Prév..., 2015 | | | Pedaria morettoi Josso & Prév..., 2003 | | | Pedaria murphyi Josso & Prév..., 2003 | | | Pedaria murphyi murphyi | | | Pedaria murphyi septentrionalis Josso & Prév..., 2009 | | | Pedaria nigra Laporte, 1832 | | | Pedaria oblonga Josso & Prév..., 2003 | | | Pedaria ouangoensis Cambefort, 1982 | | | Pedaria ovata Boucomont, 1922 | | | Pedaria picea Fahraeus, 1857 | | | Pedaria puncticollis Waterhouse, 1890 | | | Pedaria quirimbasensis Josso & Prév..., 2015 | | | Pedaria raffrayi Josso & Prév..., 2006 | | | Pedaria reichenbachi Josso & Prév..., 2006 | | | Pedaria renwarti Cambefort, 1982 | | | Pedaria reticulata Josso & Prév..., 2015 | | | Pedaria rohani Boucomont, 1922 | | | Pedaria rosemariae Josso & Prév..., 2009 | | | Pedaria segregis Péringuey, 1901 | | | Pedaria segrestis | | | Pedaria smrzi Josso & Prév..., 2003 | | | Pedaria sobrina (Péringuey, 1901) | ⇒ Pedaria picea Fahraeus, 1857 | | Pedaria spinipennis Josso & Prév..., 2003 | | | Pedaria spinithorax Paulian, Cam..., 1982 | | | Pedaria spinosa Boucomont, 1922 | | | Pedaria sudrei Josso & Prév..., 2003 | | | Pedaria tanganyika Josso & Prév..., 2003 | | | Pedaria taylori Waterhouse, 1890 | | | Pedaria tenebrosa Balthasar, 1937 | | | Pedaria tibialis Josso & Prév..., 2006 | | | Pedaria tuberculigera Waterhouse, 1890 | | | Pedaria walteri Josso & Prév..., 2003 | | | Pedaria werneri Josso & Prév..., 2003 | | | Pedaria wittei Janssens, 1939 | | | Phalops ardea Klug, 1855 | | | Phalops aurifrons Fairmaire, 1884 | | | Phalops barbicornis Lansberge, 1883 | | | Phalops batesi Koenike, 1892 | | | Phalops batesi batesi | | | Phalops batesi pseudobatesi Moretto, 2007 | | | Phalops beccarii Harold, 1875 | | | Phalops boschas Klug, 1855 | | | Phalops bubalus Harold, 1867 | | | Phalops chalybeus Klug, 1845 | | | Phalops djuricus Kuntzen, 1913 | | | Phalops dregei Harold, 1867 | | | Phalops faccai Müller, 1941 | | | Phalops fimbriatus Klug, 1835 | | | Phalops flavocinctus Klug, 1855 | | | Phalops gallanus d´Orbigny, 1904 | | | Phalops glauningi Kuntzen, 1913 | | | Phalops guttulatus Fairmaire, 1887 | | | Phalops inermis Lansberge, 1883 | | | Phalops iphis Olivier, 1789 | | | Phalops laminifrons Fairmaire, 1882 | | | Phalops lutatus d´Orbigny, 1902 | | | Phalops pauliani Barbero & Pa..., 2003 | | | Phalops plancus Erichson, 1843 | | | Phalops prasinus (Erichson, 1843) | ⇒ Phalops beccarii Harold, 1875 | | Phalops princeps d´Orbigny, 1908 | | | Phalops pyroides d´Orbigny, 1908 | | | Phalops rufosignatus (Lansberge, 1885) | ⇒ Phalops dregei Harold, 1867 | | Phalops sinuaticeps d´Orbigny, 1902 | | | Phalops smaragdinus Harold, 1875 | | | Phalops sulcatus Lansberge, 1883 | | | Phalops tricuspis d´Orbigny, 1908 | | | Phalops tuberosus d´Orbigny, 1905 | | | Phalops vanellus Lansberge, 1883 | | | Phalops wittei Harold, 1867 | | | Phalops zuninoi Barbero & Pa..., 2003 | | | Phanaeus (Notiophanaeus) pyrois Bates, 1887 | | | Phanaeus (Phanaeus) adonis Harold, 1863 | | | Phanaeus (Phanaeus) vindex MacLeay, 1819 | | | Phanaeus beltianus Bates, 1887 | | | Phanaeus daphnis Harold, 1863 | | | Phanaeus demon Castelnau, 1840 | | | Phanaeus eximius Bates, 1887 | | | Phanaeus tridens Castelnau, 1840 | | | Phanaeus wagneri Harold, 1863 | | | Pinacopodius meruensis Krikken, 1982 | | | Pinacopodius praestolatus