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| | | | Checkliste der Unterfamilie Scarabaeinae Latreille, 1802 (Pillendreher)
für die Region Eurasien (Kontinente)
Klasse
| Insecta
Linnaeus, 1758
Freikiefler, Insekten, Kerbtiere, Kerfe | 1859
65777
561707 | Familien
Gattungen
Arten |
Taxom
|
Unterklasse
| Pterygota
Lang, 1888
Fluginsekten | 1850
65672
561174 | Familien
Gattungen
Arten |
Taxom
|
Teilklasse
| Neoptera
van der Wulp, 1890
Neuflügler | 1595
64213
555201 | Familien
Gattungen
Arten |
Taxom
|
Überordnung
| Holometabola
Heider, 1889
Holometabolen Insekten | 829
47685
461834 | Familien
Gattungen
Arten |
Taxom
|
Ordnung
| Coleoptera
Linnaeus, 1758
Käfer | 209
18218
187489 | Familien
Gattungen
Arten |
Taxom
|
Unterordnung
| Polyphaga
Emery, 1886
| 167
16090
158751 | Familien
Gattungen
Arten |
Taxom
|
Teilordnung
| Scarabaeiformia
Crowson, 1960
| 18
1686
21104 | Familien
Gattungen
Arten |
Taxom
|
Überfamilie
| Scarabaeoidea
Latreille, 1802
Blatthornkäfer | 18
1686
21104 | Familien
Gattungen
Arten |
Taxom
| | Familie | Scarabaeidae
Latreille, 1802
Blatthornkäfer | 1281
16823 | Gattungen
Arten |
Taxom
| | Unterfamilie | Scarabaeinae
Latreille, 1802
Pillendreher | 126
2498 | Gattungen
Arten |
Taxom
|
Gattungen für Eurasien
126 Gattungen mit 2498 Arten #ABCDEGHIKLMNOPSTUXY
| Allogymnopleurus Janssens, 1855 | | 9 | Arten | Taxom | | Amphistomus van Lansberge, 1974 | | 10 | Arten | Taxom | Anachalcos Hope, 1837 | | 0 | Arten | Taxom | | Anoctus Sharp, 1942 | | 5 | Arten | Taxom | | Aptenocanthon Matthews, 1936 | | 1 | Art | Taxom | | Ateuchites Meunier, 1898 | | 1 | Art | Taxom | | Ateuchus Weber, 1886 | | 4 | Arten | Taxom | | Aulacopris White, 1974 | | 1 | Art | Taxom | | Bdelyropsis Pereira, Vul..., 1976 | | 1 | Art | Taxom | | Boletoscapter Matthews, 1974 | | 2 | Arten | Taxom | | Bubas Mulsant, 1811 | | 3 | Arten | Taxom | | Caccobius Thomson, 1936 | | 58 | Arten | Taxom | | Canthidium Erichson, 1981 | | 10 | Arten | Taxom | | Canthon Hoffmannsegg, 1953 | | 25 | Arten | Taxom | Canthonella Chapin, 1930 | | 0 | Arten | Taxom | | Canthonosoma MacLeay, 1871 | | 3 | Arten | Taxom | | Cassolus Sharp, 1885 | | 9 | Arten | Taxom | | Catharsius Hope, 1857 | | 37 | Arten | Taxom | | Cheironitis van Lansberge, 1789 | | 22 | Arten | Taxom | | Chironitis Lansberge, 1849 | | 5 | Arten | Taxom | | Circellium Latreille, 1781 | | 1 | Art | Taxom | | Cleptocaccobius Cambefort, 1857 | | 11 | Arten | Taxom | | Copris Geoffroy, 2009 | | 206 | Arten | Taxom | | Coproecus Reiche, 1844 | | 1 | Art | Taxom | | Coprophanaeus Olsoufieff, 1847 | | 7 | Arten | Taxom | | Coptodactyla Burmeister, 1976 | | 9 | Arten | Taxom | | Coptorhina Hope, 1833 | | 2 | Arten | Taxom | | Cretonitis Nikolajev, 2007 | | 1 | Art | Taxom | | Cretorabaeus Nikolajev, 1995 | | 1 | Art | Taxom | | Cryptocanthon Balthasar, 1973 | | 1 | Art | Taxom | | Cyobius Sharp, 1875 | | 5 | Arten | Taxom | | Cyptochirus Lesne, 1828 | | 1 | Art | Taxom | | Delopleurus Erichson, 1931 | | 3 | Arten | Taxom | | Deltochilum Eschscholtz, 1887 | | 6 | Arten | Taxom | | Demarziella Balthasar, 1976 | | 6 | Arten | Taxom | Dendropaemon Perty, 1830 | | 0 | Arten | Taxom | | Diastellopalpus van Lansberge, 1884 | | 3 | Arten | Taxom | | Dichotomius Hope, 1867 | | 10 | Arten | Taxom | | Digitonthophagus Balthasar, 2017 | | 11 | Arten | Taxom | | Diorygopyx Matthews, 1871 | | 3 | Arten | Taxom | | Disphysema Harold, 1873 | | 1 | Art | Taxom | | Drepanocerus Kirby, 1851 | | 8 | Arten | Taxom | | Eodrepanus Barbero, Pal..., 1945 | | 5 | Arten | Taxom | | Epactoides Olsoufieff, 1960 | | 1 | Art | Taxom | | Epirinus Reiche, 1880 | | 8 | Arten | Taxom | | Euoniticellus Janssens, 1873 | | 10 | Arten | Taxom | | Euonthophagus Balthasar, 1946 | | 29 | Arten | Taxom | | Eurysternus Dalman, 2009 | | 8 | Arten | Taxom | | Garreta Janssens, 1857 | | 15 | Arten | Taxom | | Glauconia Paulian, 0 | | 2 | Arten | Taxom | | Gromphas Brullé, 1869 | | 2 | Arten | Taxom | | Gymnopleurus Illiger, 1843 | | 54 | Arten | Taxom | | Haroldius Boucomont, 1993 | | 34 | Arten | Taxom | | Helictopleurus Orbigny, 2000 | | 2 | Arten | Taxom | | Heliocopris Hope, 1859 | | 21 | Arten | Taxom | | Heteronitis Gillet, 1862 | | 1 | Art | Taxom | Homalocopris Solsky, 1871 | | 0 | Arten | Taxom | | Hyalonthophagus Palestrini &..., 1855 | | 2 | Arten | Taxom | | Isocopris Pereira & Ma..., 1901 | | 1 | Art | Taxom | | Ixodina Roth, 1909 | | 2 | Arten | Taxom | | Kheper Janssens, 1798 | | 9 | Arten | Taxom | | Labroma Sharp, 1974 | | 1 | Art | Taxom | | Larhodius Balthasar, 2003 | | 3 | Arten | Taxom | | Latodrepanus Krikken, 1857 | | 2 | Arten | Taxom | | Lepanus Balthasar, 1974 | | 10 | Arten | Taxom | | Liatongus Reitter, 1933 | | 32 | Arten | Taxom | | Lithanomala Nikolajev, 1990 | | 4 | Arten | Taxom | | Litocopris Waterhouse, 1869 | | 1 | Art | Taxom | | Lobateuchus Montreuil & al., 2010 | | 1 | Art | Taxom | | Macroderes Westwood, 1880 | | 5 | Arten | Taxom | | Macropanelus | | 3 | Arten | Taxom | | Melanocanthon Halffter, 1823 | | 2 | Arten | Taxom | | Metacatharsius Paulian, 1857 | | 11 | Arten | Taxom | | Milichus Péringuey, 1871 | | 2 | Arten | Taxom | | Mimonthophagus Balthasar, 1855 | | 2 | Arten | Taxom | | Mnematidium Ritsema, 1845 | | 2 | Arten | Taxom | | Mnematium MacLeay, 1832 | | 1 | Art | Taxom | | Monoplistes Lansberge, 1888 | | 2 | Arten | Taxom | | Neosisyphus Müller, 1877 | | 9 | Arten | Taxom | | Ochicanthon Vaz-de-Mello, 2006 | | 53 | Arten | Taxom | | Odontoloma Boheman, 1868 | | 5 | Arten | Taxom | | Oniticellus Serville, 1934 | | 22 | Arten | Taxom | | Onitis Fabricius, 1886 | | 63 | Arten | Taxom | | Ontherus Erichson, 1996 | | 6 | Arten | Taxom | | Onthophagus Latreille, 1963 | Kotfresser | 1319 | Arten | Taxom | | Onychothecus Boucomont, 1992 | | 4 | Arten | Taxom | | Pachylomera Griffith & P..., 1828 | | 1 | Art | Taxom | | Pachysoma MacLeay, 1953 | | 4 | Arten | Taxom | | Panelus Lewis, 0 | | 42 | Arten | Taxom | | Parachorius Harold, 1873 | | 22 | Arten | Taxom | | Paragymnopleurus Shipp, 1997 | | 22 | Arten | Taxom | | Paraphytus Harold, 1981 | | 9 | Arten | Taxom | | Paroniticellus Balthasar, 1809 | | 1 | Art | Taxom | | Peckolus Scholtz & Ho..., 1988 | | 1 | Art | Taxom | | Pedaria Laporte, 1901 | | 5 | Arten | Taxom | | Phacosoma Boucomont, 1989 | | 2 | Arten | Taxom | | Phaedotrogus Paulian, 1972 | | 1 | Art | Taxom | | Phalops Erichson, 1867 | | 16 | Arten | Taxom | | Phanaeus MacLeay, 1994 | | 12 | Arten | Taxom | | Ponerotrogus Silvestri, 1924 | | 1 | Art | Taxom | | Prionocephale Lin, 1980 | | 1 | Art | Taxom | | Proagoderus Lansberge, 1912 | | 20 | Arten | Taxom | | Proteroscarabaeus Grabau, 1923 | | 6 | Arten | Taxom | | Pseudocanthon Bates, 1846 | | 3 | Arten | Taxom | | Pycnopanelus Arrow, 1931 | | 1 | Art | Taxom | | Sarophorus Erichson, 1840 | | 2 | Arten | Taxom | | Sauvagesinella Paulian, 2002 | | 2 | Arten | Taxom | | Scarabaeus Linnaeus, 1901 | | 60 | Arten | Taxom | | Sceliages Westwood, 2002 | | 1 | Art | Taxom | | Scelocopris Zhang, 1989 | | 1 | Art | Taxom | | Scybalocanthon Martínez, 1855 | | 2 | Arten | Taxom | | Sinapisoma Boucomont, 1928 | | 1 | Art | Taxom | | Sinodrepanus Simonis, 2004 | | 9 | Arten | Taxom | | Sisyphus Latreille, 0 | | 35 | Arten | Taxom | | Stiptopodius Harold, 1923 | | 1 | Art | Taxom | | Sulcophanaeus Olsoufieff, 1856 | | 2 | Arten | Taxom | Sylvicanthon Halffter & M..., 1977 | | 0 | Arten | Taxom | | Synapsis Forster, 1933 | | 28 | Arten | Taxom | | Temnoplectron Westwood, 2017 | | 6 | Arten | Taxom | | Tesserodon Hope, 2017 | | 3 | Arten | Taxom | | Tibiodrepanus Krikken, 1840 | | 6 | Arten | Taxom | | Tiniocellus Péringuey, 1851 | | 6 | Arten | Taxom | Trichillum Harold, 1868 | | 0 | Arten | Taxom | | Uroxys Westwood, 1874 | | 4 | Arten | Taxom | | Xinidium Harold, 1869 | | 1 | Art | Taxom | | Yvescambefortius Ochi & Kon, 1926 | | 1 | Art | Taxom |
Artenliste für Eurasien
2498 Arten und 112 Unterarten | Allogymnopleurus aeneus Harold, 1875 | | | Allogymnopleurus alluaudi Garreta, 1914 | | | Allogymnopleurus anthracinus Klug, 1845 | | | Allogymnopleurus consocius Péringuey, 1901 | | | Allogymnopleurus maculosus MacLeay, 1821 | | | Allogymnopleurus signaticollis Waterhouse, 1890 | | | Allogymnopleurus spilotus MacLeay, 1821 | | | Allogymnopleurus splendidus Bertoloni, 1849 | | | Allogymnopleurus thalassinus Klug, 1855 | | | Amphistomus alfurorum Huijbregts &..., 2007 | | | Amphistomus calcaratus MacLeay, 1871 | | | Amphistomus complanatus Matthews, 1974 | | | Amphistomus fasciculatus Gillet, 1927 | | | Amphistomus primonactus Matthews, 1974 | | | Amphistomus speculatus Gillet, 1927 | | | Amphistomus speculifer Matthews, 1974 | | | Amphistomus squalidus MacLeay, 1887 | | | Amphistomus storeyi Matthews, 1974 | | | Amphistomus trispiculatus Matthews, 1974 | | | Anoctus laevis Sharp, 1875 | | | Anoctus myrmecophilus Arrow, 1907 | | | Anoctus rasilis Boucomont, 1921 | | | Anoctus scatonomoides Lansberge, 1885 | | | Anoctus siamensis Balthasar, 1942 | | | Aptenocanthon hopsoni Carter, 1936 | | | Ateuchites grandis ♰ Meunier, 1898 | | | Ateuchus aeneomicans Harold, 1868 | | | Ateuchus histeroides Weber, 1801 | | | Ateuchus pygidialis Harold, 1868 | | | Ateuchus rodriguezi Preudhomme d..., 1886 | | | Aulacopris maximus Matthews, 1974 | | | Bdelyropsis venezuelensis Howden, 1976 | | | Boletoscapter cornutus MacLeay, 1887 | | | Boletoscapter furcatus Matthews, 1974 | | | Bubas bison Linnaeus, 1767 | | | Bubas bubaloides Janssens, 1938 | | | Bubas bubalus Olivier, 1811 | | | Caccobius (Caccobius) chujoi Paulian, 1942 | | | Caccobius (Caccophilus) scrofa (Balthasar, 1942) | ⇒ Cleptocaccobius scrofa Balthasar, 1942 | | Caccobius (Caccophilus) tortus Sharp, 1875 | | | Caccobius tortus matsudai Ochi & Araya, 1996 | | | Caccobius tortus tortus | | | Caccobius (Caccophilus) unicornis Fabricius, 1798 | | | Caccobius aterrimus Fabricius, 1798 | | | Caccobius balthasari (Tesar, 1969) | ⇒ Cleptocaccobius balthasarianus Cambefort, 1979 | | Caccobius bawangensis Ochi & Kon &..., 1997 | | | Caccobius bidentatus Boucomont, 1923 | | | Caccobius binodulus Harold, 1877 | | | Caccobius boucomonti Balthasar, 1935 | | | Caccobius brevis Waterhouse, 1875 | | | Caccobius castaneus (Klug, 1855) | ⇒ Caccocnemus castaneus Klug, 1855 | | Caccobius christophi