Branco & Josso, 2007 | | | Pinacotarsus dohrni Harold, 1875 | | | Pinacotarsus proximus Ferreira, 1969 | | | Platyonitis bicuariensis Ferreira, 1976 | | | Platyonitis oberthueri Janssens, 1942 | | | Platyonitis smeenkorum Krikken, 1974 | | | Pleuronitis boucomonti Janssens, 1937 | | | Pleuronitis favareli Garreta, 1914 | | | Pleuronitis fulgidus Castelnau, 1840 | | | Pleuronitis minettii Josso, 2011 | | | Proagoderus aciculatus Fahraeus, 1857 | | | Proagoderus albicapillus d´Orbigny, 1902 | | | Proagoderus amblychromatus Marcus, 1917 | | | Proagoderus armicollis d´Orbigny, 1904 | | | Proagoderus atriclaviger d´Orbigny, 1905 | | | Proagoderus atrosetosus d´Orbigny, 1902 | | | Proagoderus auratus Fabricius, 1801 | | | Proagoderus aureiceps d´Orbigny, 1902 | | | Proagoderus biarmatus d´Orbigny, 1908 | | | Proagoderus bicallosus Klug, 1855 | | | Proagoderus boranus Gestro, 1895 | | | Proagoderus bottegi Gestro, 1895 | | | Proagoderus brucei (d´Orbigny, 1902) | ⇒ Onthophagus chalcostolus d´Orbigny, 1902 | | Proagoderus brucei brucei | | | Proagoderus brucei cyanostolus d´Orbigny, 1902 | | | Proagoderus brucei njombeensis Moretto, 2004 | | | Proagoderus cambeforti Palestrini, 1992 | | | Proagoderus camiadei Josso, 2014 | | | Proagoderus cavidorsis d´Orbigny, 1902 | | | Proagoderus ceratotherium Janssens, 1953 | | | Proagoderus chalcostolus d´Orbigny, 1902 | | | Proagoderus chrysopes Bates, 1888 | | | Proagoderus colmanti d´Orbigny, 1908 | | | Proagoderus cometes Bates, 1888 | | | Proagoderus confossus d´Orbigny, 1915 | | | Proagoderus cuspidatus d´Orbigny, 1913 | | | Proagoderus cyanopterus d´Orbigny, 1905 | | | Proagoderus dargei Moretto, 2013 | | | Proagoderus dives Harold, 1877 | | | Proagoderus dudleyi Cambefort, 1980 | | | Proagoderus edithae Josso, 2014 | | | Proagoderus elgoni d´Orbigny, 1915 | | | Proagoderus euchlorus d´Orbigny, 1902 | | | Proagoderus extensus Harold, 1878 | | | Proagoderus fastosus d´Orbigny, 1913 | | | Proagoderus flexicollis d´Orbigny, 1904 | | | Proagoderus fossidorsis d´Orbigny, 1905 | | | Proagoderus freyi Moretto, 2004 | | | Proagoderus furcifer Boheman, 1860 | | | Proagoderus furciramus d´Orbigny, 1902 | | | Proagoderus gangloffi Josso & Pré..., 2001 | | | Proagoderus gemmatus Péringuey, 1901 | | | Proagoderus gerstaeckeri Harold, 1867 | | | Proagoderus gibbiramus d´Orbigny, 1902 | | | Proagoderus harpax Fabricius, 1801 | | | Proagoderus hemicyaneus | | | Proagoderus hemicyanus d´Orbigny, 1902 | | | Proagoderus ignitus d´Orbigny, 1905 | | | Proagoderus incostatus d´Orbigny, 1902 | | | Proagoderus kachowskii Olsoufief, 1900 | | | Proagoderus kafueensis Josso, 2014 | | | Proagoderus katualensis Quedenfeldt, 1888 | | | Proagoderus kilimanus Kolbe, 1891 | | | Proagoderus kolbei Marcus, 1921 | | | Proagoderus kwaleensis Moretto & Ni..., 2002 | | | Proagoderus lallieri d´Orbigny, 1902 | | | Proagoderus lallierianus Janssens, 1953 | | | Proagoderus lanista Castelnau, 1840 | | | Proagoderus laticollis Klug, 1835 | | | Proagoderus longefossus d´Orbigny, 1905 | | | Proagoderus loricatus Klug, 1855 | | | Proagoderus louisi Josso, 2014 | | | Proagoderus lucasi d´Orbigny, 1902 | | | Proagoderus lujendae Bates, 1888 | | | Proagoderus marginidens d´Orbigny, 1904 | | | Proagoderus maringouensis Palestrini, 1992 | | | Proagoderus metallites (d´Orbigny, 1905) | ⇒ Proagoderus nigricornis Fairmaire, 1887 | | Proagoderus minettii Moretto & Ni..., 2002 | | | Proagoderus monochromus d´Orbigny, 1913 | | | Proagoderus moorei Moretto, 2013 | | | Proagoderus multicornis d´Orbigny, 1908 | | | Proagoderus nasidens d´Orbigny, 1905 | | | Proagoderus negus Raffray, 1882 | | | Proagoderus ngottoensis Moretto & Ni..., 2002 | | | Proagoderus nicolasi Moretto, 2004 | | | Proagoderus nigricornis Fairmaire, 1887 | | | Proagoderus nuba d´Orbigny, 1902 | | | Proagoderus opulentus d´Orbigny, 1902 | | | Proagoderus overlaeti Janssens, 1953 | | | Proagoderus palestriniae Moretto & Ni..., 2004 | | | Proagoderus panoplus Bates, 1888 | | | Proagoderus paolae Palestrini, 1992 | | | Proagoderus patinatus Marcus, 1917 | | | Proagoderus pegesimallus Marcus, 1917 | | | Proagoderus plato Bates, 1888 | | | Proagoderus porrectus Reiche, 1847 | | | Proagoderus praecavatus d´Orbigny, 1913 | | | Proagoderus praedentatus d´Orbigny, 1904 | | | Proagoderus praefossus d´Orbigny, 1913 | | | Proagoderus prostans Reiche, 1847 | | | Proagoderus pseudoflexicollis Moretto, 2004 | | | Proagoderus pyramidalis Klug, 1855 | | | Proagoderus quadriarmatus Fairmaire, 1892 | | | Proagoderus quadricristatus d´Orbigny, 1908 | | | Proagoderus quadrispinosus d´Orbigny, 1908 | | | Proagoderus quadrituber d´Orbigny, 1908 | | | Proagoderus ramosicornis d´Orbigny, 1902 | | | Proagoderus rangifer Klug, 1855 | | | Proagoderus rarus Guérin-Mén..., 1829 | | | Proagoderus rasidorsis d´Orbigny, 1905 | | | Proagoderus rectefurcatus Fairmaire, 1891 | | | Proagoderus revoili Lansberge, 1882 | | | Proagoderus ritsemai Lansberge, 1883 | | | Proagoderus sapphirinus (Péringuey, 1901) | ⇒ Onthophagus aureiceps d´Orbigny, 1902 | | Proagoderus semiiris Thomson, 1858 | | | Proagoderus sexcornutus d´Orbigny, 1902 | | | Proagoderus somalicus d´Orbigny, 1902 | | | Proagoderus speculatus d´Orbigny, 1908 | | | Proagoderus speculicollis Quedenfeldt, 1884 | | | Proagoderus subextensus Kolbe, 1895 | | | Proagoderus superbus d´Orbigny, 1904 | | | Proagoderus taymansi d´Orbigny, 1908 | | | Proagoderus tersidorsis d´Orbigny, 1902 | | | Proagoderus trianguliceps d´Orbigny, 1905 | | | Proagoderus triarmatus d´Orbigny, 1904 | | | Proagoderus tricornifrons d´Orbigny, 1908 | | | Proagoderus uluguru Frey, 1975 | | | Proagoderus unidens Quedenfeldt, 1884 | | | Proagoderus versus d´Orbigny, 1913 | | | Proagoderus werneri Moretto & Ni..., 2002 | | | Proagoderus worsissa Roth, 1851 | | | Proagoderus yvescambeforti Moretto, 2004 | | | Pseudocanthon perplexus LeConte, 1847 | | | Pseudochironitis bennigseni Felsche, 1907 | | | Pseudochironitis benningseni Felsche, 1907 | | | Pseudochironitis stuhlmanni Kolbe, 1897 | | | Pseudocopris lunaticeps Gillet, 1910 | | | Pseudocopris tubericollis Felsche, 1910 | | | Pseudosaproecius baraudi Branco, 1995 | | | Pseudosaproecius bateke Cambefort & ..., 1977 | | | Pseudosaproecius binasus d´Orbigny, 1907 | | | Pseudosaproecius cambeforti Branco, 1995 | | | Pseudosaproecius comatus d´Orbigny, 1902 | | | Pseudosaproecius cylindroides d´Orbigny, 1908 | | | Pseudosaproecius dicerus d´Orbigny, 1915 | | | Pseudosaproecius falcatus Branco, 1995 | | | Pseudosaproecius mirepunctatus d´Orbigny, 1905 | | | Pseudosaproecius ocellatus Frey, 1958 | | | Pseudosaproecius portentosus Felsche, 1907 | | | Pseudosaproecius validicornis Quedenfeldt, 1884 | | | Pycnopanelus krikkeni Cambefort, 1978 | | | Pycnopanelus parvicollis d´Orbigny, 1913 | | | Sarophorus angolensis Frolov, 2004 | | | Sarophorus bidentatus Frolov & Sch..., 2003 | | | Sarophorus carinatus Frolov & Sch..., 2003 | | | Sarophorus cicatricosus Péringuey, 1901 | | | Sarophorus costatus Fahraeus, 1857 | | | Sarophorus diabolus Roets, 2017 | | | Sarophorus frolovi Roets, 2017 | | | Sarophorus latus Frolov & Sch..., 2003 | | | Sarophorus nitidus Frolov & Sch..., 2003 | | | Sarophorus punctatus Frolov & Sch..., 2003 | | | Sarophorus striatus Frolov & Sch..., 2003 | | | Sarophorus tuberculatus Castelnau, 1840 | | | Scaptocnemis segregis Péringuey, 1901 | | | Scarabaeus (Ateuchetus) cicatricosus (Lucas, 1846) | ⇒ Scarabaeus palemo Olivier, 1789 | | Scarabaeus (Ateuchetus) laticollis Linnaeus, 1767 | | | Scarabaeus (Ateuchetus) puncticollis (Latreille, 1819) | ⇒ Geotrupes spiniger Marsham, 1802 | | Scarabaeus (Ateuchetus) semipunctatus Fabricius, 1792 | | | Scarabaeus (Ateuchetus) variolosus (Fabricius, 1787) | ⇒ Ateuchetus semipunctatus Fabricius, 1792 | | Scarabaeus (Scarabaeus) bannuensis Janssens, 1940 | | | Scarabaeus (Scarabaeus) pius Illiger, 1803 | | | Scarabaeus (Scarabaeus) sacer (Linnaeus, 1758) | ⇒ Scarabaeus pius Illiger, 1803 | | Scarabaeus (Scarabaeus) winkleri Stolfa, 1938 | | | Scarabaeus aegaeon Drury, 1773 | | | Scarabaeus aegyptiacus Stolfa, 1938 | | | Scarabaeus aegyptiorum Latreille, 1827 | | | Scarabaeus aegyptiorum purpurascens Gerstaecker, 1871 | | | Scarabaeus aesculapius Olivier, 1789 | | | Scarabaeus alienus Péringuey, 1901 | | | Scarabaeus ambiguus (Boheman, 1857) | ⇒ Gnorimus variabilis Linnaeus, 1758 | | Scarabaeus anderseni Waterhouse, 1890 | | | Scarabaeus andreaei zur Strassen, 1963 | | | Scarabaeus auratus (Schrank, 1781) | ⇒ Gnorimus nobilis Linnaeus, 1758 | | Scarabaeus basuto zur Strassen, 1962 | | | Scarabaeus bennigseni Felsche, 1907 | | | Scarabaeus bohemani Harold, 1868 | | | Scarabaeus bornemisszai zur Strassen, 1980 | | | Scarabaeus bornemizzai zur Strassen, 1980 | | | Scarabaeus brahminus Castelnau, 1840 | | | Scarabaeus caffer Boheman, 1857 | | | Scarabaeus canaliculatus Fairmaire, 1888 | | | Scarabaeus cancer Arrow, 1919 | | | Scarabaeus carniphilus Davis & Desc..., 2015 | | | Scarabaeus catenatus Gerstaecker, 1871 | | | Scarabaeus clanceyi Ferreira, 1954 | | | Scarabaeus cognatus Péringuey, 1901 | | | Scarabaeus confusus Müller, 1942 | | | Scarabaeus convexus Hausmann, 1807 | | | Scarabaeus costatus Wiedemann, 1823 | | | Scarabaeus damarensis Janssens, 1940 | | | Scarabaeus deludens zur Strassen, 1961 | | | Scarabaeus denticollis Péringuey, 1888 | | | Scarabaeus dioscoridis Petrovitz, 1962 | | | Scarabaeus ebenus Klug, 1855 | | | Scarabaeus endroedyi Harrison & S..., 2003 | | | Scarabaeus ermienae Deschodt & D..., 2015 | | | Scarabaeus festivus Harold, 1868 | | | Scarabaeus fitzsimonsi Ferreira, 1953 | | | Scarabaeus flavicornis Boheman, 1860 | | | Scarabaeus fraterculus Kolbe, 1895 | | | Scarabaeus fritschi Harold, 1868 | | | Scarabaeus funebris Boheman, 1857 | | | Scarabaeus galenus Westwood, 1847 | | | Scarabaeus gangeticus Castelnau, 1840 | | | Scarabaeus gariepinus Ferreira, 1953 | | | Scarabaeus