Harold, 1879 | | | Caccobius demangei Boucomont, 1919 | | | Caccobius denticollis Harold, 1867 | | | Caccobius devagiriensis Nithya & Sabu, 2013 | | | Caccobius dickinsoni Masumoto, Oc..., 2022 | | | Caccobius diminutivus Walker, 1858 | | | Caccobius excavatus Bai, Zhang &..., 2007 | | | Caccobius ferrugineus Fahraeus, 1857 | | | Caccobius flavolimbatus Balthasar, 1942 | | | Caccobius fukiensis Balthasar, 1942 | | | Caccobius gallinus Arrow, 1907 | | | Caccobius gilleti Balthasar, 1933 | | | Caccobius gonoderus Fairmaire, 1888 | | | Caccobius himalayanus Jekel, 1872 | | | Caccobius hirayamai (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccobius histeroides Ménétriés, 1832 | | | Caccobius imitans Balthasar, 1933 | | | Caccobius indicus Harold, 1867 | | | Caccobius jessoensis Harold, 1867 | | | Caccobius jononis (Matsumura, 1936) | ⇒ Gibbonthophagus viduus Harold, 1875 | | Caccobius kelleri Olsoufieff, 1907 | | | Caccobius maruyamai Masumoto, Oc..., 2012 | | | Caccobius masumotoi Cambefort, 1990 | | | Caccobius meridionalis Boucomont, 1914 | | | Caccobius mundus Ménétriés, 1838 | | | Caccobius nigritulus Klug, 1855 | | | Caccobius nikkoensis Lewis, 1895 | | | Caccobius obtusus Fahraeus, 1857 | | | Caccobius pantherinus Arrow, 1931 | | | Caccobius pulicarius Harold, 1875 | | | Caccobius pullus Jekel, 1872 | | | Caccobius rufipennis Motschulsky, 1858 | | | Caccobius sappoensis Matsumura, 1936 | | | Caccobius sapporensis (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccobius scheuerni Cambefort, 1990 | | | Caccobius schreberi Linnaeus, 1767 | | | Caccobius sordidus Harold, 1886 | | | Caccobius suzukii (Matsumura, 1936) | ⇒ Caccobius boucomonti Balthasar, 1935 | | Caccobius torticornis Arrow, 1931 | | | Caccobius tsunoellus (Matsumara, 1938) | ⇒ Caccophilus tortus Sharp, 1875 | | Caccobius ultor Sharp, 1875 | | | Caccobius vulcanus Fabricius, 1801 | | | Caccobius yamauchii (Matsumura, 1936) | ⇒ Caccobius unicornis Fabricius, 1798 | | Caccobius yubariensis (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Canthidium (Eucanthidium) macclevei Kohlmann & S..., 2006 | | | Canthidium annagabrielae Solis & Kohl..., 2004 | | | Canthidium ardens Bates, 1887 | | | Canthidium delgadoi Kohlmann & S..., 2006 | | | Canthidium hespenheidei Howden & Young, 1981 | | | Canthidium pallidoalatum Howden & Young, 1981 | | | Canthidium perceptibile Howden & Young, 1981 | | | Canthidium smaragdinum Harold, 1867 | | | Canthidium subdopuncticolle Howden & Young, 1981 | | | Canthidium vespertinum Howden & Young, 1981 | | | Canthon (Boreocanthon) depressipennis LeConte, 1859 | | | Canthon (Boreocanthon) ebenus Say, 1823 | | | Canthon (Boreocanthon) lecontei Harold, 1868 | | | Canthon (Boreocanthon) mixtus Robinson, 1948 | | | Canthon (Boreocanthon) praticola Le Conte, 1859 | | | Canthon (Boreocanthon) probus Germar, 1824 | | | Canthon (Boreocanthon) puncticollis (LeConte, 1866) | ⇒ Canthon punctatus Schmidt, 1922 | | Canthon (Boreocanthon) simplex Le Conte, 1857 | | | Canthon (Canthon) chalcites Haldeman, 1843 | | | Canthon (Canthon) humectus Say, 1831 | | | Canthon (Canthon) imitator Brown, 1946 | | | Canthon (Canthon) indigaceus (LeConte, 1866) | ⇒ Canthon humectus Say, 1831 | | Canthon (Canthon) pilularius Linnaeus, 1758 | Pillenkäfer | | Canthon (Canthon) vigilans LeConte, 1858 | | | Canthon (Glaphyrocanthon) femoralis Chevrolat, 1834 | | | Canthon (Glaphyrocanthon) viridis (Palisot de B..., 1805) | ⇒ Epilissus viridis Klug, 1832 | | Canthon bicolor Castelnau, 1840 | | | Canthon deyrollei Harold, 1868 | | | Canthon dives Harold, 1868 | | | Canthon euryscelis Bates, 1887 | | | Canthon fulgidus Redtenbacher, 1867 | | | Canthon leechi Martinez, Ha..., 1964 | | | Canthon semiopacus Harold, 1868 | | | Canthon simulans Martínez, 1950 | | | Canthon staigi Pereira, 1953 | | | Canthonosoma castelnaui Harold, 1868 | | | Canthonosoma macleayi Harold, 1868 | | | Canthonosoma mastersi MacLeay, 1871 | | | Cassolus fukiensis Balthasar, 1960 | | | Cassolus gotoi Masumoto, 1986 | | | Cassolus humeralis Arrow, 1907 | | | Cassolus javanus Boucomont, 1914 | | | Cassolus minutus Boucomont, 1914 | | | Cassolus nudus Sharp, 1875 | | | Cassolus peninsularis Arrow, 1907 | | | Cassolus pongchaii Masumoto, 1989 | | | Cassolus sumatranus Lansberge, 1885 | | | Catharsius aethiops Sharp, 1875 | | | Catharsius alpheus Gillet, 1932 | | | Catharsius birmanensis Lansberge, 1874 | | | Catharsius borneensis (Paulian, 1936) | ⇒ Catharsius molossus Linnaeus, 1758 | | Catharsius bradshawi Lansberge, 1887 | | | Catharsius brutus Harold, 1880 | | | Catharsius calaharicus Kolbe, 1893 | | | Catharsius capucinus Fabricius, 1781 | | | Catharsius dayacus Lansberge, 1886 | | | Catharsius dubius (Péringuey, 1901) | ⇒ Catharsius molossus Linnaeus, 1758 | | Catharsius dux Harold, 1878 | | | Catharsius eteocles (Laporte, 1840) | ⇒ Catharsius alpheus Gillet, 1932 | | Catharsius furcillatus Guérin-Ménev..., 1847 | | | Catharsius granulatus Sharp, 1875 | | | Catharsius guineensis Ferreira, 1963 | | | Catharsius harpagus Harold, 1877 | | | Catharsius heros Boheman, 1860 | | | Catharsius javanus Lansberge, 1886 | | | Catharsius kangeanus (Paulian, 1936) | ⇒ Catharsius molossus Linnaeus, 1758 | | Catharsius marcellus Kolbe, 1893 | | | Catharsius melancholicus (Boheman, 1860) | ⇒ Catharsius longiceps Gillet, 1907 | | Catharsius molossus Linnaeus, 1758 | Dickkopf | | Catharsius nathani Frey, 1956 | | | Catharsius pandion Harold, 1877 | | | Catharsius phidias (Olivier, 1789) | ⇒ Catharsius platicornis Ferreira, 1960 | | Catharsius philus Kolbe, 1893 | | | Catharsius pithecius Fabricius, 1775 | Indianische Schulteraale | | Catharsius platicornis Ferreira, 1960 | | | Catharsius platypus Sharp, 1875 | | | Catharsius quadridentatus Lansberge, 1885 | | | Catharsius renaudpauliani Ochi & Kon, 1996 | | | Catharsius sagax Quenstedt, 1806 | | | Catharsius satyrus Kolbe, 1893 | | | Catharsius sesostris Waterhouse, 1888 | | | Catharsius timorensis (Lansberge, 1879) | ⇒ Catharsius molossus Linnaeus, 1758 | | Catharsius tricornutus DeGeer, 1778 | | | Catharsius ulysses Boheman, 1857 | | | Cheironitis arrowi Janssens, 1937 | | | Cheironitis audens Péringuey, 1901 | | | Cheironitis candezei Lansberge, 1875 | | | Cheironitis eumenes Gebler, 1860 | | | Cheironitis furcifer Rossi, 1792 | | | Cheironitis haroldi Ballion, 1870 | | | Cheironitis hoplosternus Harold, 1868 | | | Cheironitis hungaricus Herbst, 1789 | | | Cheironitis indicus (Lansberge, 1875) | ⇒ Cheironitis arrowi Janssens, 1937 | | Cheironitis irroratus Rossi, 1790 | | | Cheironitis klapperichi (Balthasar, 1955) | ⇒ Cheironitis furcifer Rossi, 1792 | | Cheironitis moeris Pallas, 1781 | | | Cheironitis osiridis Reiche, 1856 | | | Cheironitis pamphilus Ménétriés, 1849 | | | Cheironitis petraensis Coppo, 2021 | | | Cheironitis phoebus Reitter, 1892 | | | Cheironitis scabrosus Fabricius, 1776 | | | Cheironitis socotranus Gahan, 1900 | | | Cheironitis sterculius Ballion, 1870 | | | Cheironitis ungaricus Herbst, 1789 | | | Cheironitis ungaricus irroratus (Rossi, 1790) | ⇒ Cheironitis irroratus Rossi, 1790 | | Cheironitis ungaricus ungaricus Herbst, 1789 | | | Chironitis furcifer Rossi, 1792 | | | Chironitis hungaricus (Herbst, 1789) | ⇒ Cheironitis ungaricus Herbst, 1789 | | Chironitis irroratus Rossi, 1790 | | | Chironitis moeris Pallas, 1781 | | | Chironitis pamphilus Ménétriése, 1849 | | | Circellium bacchus Fabricius, 1781 | Riesen-Pillendreher | | Cleptocaccobius arrowi Cambefort, 1985 | | | Cleptocaccobius balthasarianus Cambefort, 1979 | | | Cleptocaccobius boucomonti Cambefort, 1985 | | | Cleptocaccobius durantoni Cambefort, 1985 | | | Cleptocaccobius inermis Arrow, 1931 | | | Cleptocaccobius khatimae Cambefort, 1985 | | | Cleptocaccobius scrofa Balthasar, 1942 | | | Cleptocaccobius simplex Boucomont, 1923 | | | Cleptocaccobius simplex meridionalis Cambefort, 1985 | | | Cleptocaccobius simplex simplex | | | Cleptocaccobius viridicollis Fahraeus, 1857 | | | Copris (Copris) cambodiensis Ochi, Kon & ..., 2008 | | | Copris cambodiensis cambodiensis | | | Copris cambodiensis masumotoi Ochi, Kon & ..., 2011 | | | Copris (Copris) fukiensis Balthasar, 1952 | | | Copris (Copris) kachinensis Ochi, Kon & ..., 2008 | | | Copris (Copris) kartlinus ♰ O. N. Kabakov, 1988 | | | Copris (Copris) punctatus Gillet, 1910 | | | Copris punctatus formosanus Gillet, 1922 | | | Copris punctatus punctatus | | | Copris (Copris) schoolmeestersi Ochi & Kon, 2010 | | | Copris (Copris) tripartitus Waterhouse, 1875 | | | Copris (Microcopris) propinquus Felsche, 1910 | | | Copris (Paracopris) cariniceps Felsche, 1910 | | | Copris (Sinocopris) barclayi Ochi, Kon & Bai, 2009 | | | Copris (Sinocopris) pecuarius Lewis, 1884 | | | Copris pecuarius continentalis Ochi & Kon, 2010 | | | Copris (Sinocopris) uenoi Hanboonsong ..., 2003 | | | Copris acutidens Motschulsky, 1860 | | | Copris agnus Sharp, 1875 | | | Copris amyntor (Klug, 1855) | ⇒ Copris harrisi Waterhouse, 1891 | | Copris anceus Olivier, 1789 | | | Copris andrewesi Waterhouse, 1891 | | | Copris angusticornis Arrow, 1933 | | | Copris anomiopseoides (Boucomont, 1924) | ⇒ Paracopris anomiopseoides Boucomont, 1924 | | Copris antares Bacescu & Ca..., 1947 | | | Copris apicepunctatus Balthasar, 1942 | | | Copris arimotoi Ochi & Araya, 1994 | | | Copris armeniacus Falderman, 1835 | | | Copris armiger Gillet, 1910 | | | Copris arrowi Felsche, 1910 | | | Copris bartolozzi Ochi, Kon & ..., 2019 | | | Copris basipunctatus Balthasar, 1942 | | | Copris bellator Chevrolat, 1844 | | | Copris bengalensis Shiino, 1978 | | | Copris brachypterus Nomura, 1964 | | | Copris brachypterus brachypterus | | | Copris brachypterus toguchii Ochi & Kawahara, 2012 | | | Copris caelatus Fabricius, 1794 | | | Copris calvus Sharp, 1875 | | | Copris caobangensis Bui, Dumack ..., 2018 | | | Copris capensis Waterhouse, 1891 | | | Copris carinicus Gillet, 1910 | | | Copris cassius Péringuey, 1901 | | | Copris celebensis Lansberge, 1886 | | | Copris cheni Ochi, Kon & Bai, 2007 | | | Copris chinmontanus Ochi, Kon & ..., 2019 | | | Copris compressipennis (Gillet, 1910) | ⇒ Paracopris compressipennis Gillet, 1910 | | Copris confucius Harold, 1877 | | | Copris continentalis Ochi & Kon, 2010 | | | Copris corniculatus (Mulsant, 1842) | ⇒ Copris lunaris Linnaeus, 1758 | | Copris cornifrons Boheman, 1860 | | | Copris corniger (Sahlberg, 1823) | ⇒ Onthophagus taurus Schreber, 1759 | | Copris corpulentus Gillet, 1910 | | | Copris costaricensis Gahan, 1894 | | | Copris cribratus Gillet, 1927 | | | Copris davidis Deyrolle, 1878 | | | Copris davisoni Waterhouse, 1891 | | | Copris delicatus Arrow, 1931 | | | Copris denticulatus Nguyen-phung, 1988 | | | Copris doriae Harold, 1877 | | | Copris druidum ♰ O. Heer, 1862 | | | Copris elphenor Klug, 1855 | | | Copris erratus Lansberge, 1886 | | | Copris erratus erratus | | | Copris erratus masayukii Ochi & Kon, 2004 | | | Copris evanidus Klug, 1855 | | | Copris excisus Waterhouse, 1891 | | | Copris fallaciosus Gillet, 1907 | | | Copris felschei (Reitter, 1893) | ⇒ Copris armeniacus Falderman, 1835 | | Copris fidius Olivier, 1789 | | | Copris frankenbergerianus Balthasar, 1958 | | | Copris fricator Fabricius, 1787 | | | Copris fricator fricator | | | Copris fujiokai Ochi & Kon, 1996 | | | Copris fujiokai poringensis Ochi & Kon, 2005 | | | Copris furciceps Felsche, 1910 | | | Copris gibbulus Lansberge, 1886 | | | Copris gibbulus borneensis Ochi & Kon, 2005 | | | Copris gibbulus gibbulus | | | Copris gladiator Arrow, 1933 | | | Copris gracilis Waterhouse, 1891 | | | Copris harrisi Waterhouse, 1891 | | | Copris hatayamai Ochi & Araya, 1992 | | | Copris hidakai Ochi & Kon, 1996 | | | Copris hispanus Linnaeus, 1764 | Schwarzglänzende Spanische Nashornkäfer | | Copris hispanus cavolinii Petagna, 1792 | | | Copris hispanus hispanus Linné, 1764 | | | Copris hispanus persianus Miksic, 1958 | | | Copris hosakai Ochi & Kon, 2014 | | | Copris howdeni Matthews & H..., 1959 | | | Copris illotus Balthasar, 1942 | | | Copris imitans Felsche, 1910 | | | Copris inaequabilis Zhang, 1997 | | | Copris incertus Say, 1835 | | | Copris indicus (Gillet, 1910) | ⇒ Copris fricator Fabricius, 1787 | | Copris interioris Kolbe, 1897 | | | Copris iris Sharp, 1875 | | | Copris jacchoides Nguyen-phung..., 1987 | | | Copris javanensis Bui, Ziegler..., 2019 | | | Copris johkii Ochi & Kon, 2014 | | | Copris jucundus Gillet, 1932 | | | Copris kasagii Ochi & Kon, 1996 | | | Copris keralensis Gill, 1986 | | | Copris kimioi Ochi & Kon, 2021 | | | Copris kiriromensis Ochi & Kon, 2021 | | | Copris kiuchii Masumoto, 1989 | | | Copris klapperichi Balthasar, 1942 | | | Copris kusakabei Ochi, Kon & ..., 2005 | | | Copris laevigatus (Gillet, 1927) | ⇒ Onthophagus laevigatus Fabricius, 1798 | | Copris lannathai Hanboonsong ..., 2003 | | | Copris laosensis Ochi & Kon, 2021 | | | Copris lugubris Boheman, 1858 | | | Copris lunaris Linnaeus, 1758 | Kleine Mondhornkäfers | | Copris macacus Lansberge, 1886 | | | Copris macer Péringuey, 1901 | | | Copris magicus Harold, 1881 | | | Copris manni Ochi, Kon & Bai, 2009 | | | Copris marcoberrai Ochi, Kon & ..., 2019 | | | Copris marcus Gillet, 1921 | | | Copris medogensis Y.-W. Zhang, 1988 | | | Copris mesacanthus Harold, 1878 | | | Copris micros Ochi & Kon, 2021 | | | Copris mindorensis Ochi, 1992 | | | Copris minutus Drury, 1773 | | | Copris misellus Péringuey, 1901 | | | Copris miyakei Ochi & Kon, 2004 | | | Copris moffartsi Gillet, 1907 | | | Copris mongkhoni Hanboonsong ..., 2003 | | | Copris montreuili Ochi, Kon & ..., 2019 | | | Copris neglectus Moxey, 1963 | | | Copris nepos Gillet, 1908 | | | Copris nevinsoni Waterhouse, 1891 | | | Copris nonarmatus Ochi & Kon, 2021 | | | Copris novaki Zelenka, 1992 | | | Copris numa Lansberge, 1886 | | | Copris obenbergeri Balthasar, 1933 | | | Copris obesus Boheman, 1857 | | | Copris ochus Motschulsky, 1860 | | | Copris olivieri Illiger, 1800 | | | Copris orion Klug, 1835 | | | Copris orphanus (Guérin-Ménev..., 1847) | ⇒ Copris evanidus Klug, 1855 | | Copris parapecuarius Ochi, Kon & ..., 2008 | | | Copris parasonensis Ochi, Kon & ..., 2019 | | | Copris paraszechouanicus Ochi, Kon & ..., 2018 | | | Copris peculiariprotibialis Ochi & Kon, 2023 | | | Copris pedarioides Lansberge, 1886 | | | Copris poggii Ochi & Kon, 2005 | | | Copris popaensis Ochi, Kon & ..., 2019 | | | Copris potanini Semenov, 1891 | | | Copris pseudochus Ochi & Kon, 2004 | | | Copris pseudochus basipunctulatus Ochi, Kon & ..., 2020 | | | Copris pseudochus chuduraziensis Ochi, Kon & ..., 2019 | | | Copris pseudochus pseudochus | | | Copris pseudosinicus Balthasar, 1958 | | | Copris pueli Mollandin De..., 1905 | | | Copris puncticollis Boheman, 1857 | | | Copris punctipennis Boucomont, 1914 | | | Copris punctulatus Wiedemann, 1823 | | | Copris punctulatus punctulatus | | | Copris punctulatus septentrionalis Ochi, Kon & ..., 2019 | | | Copris punjabensis Gillet, 1921 | | | Copris quasilaevigatus Ochi, Kon & Bai, 2007 | | | Copris ramosiceps Gillet, 1921 | | | Copris reflexus (Fabricius, 1787) | ⇒ Copris minutus Drury, 1773 | | Copris repertus Walker, 1858 | | | Copris ribbei Lansberge, 1886 | | | Copris riekoae Ochi & Kon, 2003 | | | Copris ruricola Balthasar, 1933 | | | Copris sabinus Gillet, 1910 | | | Copris sacontala Redtenbacher, 1848 | | | Copris sarpedon Harold, 1868 | | | Copris saundersi Harold, 1869 | | | Copris scorpio Zelenka, 1993 | | | Copris siamensis Gillet, 1910 | | | Copris siangensis Biswas, 1980 | | | Copris signatus Walker, 1858 | | | Copris sinicus Hope, 1842 | | | Copris sinon Olivier, 1789 | | | Copris sodalis Walker, 1858 | | | Copris sonensis Bui, Dumack ..., 2018 | | | Copris sorex Balthasar, 1942 | | | Copris sphaeropterus Harold, 1877 | | | Copris spinator Harold, 1881 | | | Copris subdolus Balthasar, 1958 | | | Copris subsidens Péringuey, 1901 | | | Copris subterranea ♰ O. Heer, 1862 | | | Copris subterraneus Heer, 1862 | | | Copris sumatrensis Gillet, 1921 | | | Copris surdus Arrow, 1931 | | | Copris szechouanicus Balthasar, 1958 | | | Copris takanoi Ochi, Kon & Bai, 2020 | | | Copris teramachii Ochi, Kon & Bai, 2020 | | | Copris tetraodon Gillet, 1910 | | | Copris tricorniprothoracicus Ochi, Kon & ..., 2020 | | | Copris tsukamotoi Ochi, Kon & ..., 2008 | | | Copris typhoeus Gerstaecker, 1884 | | | Copris umbilicatus Abeille de P..., 1901 | | | Copris urus (Boheman, 1857) | ⇒ Catharsius molossus Linnaeus, 1758 | | Copris vietnamicus Kabakov, 1994 | | | Copris vietnamimontanus Ochi, Kon & ..., 2018 | | | Copris vilhenai Ferreira, 1962 | | | Copris vitalisi Gillet, 1921 | | | Copris wiesei Kolbe, 1914 | | | Copris xingkeyangi Ochi, Kon & Bai, 2020 | | | Copris yangi Ochi, Kon & Bai, 2007 | | | Copris zhangi Ochi, Kon & Bai, 2009 | | | Coproecus hemisphaericus Guérin-Ménev..., 1844 | | | Coprophanaeus (Coprophanaeus) pluto Harold, 1863 | | | Coprophanaeus (Megaphanaeus) bellicosus Olivier, 1789 | | | Coprophanaeus (Megaphanaeus) ensifer Germar, 1821 | | | Coprophanaeus (Megaphanaeus) lancifer Linnaeus, 1767 | Pikenträger | | Coprophanaeus boucardi Nevinson, 1891 | | | Coprophanaeus corythus Harold, 1863 | | | Coprophanaeus telamon Erichson, 1847 | | | Coptodactyla depressa Paulian, 1933 | | | Coptodactyla glabricollis Hope, 1842 | | | Coptodactyla merdeka Reid, 2000 | | | Coptodactyla meridionalis Matthews, 1976 | | | Coptodactyla monstrosa Felsche, 1909 | | | Coptodactyla nitida (Paulian, 1933) | ⇒ Coptodactyla subaenea Harold, 1877 | | Coptodactyla onitoides Gillet, 1925 | | | Coptodactyla subaenea Harold, 1877 | | | Coptodactyla torresica Mathews, 1976 | | | Coptorhina auspicata Péringuey, 1901 | | | Coptorhina klugi Hope, 1833 | | | Cretonitis copripes ♰ Nikolajev, 2007 | | | Cretorabaeus elongatus ♰ G. V. Nikolajev, 1995 | | | Cryptocanthon nebulinus Howden, 1973 | | | Cyobius brendelli Ochi, Kon & ..., 2010 | | | Cyobius cheyi Ochi, Kon & ..., 2006 | | | Cyobius mendeli Ochi, Kon & ..., 2010 | | | Cyobius tsukamotoi Ochi, Kon & ..., 2008 | | | Cyobius wallacei Sharp, 1875 | | | Cyptochirus ambiguus (Kirby, 1828) | ⇒ Drepanocerus trogiformis Roth, 1851 | | Delopleurus mencli Kral, 2014 | | | Delopleurus parvus Sharp, 1875 | | | Delopleurus striatus Arrow, 1931 | | | Deltochilum gibbosum Fabricius, 1775 | Amerikanische Höckerflügel | | Deltochilum hypponum Buquet, 1844 | | | Deltochilum lobipes Bates, 1887 | | | Deltochilum mexicanum Burmeister, 1848 | | | Deltochilum scabriusculum Bates, 1887 | | | Deltochilum valgum acropyge Bates, 1887 | | | Demarziella geminata MacLeay, 1871 | | | Demarziella imitatrix Balthasar, 1967 | | | Demarziella interrupta Carter, 1936 | | | Demarziella metallica Carter, 1936 | | | Demarziella mirifica Balthasar, 1961 | | | Demarziella pratensis Matthews, 1976 | | | Diastellopalpus infernalis Lansberge, 1887 | | | Diastellopalpus johnstoni Waterhouse, 1885 | | | Diastellopalpus lamellicollis Quedenfeldt, 1884 | | | Dichotomius (Luederwaldtinia) centralis Harold, 1869 | | | Dichotomius (Luederwaldtinia) schiffleri Vaz-de-Mello..., 2001 | | | Dichotomius (Luederwaldtinia) yucatanus Bates, 1887 | | | Dichotomius amicitiae Kohlmann & S..., 1997 | | | Dichotomius annae Linnaeus, 1767 | | | Dichotomius carolinus Linnaeus, 1767 | | | Dichotomius colonicus (Say, 1835) | ⇒ Dichotomius carolinus Linnaeus, 1767 | | Dichotomius melzeri Luederwaldt, 1922 | | | Dichotomius robustus Luederwaldt, 1935 | | | Dichotomius satanas Harold, 1867 | | | Digitonthophagus aksumensis Génier, 2017 | | | Digitonthophagus biflagellatus Génier, 2017 | | | Digitonthophagus bonasus Fabricius, 1775 | Tranquebarische dreygehörnte Kopf | | Digitonthophagus catta Fabricius, 1787 | | | Digitonthophagus eucatta Génier, 2017 | | | Digitonthophagus falciger Génier, 2017 | | | Digitonthophagus fimator Génier, 2017 | | | Digitonthophagus gazella Fabricius, 1787 | | | Digitonthophagus sahelicus Moretto, 2017 | | | Digitonthophagus uks Génier, 2017 | | | Digitonthophagus ulcerosus Génier, 2017 | | | Diorygopyx asciculifer Matthews, 1974 | | | Diorygopyx simpliciclunis Matthews, 1974 | | | Diorygopyx tibialis MacLeay, 1871 | | | Disphysema candezei Harold, 1873 | | | Drepanocerus ambiguus Kirby, 1828 | | | Drepanocerus arrowi (Bathasar, 1932) | ⇒ Sinodrepanus rex Boucomont, 1912 | | Drepanocerus kirbyi Kirby, 1828 | | | Drepanocerus planus Fabricius, 1801 | | | Drepanocerus rex (Boucomont, 1912) | ⇒ Sinodrepanus rex Boucomont, 1912 | | Drepanocerus runicus (Arrow, 1909) | ⇒ Ixodina runica Arrow, 1909 | | Drepanocerus sinicus (Harold, 1868) | ⇒ Tibiodrepanus sinicus Harold, 