geminogalenus Davis & Desc..., 2017 | | | Scarabaeus gilleti Janssens, 1940 | | | Scarabaeus glentoni Harrison & S..., 2003 | | | Scarabaeus goryi Castelnau, 1840 | | | Scarabaeus gracai Ferreira, 1952 | | | Scarabaeus heqvisti zur Strassen, 1962 | | | Scarabaeus hippocrates MacLeay, 1821 | | | Scarabaeus hottentorum Péringuey, 1901 | | | Scarabaeus indus (Linnaeus, 1758) | ⇒ Euphoria inda Linnaeus, 1758 | | Scarabaeus inguisitus | | | Scarabaeus inoportunus Ferreira, 1953 | | | Scarabaeus inquisitus Péringuey, 1908 | | | Scarabaeus interstitialis Boheman, 1857 | | | Scarabaeus intricatus Fabricius, 1801 | | | Scarabaeus isidis Castelnau, 1840 | | | Scarabaeus jalof Castelnau, 1840 | | | Scarabaeus janssensi Balthasar, 1940 | | | Scarabaeus karae Davis & Desc..., 2016 | | | Scarabaeus karlwerneri Nicolas & Mo..., 2002 | | | Scarabaeus karrooensis zur Strassen, 1961 | | | Scarabaeus knobeli Ferreira, 1958 | | | Scarabaeus kochi Ferreira, 1952 | | | Scarabaeus kwiluensis Janssens, 1940 | | | Scarabaeus laevifrons Fairmaire, 1884 | | | Scarabaeus laevigatus (Kolbe, 1887) | ⇒ Scarabaeus goryi Castelnau, 1840 | | Scarabaeus lamarcki MacLeay, 1821 | | | Scarabaeus licitus Ferreira, 1954 | | | Scarabaeus lucidulus Boheman, 1860 | | | Scarabaeus megaparvulus Davis & Desc..., 2015 | | | Scarabaeus montreuili meridionalis Moretto, 2023 | | | Scarabaeus montreuili montreuili | | | Scarabaeus natalensis zur Strassen, 1962 | | | Scarabaeus nepos (Fairmaire, 1884) | ⇒ Scarabaeus goryi Castelnau, 1840 | | Scarabaeus niemandi Deschodt & D..., 2015 | | | Scarabaeus nigroaeneus Boheman, 1857 | | | Scarabaeus nitidicollis Lansberge, 1882 | | | Scarabaeus obsoletepunctatus Balthasar, 1940 | | | Scarabaeus opacipennis Fairmaire, 1884 | | | Scarabaeus osiridis Moretto, 2023 | | | Scarabaeus pabulator Péringuey, 1908 | | | Scarabaeus palemo Olivier, 1789 | | | Scarabaeus parvulus Boheman, 1860 | | | Scarabaeus piliventris zur Strassen, 1962 | | | Scarabaeus planifrons Fairmaire, 1884 | | | Scarabaeus planipennis Davis & Desc..., 2015 | | | Scarabaeus plansibilis | | | Scarabaeus platynotus Bates, 1888 | | | Scarabaeus plausibilis Péringuey, 1892 | | | Scarabaeus poggei Waterhouse, 1890 | | | Scarabaeus proboscideus Guérin-Ménev..., 1844 | | | Scarabaeus prodigiosus Erichson, 1843 | | | Scarabaeus profanus Fabricius, 1792 | | | Scarabaeus proximus (Péringuey, 1901) | ⇒ Drepanopodus proximus Péringuey, 1901 | | Scarabaeus pubiventris (Lansberge, 1874) | ⇒ Kheper prodigiosus Erichson, 1843 | | Scarabaeus radama Fairmaire, 1895 | | | Scarabaeus ritchiei MacLeay, 1821 | | | Scarabaeus rixosus Péringuey, 1901 | | | Scarabaeus rodriguesi Ferreira, 1953 | | | Scarabaeus rotundigenus Felsche, 1907 | | | Scarabaeus rubripennis Boheman, 1860 | | | Scarabaeus ruficornis | | | Scarabaeus rugosus (Hausmann, 1807) | ⇒ Onthophagus taurus Schreber, 1759 | | Scarabaeus rusticus Boheman, 1857 | | | Scarabaeus satyrus (Boheman, 1860) | ⇒ Xyloryctes jamaicensis Drury, 1773 | | Scarabaeus savignyi MacLeay, 1821 | | | Scarabaeus schinzi Fairmaire, 1888 | | | Scarabaeus scholtzi Mostert & Holm, 1982 | | | Scarabaeus schulzeae Ferreira, 1969 | | | Scarabaeus sevoistra Alluaud, 1902 | | | Scarabaeus soutpansbergensis Deschodt & D..., 2015 | | | Scarabaeus spretus zur Strassen, 1962 | | | Scarabaeus striatus Castelnau, 1840 | | | Scarabaeus sulcipennis Quedenfeldt, 1888 | | | Scarabaeus suri Hausmann, 1807 | | | Scarabaeus tonckeri Boucomont, 1923 | | | Scarabaeus transcaspicus Stolfa, 1938 | | | Scarabaeus vansoni Ferreira, 1958 | | | Scarabaeus venerabilis Harold, 1871 | | | Scarabaeus vethi Lansberge, 1886 | | | Scarabaeus viator Péringuey, 1901 | | | Scarabaeus vicinus (Janssens, 1940) | ⇒ Lucanus cervus Linnaeus, 1758 | | Scarabaeus viettei Paulian, 1953 | | | Scarabaeus westwoodi (Harold, 1869) | ⇒ Scarabaeus galenus Westwood, 1847 | | Scarabaeus xavieri Ferreira, 1968 | | | Scarabaeus zambesianus Péringuey, 1901 | | | Scarabaeus zambezianus Péringuey, 1901 | | | Scatimus erinnyos Kohlmann & S..., 1996 | | | Scatimus ovatus Harold, 1862 | | | Sceliages adamastor Lepeletier &..., 1828 | | | Sceliages augias Gillet, 1908 | | | Sceliages brittoni zur Strassen, 1965 | | | Sceliages curvipes (Gillet, 1911) | ⇒ Sceliages adamastor Lepeletier &..., 1828 | | Sceliages difficilis zur Strassen, 1965 | | | Sceliages gagates Shipp, 1895 | | | Sceliages granulatus Forgie & Gre..., 2002 | | | Sceliages hippias Westwood, 1844 | | | Silvaphilus joselmae Daniel, Strü..., 2022 | | | Silvaphilus oubosiensis Roets & Ober..., 2010 | | | Sisyphus (Sisyphus) struwei Schafer & Fi..., 1992 | | | Sisyphus struwei kivuensis Schafer & Fi..., 2001 | | | Sisyphus struwei struwei | | | Sisyphus africanus africanus | | | Sisyphus africanus occidentalis Montreuil, 2017 | | | Sisyphus africanus orientalis Montreuil, 2017 | | | Sisyphus alveatus Boucomont, 1935 | | | Sisyphus angulicollis (Felsche, 1909) | ⇒ Neosisyphus angulicollis Felsche, 1909 | | Sisyphus arboreus Walter, 1982 | | | Sisyphus armatus (Gory, 1833) | ⇒ Neosisyphus confrater Kolbe, 1914 | | Sisyphus astriatus Schafer & Fi..., 2001 | | | Sisyphus atratus (Klug, 1855) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Sisyphus auricomus Daniel & Davis, 2018 | | | Sisyphus australis Daniel & Davis, 2018 | | | Sisyphus baoule Cambefort, 1984 | | | Sisyphus barbarossa (Wiedemann, 1823) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Sisyphus basilewskyi (Haaf, 1956) | ⇒ Neosisyphus basilewskyi Haaf, 1956 | | Sisyphus bayanga Montreuil, 2016 | | | Sisyphus biarmatus Felsche, 1909 | | | Sisyphus bicuariensis Daniel & Davis, 2018 | | | Sisyphus bornemisszanus Endrödi, 1983 | | | Sisyphus bouyeri Montreuil, 2015 | | | Sisyphus caffer Boheman, 1857 | | | Sisyphus caffer goryi Harold, 1859 | | | Sisyphus calcaratus (Boucomont, 1937) | ⇒ Neosisyphus rugosus Gory, 1833 | | Sisyphus cambeforti Montreuil, 2015 | | | Sisyphus costatus Thunberg, 1818 | | | Sisyphus crispatus Gory, 1833 | | | Sisyphus crispatus crispatus | | | Sisyphus desaegeri Haaf, 1959 | | | Sisyphus dierkensi Montreuil, 2015 | | | Sisyphus drumonti Montreuil, 2015 | | | Sisyphus eburneus Cambefort, 1984 | | | Sisyphus ethiopicus Montreuil, 2017 | | | Sisyphus fasciculatus Boheman, 1857 | | | Sisyphus felschei Montreuil, 2015 | | | Sisyphus fortuitus Péringuey, 1900 | | | Sisyphus gazanus Arrow, 1909 | | | Sisyphus genierorum Montreuil, 2015 | | | Sisyphus gladiator (Arrow, 1927) | ⇒ Neosisyphus gladiator Arrow, 1927 | | Sisyphus goryi (Harold, 1859) | ⇒ Sisyphus caffer goryi Harold, 1859 | | Sisyphus granulifer Montreuil, 2014 | | | Sisyphus