1868 | | Drepanocerus trogiformis Roth, 1851 | | | Eodrepanus coopei Barbero, Pal..., 2009 | | | Eodrepanus fastiditus Péringuey, 1901 | | | Eodrepanus integriceps Janssens, 1953 | | | Eodrepanus liuchungloi Kryzhanovsky..., 1966 | | | Eodrepanus striatulus Paulian, 1945 | | | Epactoides basilewskyi Balthasar, 1960 | | | Epirinus aeneus Wiedemann, 1823 | | | Epirinus asper Péringuey, 1901 | | | Epirinus comosus Péringuey, 1901 | | | Epirinus flagellatus Fabricius, 1775 | | | Epirinus gratus Péringuey, 1901 | | | Epirinus obtusus Boheman, 1857 | | | Epirinus pseudorugosus Medina & Sch..., 2005 | | | Epirinus scrobiculatus Harold, 1880 | | | Euoniticellus africanus Harold, 1873 | | | Euoniticellus cambeforti Masumoto, 1987 | | | Euoniticellus fulvus Goeze, 1777 | | | Euoniticellus inaequalis Reiche, 1850 | | | Euoniticellus intermedius Reiche, 1848 | | | Euoniticellus kawanus Janssens, 1939 | | | Euoniticellus nasicornis Reiche, 1849 | | | Euoniticellus pallens Olivier, 1789 | | | Euoniticellus pallipes Fabricius, 1781 | | | Euoniticellus triangulatus Harold, 1873 | | | Euonthophagus aeneobrunneus Kabakov, 1977 | | | Euonthophagus amyntas Olivier, 1789 | | | Euonthophagus amyntas alces Fabricius, 1792 | | | Euonthophagus amyntas amyntas Olivier, 1789 | | | Euonthophagus amyntas auchenia Redtenbacher, 1850 | | | Euonthophagus amyntas subviolaceus Ménétries, 1832 | | | Euonthophagus atramentarius Ménétriés, 1832 | | | Euonthophagus bedeli Reitter, 1891 | | | Euonthophagus carbonarius Klug, 1855 | | | Euonthophagus conterminus Petrovitz, 1971 | | | Euonthophagus crocatus Mulsant & Go..., 1870 | | | Euonthophagus crocatus crocatus | | | Euonthophagus dorbignyi Olsoufieff, 1900 | | | Euonthophagus dorbignyi neutto Olsoufieff, 1900 | | | Euonthophagus gibbosus Scriba, 1790 | | | Euonthophagus hazariensis Kabakov, 1977 | | | Euonthophagus koshantschikoffi Reitter, 1891 | | | Euonthophagus laghmanicus Kabakov, 1977 | | | Euonthophagus loeffleri Petrovitz, 1965 | | | Euonthophagus maindroni d´Orbigny, 1897 | | | Euonthophagus mostafatsairii Palestrini, ..., 1979 | | | Euonthophagus pentaceros Olsoufieff, 1900 | | | Euonthophagus pertinax Balthasar, 1959 | | | Euonthophagus schnabeli Splichal, 1910 | | | Euonthophagus sulcicollis Reitter, 1892 | | | Euonthophagus tissoni Reitter, 1906 | | | Euonthophagus verberatus Ziani, 2006 | | | Euonthophagus yama Ziani, 2006 | | | Euonthophagus yemenicus Balthasar, 1946 | | | Eurysternus arnaudi Génier, 2009 | | | Eurysternus caribaeus Herbst, 1789 | | | Eurysternus cayennensis Castelnau, 1840 | | | Eurysternus hirtellus Dalman, 1824 | | | Eurysternus magnus Castelnau, 1840 | | | Eurysternus marmoreus Castelnau, 1840 | | | Eurysternus mexicanus Harold, 1869 | | | Eurysternus superbus Génier, 2009 | | | Garreta azareus | | | Garreta dejeani Castelnau, 1840 | | | Garreta gilleti Garreta, 1914 | | | Garreta matabelensis Janssens, 1938 | | | Garreta mombelgi Boucomont, 1929 | | | Garreta morosus Fairmaire, 1886 | | | Garreta mundus Wiedemann, 1819 | | | Garreta nitens Olivier, 1789 | | | Garreta opacus Redtenbacker, 1848 | | | Garreta ruficornis Motschulsky, 1854 | | | Garreta smaragdifer Walker, 1858 | | | Garreta sumptuosus Castelnau, 1840 | | | Garreta sylvestris Mittal, 2011 | | | Garreta unicolor Fahraeus, 1857 | | | Garreta wahlbergi Fahraeus, 1857 | | | Glauconia kefersteini | | | Glauconia undulata | | | Gromphas aeruginosa Perty, 1830 | | | Gromphas inermis Harold, 1869 | | | Gymnopleurus aciculatus Gebler, 1841 | | | Gymnopleurus aenescens Wiedemann, 1821 | | | Gymnopleurus aeruginosus Harold, 1867 | | | Gymnopleurus aethiops (Sharp, 1875) | ⇒ Paragymnopleurus melanarius Harold, 1867 | | Gymnopleurus assamensis (Waterhouse, 1890) | ⇒ Paragymnopleurus sinuatus assamensis Waterhouse, 1890 | | Gymnopleurus atratus Klug, 1845 | | | Gymnopleurus barovskyi (Kieseritsky, 1928) | ⇒ Gymnopleurus flagellatus Fabricius, 1787 | | Gymnopleurus bicallosus Felsche, 1909 | | | Gymnopleurus biharensis Arrow, 1931 | | | Gymnopleurus bombayensis Arrow, 1931 | | | Gymnopleurus coriarius (Herbst, 1789) | ⇒ Gymnopleurus flagellatus Fabricius, 1787 | | Gymnopleurus cyaneus (Fabricius, 1798) | ⇒ Gymnopleurus sericeifrons Fairmaire, 1887 | | Gymnopleurus deperditus ♰ O. Heer, 1862 | | | Gymnopleurus elegans Klug, 1845 | | | Gymnopleurus eocaenicus ♰ F. Meunier, 1921 | | | Gymnopleurus flagellatus Fabricius, 1787 | | | Gymnopleurus flagellatus asperatus Motschulsky, 1849 | | | Gymnopleurus flagellatus calignosus Balthasar, 1934 | | | Gymnopleurus flagellatus flagellatus Fabricius, 1787 | | | Gymnopleurus flagellatus hornei Waterhouse, 1890 | | | Gymnopleurus flagellatus serratus Fischer von ..., 1821 | | | Gymnopleurus gemmatus Harold, 1871 | | | Gymnopleurus geoffroyi Fuessly, 1775 | | | Gymnopleurus geoffroyi cyanescens Motschulsky, 1849 | | | Gymnopleurus geoffroyi geoffroyi | | | Gymnopleurus geoffroyi virescens Müller, 1902 | | | Gymnopleurus humanus MacLeay, 1821 | | | Gymnopleurus humeralis Klug, 1855 | | | Gymnopleurus hypocrita Balthasar, 1960 | | | Gymnopleurus ignitus Klug, 1855 | | | Gymnopleurus koenigi Fabricius, 1775 | | | Gymnopleurus