ibaya Montreuil, 2017 | | | Sisyphus impressipennis Lansberge, 1886 | | | Sisyphus inconspicuus Daniel & Davis, 2018 | | | Sisyphus infuscatus (Klug, 1855) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Sisyphus janssensi Montreuil, 2015 | | | Sisyphus jossoi Montreuil, 2015 | | | Sisyphus juheli Montreuil, 2015 | | | Sisyphus kibale Montreuil, 2016 | | | Sisyphus kuehni Haaf, 1955 | | | Sisyphus latus (Boucomont, 1928) | ⇒ Sisyphus costatus Thunberg, 1818 | | Sisyphus lobi Cambefort, 1984 | | | Sisyphus macrorubrus (Paschalidis, 1974) | ⇒ Neosisyphus macrorubrus Paschalidis, 1974 | | Sisyphus manni Montreuil, 2015 | | | Sisyphus minettii Montreuil, 2014 | | | Sisyphus mirabilis Arrow, 1927 | | | Sisyphus mkomazi Montreuil, 2017 | | | Sisyphus moorei Montreuil, 2015 | | | Sisyphus morenoi Montreuil, 2015 | | | Sisyphus morettoi Montreuil, 2015 | | | Sisyphus muricatus Olivier, 1789 | | | Sisyphus nanniscus Péringuey, 1901 | | | Sisyphus natalensis Balthasar, 1968 | | | Sisyphus negus Montreuil, 2015 | | | Sisyphus neobornemisszanus Daniel & Davis, 2016 | | | Sisyphus nodifer Gerstaecker, 1871 | | | Sisyphus ocellatus Reiche, 1847 | | | Sisyphus ocellatus nanniscus Péringuey, 1901 | | | Sisyphus ongo Montreuil, 2017 | | | Sisyphus oralensis Daniel & Davis, 2016 | | | Sisyphus paschalidisae Cambefort, 1984 | | | Sisyphus penicillatus Harold, 1880 | | | Sisyphus perissinottoi Montreuil, 2015 | | | Sisyphus quadricollis (Péringuey, 1901) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Sisyphus rubripes (Boheman, 1857) | ⇒ Neosisyphus confrater Kolbe, 1914 | | Sisyphus rubrus (Paschalidis, 1974) | ⇒ Neosisyphus rubrus Paschalidis, 1974 | | Sisyphus rugosus (Roth, 1851) | ⇒ Sisyphus costatus Thunberg, 1818 | | Sisyphus schaefferi Linnaeus, 1758 | Lanbeiniger Pillendreher | | Sisyphus schaefferi schaefferi Linnaeus, 1758 | | | Sisyphus seminulum (Gerstaecker, 1871) | ⇒ Sisyphus costatus Thunberg, 1818 | | Sisyphus setiger (Roth, 1851) | ⇒ Neosisyphus confrater Kolbe, 1914 | | Sisyphus sordidus (Boheman, 1857) | ⇒ Sisyphus caffer Boheman, 1857 | | Sisyphus spinipes (Thunberg, 1818) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Sisyphus splendidus Montreuil, 2015 | | | Sisyphus swazi Daniel & Davis, 2018 | | | Sisyphus tai Cambefort, 1984 | | | Sisyphus tembyi Daniel & Davis, 2018 | | | Sisyphus tibialis Raffray, 1877 | | | Sisyphus transvaalensis (Péringuey, 1908) | ⇒ Sisyphus impressipennis Lansberge, 1886 | | Sisyphus trichodichromicus Montreuil, 2015 | | | Sisyphus trochantericus (Fairmaire, 1891) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Sisyphus umbraphilus Daniel & Davis, 2016 | | | Sisyphus usambaricus Montreuil, 2015 | | | Sisyphus walteri Montreuil, 2016 | | | Sisyphus werneri Montreuil, 2017 | | | Sisyphus youngai Endrödi, 1973 | | | Stiptocnemis gilleti Boucomont, 1923 | | | Stiptocnemis hiekei Branco, 1991 | | | Stiptopodius brancoi Strümpher & ..., 2021 | | | Stiptopodius doriae Harold, 1871 | | | Stiptopodius gaillardi Boucomont, 1923 | | | Stiptopodius glaber Branco, 1992 | | | Stiptopodius glabricollis Müller, 1942 | | | Stiptopodius granulosus Branco, 2009 | | | Stiptopodius krausei Branco, 1991 | | | Stiptopodius longipedis Branco, 1991 | | | Stiptopodius muellerae Daniel & Des..., 2020 | | | Stiptopodius nitidus Boucomont, 1923 | | | Stiptopodius nodieri Boucomont, 1923 | | | Stiptopodius patrizii Boucomont, 1923 | | | Stiptopodius peringueyi Daniel & Des..., 2020 | | | Stiptopodius savuti Daniel & Des..., 2020 | | | Stiptopodius singularis Péringuey, 1901 | | | Stiptotarsus latetarsatus Boucomont, 1923 | | | Stiptotarsus mocquerysi Boucomont, 1923 | | | Stiptotarsus tersus Balthasar, 1963 | | | Strandius obliquus (Cambefort, 1980) | ⇒ Onthophagus obliquus Olivier, 1789 | | Sukelus jessopi Branco, 1991 | | | Sulcodrepanus sulcicollis Castelnau, 1840 | | | Sulcophanaeus noctis Bates, 1887 | | | Sulcophanaeus velutinus Murray, 1856 | | | Tanzanolus leleupi Balthasar, 1960 | | | Tesserodon pilicrepus Matthews, 1974 | | | Tibiodrepanus sulcicollis Castelnau, 1840 | | | Tibiodrepanus tagliaferrii Barbero, Pal..., 2011 | | | Tiniocellus dolosus Branco, 2010 | | | Tiniocellus eurypygus Branco, 2010 | | | Tiniocellus eurypygus eurypygus | | | Tiniocellus eurypygus transdrakensbergensis Branco, 2010 | | | Tiniocellus praetermissus Branco, 2010 | | | Tiniocellus setifer Kraatz, 1895 | | | Tiniocellus spinipes Roth, 1851 | | | Tomogonus cambeforti Nicolas, 2002 | | | Tomogonus chauvetorum Walter, 2012 | | | Tomogonus crassoides Boucomont, 1923 | | | Tomogonus crassus d´Orbigny, 1902 | | | Tomogonus implicatus Demange, 1978 | | | Tomogonus intortus Demange, 1971 | | | Tomogonus involutus Demange & Ma..., 1975 | | | Tomogonus lamottei Demange & Ma..., 1975 | | | Tomogonus samburu Walter, 2012 | | | Tomogonus schubarti Demange & Ma..., 1975 | | | Tomogonus subgrundus Demange, 1971 | | | Tragiscus dimidiatus Klug, 1855 | | | Trichillidium pilosum Robinson, 1948 | | | Tropidonitis paradoxus Boheman, 1857 | | | Unidentis vwaza Josso & Prév..., 2008 | | | Uroxys boneti Pereira & Ha..., 1961 | | | Uroxys deavilai Delgado & Ko..., 2007 | | | Uroxys dybasi Howden & Young, 1981 | | | Uroxys micros Bates, 1887 | | | Uroxys pauliani Balthasar, 1940 | | | Versicorpus daures Deschodt, 2019 | | | Versicorpus erongoense Deschodt, Da..., 2011 | | | Versicorpus erongoensis Deschodt, Da..., 2011 | | | Versicorpus streyi Frolov, 2005 | | | Walterantus rufoclypeatus Frey, 1960 | | | Xinidium davisi Cambefort, 1985 | | | Xinidium dentilabris Harold, 1869 | | | Xinidium dewitzi Harold, 1878 | | | Xinidium howdeni Cambefort, 1993 | |
Unterknoten von Scarabaeinae mit Vorkommen in Afrika | Tribus | Ateuchini
Montreuil, 1998
| 9
118 | Gattungen
Arten |
Taxom
| | Tribus | Canthonini
Péringuey, 1900
| 36
345 | Gattungen
Arten |
Taxom
| | Tribus | Coprini
Kolbe, 1805
| 14
425 | Gattungen
Arten |
Taxom
| | Tribus | Gymnopleurini
Lacordaire, 1856
| 3
87 | Gattungen
Arten |
Taxom
| | Tribus | Oniticellini
d´Orbigny, 1916
| 20
172 | Gattungen
Arten |
Taxom
| | Tribus | Onitini
Hope, 1837
| 18
209 | Gattungen
Arten |
Taxom
| | Tribus | Onthophagini
Lacordaire, 1856
Kotkäfer | 32
1468 | Gattungen
Arten |
Taxom
| | Tribus | Phanaeini
MacLeay, 1819
| 4
17 | Gattungen
Arten |
Taxom
| | Tribus | Scarabaeini
Latreille, 1802
| 10
194 | Gattungen
Arten |
Taxom
| | Tribus | Sisyphini
Mulsant, 1842
| 3
118 | Gattungen
Arten |
Taxom
|
Links, Quellen und weitere Informationen Scarabaeinae bei artfakta.se
Scarabaeinae bei dyntaxa.se
Scarabaeinae bei itis.gov
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