koenigii Fabricius, 1775 | Tranquebarische Königskäfer | | Gymnopleurus lacunosus Klug, 1845 | | | Gymnopleurus leei Fabricius, 1792 | | | Gymnopleurus miliaris Fabricius, 1775 | | | Gymnopleurus mimus (Balthasar, 1934) | ⇒ Gymnopleurus geoffroyi Fuessly, 1775 | | Gymnopleurus mopsus Pallas, 1781 | Breiter Pillendreher | | Gymnopleurus mopsus mopsus | | | Gymnopleurus mopsus sinensis Balthasar, 1934 | | | Gymnopleurus naviauxi Montreuil, 2009 | | | Gymnopleurus parvus MacLeay, 1821 | | | Gymnopleurus persianus (Reitter, 1909) | ⇒ Gymnopleurus mopsus Pallas, 1781 | | Gymnopleurus puncticollis (Gillet, 1909) | ⇒ Gymnopleurus virens Erichson, 1843 | | Gymnopleurus qurosh Montreuil, 2011 | | | Gymnopleurus rotundatus ♰ O. Heer, 1862 | | | Gymnopleurus sericeifrons Fairmaire, 1887 | | | Gymnopleurus serratus (Fischer von ..., 1821) | ⇒ Gymnopleurus flagellatus Fabricius, 1787 | | Gymnopleurus sindensis Arrow, 1931 | | | Gymnopleurus sinuatus (Geoffroy, 1785) | ⇒ Gymnopleurus geoffroyi Fuessly, 1775 | | Gymnopleurus sisyphus ♰ O. Heer, 1847 | | | Gymnopleurus sturmi (MacLeay, 1821) | ⇒ Gymnopleurus sturmii MacLeay, 1821 | | Gymnopleurus sturmii MacLeay, 1821 | | | Gymnopleurus tuxeni Petrovitz, 1955 | | | Gymnopleurus virens Erichson, 1843 | | | Haroldius annandalai Silvestri, 1924 | | | Haroldius annandalei Silvestri, 1924 | | | Haroldius borneensis Paulian, 1993 | | | Haroldius brendelli Krikken & Hu..., 2009 | | | Haroldius cambeforti Krikken & Hu..., 2006 | | | Haroldius cardoni Boucomont, 1923 | | | Haroldius celebensis Krikken & Hu..., 2006 | | | Haroldius discoidalis Paulian, 1993 | | | Haroldius fairmairei Boucomont, 1914 | | | Haroldius fleutiauxi Paulian, 1945 | | | Haroldius globosus Boucomont, 1925 | | | Haroldius heimi Wasmann, 1918 | | | Haroldius herrenorum Paulian, 1985 | | | Haroldius hwangi Masumoto, Le..., 2005 | | | Haroldius jechaer Král & Hájek, 2012 | | | Haroldius kawadai Masumoto, 1995 | | | Haroldius kolaka Krikken & Hu..., 2006 | | | Haroldius krali Utsunomiya &..., 2000 | | | Haroldius lassallei Cambefort, 1986 | | | Haroldius loebli Paulian, 1987 | | | Haroldius maruyamai Utsunomiya &..., 2005 | | | Haroldius oharai Ochi, Kon & ..., 2009 | | | Haroldius pahangensis Kral, 2003 | | | Haroldius pauliani Scheuern, 1995 | | | Haroldius penelopae Krikken & Hu..., 2006 | | | Haroldius perroti Paulian, 1939 | | | Haroldius philippinensis Pereira, 1954 | | | Haroldius rugatulus Boucomont, 1914 | | | Haroldius stevensi Arrow, 1931 | | | Haroldius sumatranus Paulian & Sc..., 1994 | | | Haroldius tangkoko Krikken & Hu..., 2006 | | | Haroldius thailandensis Paulian & Sc..., 1994 | | | Haroldius turnai Kral, 2003 | | | Haroldius uenoi Masumoto & Yin, 1993 | | | Helictopleurus giganteus Harold, 1869 | | | Helictopleurus neoamplicollis Krell, 2000 | | | Heliocopris andersoni Bates, 1868 | | | Heliocopris antiquus ♰ I. Fujiyama, 1968 | | | Heliocopris ares Moretto & Mi..., 2022 | | | Heliocopris atropos Boheman, 1860 | | | Heliocopris bucephalus Fabricius, 1775 | Indianische Ochsenkopf | | Heliocopris dominus Bates, 1868 | | | Heliocopris erycoides Felsche, 1907 | | | Heliocopris faunus Boheman, 1857 | | | Heliocopris gigas Linnaeus, 1758 | Ägyptische Riese | | Heliocopris gigas gigas | | | Heliocopris hamadryas Fabricius, 1775 | | | Heliocopris hermes Gillet, 1911 | | | Heliocopris isidis (Boheman, 1857) | ⇒ Heliocopris andersoni Bates, 1868 | | Heliocopris japetus Klug, 1855 | | | Heliocopris marshalli Péringuey, 1901 | | | Heliocopris midas Fabricius, 1775 | | | Heliocopris neptunus Boheman, 1857 | | | Heliocopris pauliani Janssens, 1939 | | | Heliocopris pirmal Fabricius, 1798 | | | Heliocopris staudingeri Kolbe, 1893 | | | Heliocopris tyrannus Thomson, 1859 | | | Heteronitis castelnaui Harold, 1862 | | | Hyalonthophagus alcedo Orbigny, 1913 | | | Hyalonthophagus alcyon Klug, 1855 | | | Isocopris imitator Felsche, 1901 | | | Ixodina abyssinica Roth, 1851 | | | Ixodina runica Arrow, 1909 | | | Kheper aegyptiorum Latreille, 1823 | | | Kheper aegyptiorum aegyptiorum | | | Kheper bonellii MacLeay, 1821 | | | Kheper devotus Redtenbacher, 1848 | | | Kheper erichsoni Harold, 1867 | | | Kheper lamarcki MacLeay, 1821 | | | Kheper nigroaeneus Boheman, 1857 | | | Kheper prodigiosus Erichson, 1843 | | | Kheper sanctus Fabricius, 1798 | | | Labroma umbratilis Matthews, 1974 | | | Larhodius calcaratus Janssens, 1934 | | | Larhodius hashimi Masumoto & U..., 2003 | | | Larhodius maruyamai Masumoto & U..., 2003 | | | Latodrepanus caelatus Gerstaecker, 1871 | | | Latodrepanus laticollis Fahraeus, 1857 | | | Lepanus australis Matthews, 1974 | | | Lepanus furcifer Matthews, 1974 | | | Lepanus globulus MacLeay, 1887 | | | Lepanus latheticus Matthews, 1974 | | | Lepanus monteithi Matthews, 1974 | | | Lepanus occidentalis Matthews, 1974 | | | Lepanus pygmaeus MacLeay, 1888 | | | Lepanus tozerensis Gunter & Weir, 2019 | | | Lepanus ustulatus Lansberge, 1874 | | | Lepanus villosus Matthews, 1974 | | | Liatongus affinis Arrow, 1908 | | | Liatongus ancorifer Kral & Rejsek, 1999 | | | Liatongus appositicornis Kral & Rejsek, 1999 | | | Liatongus bucerus Fairmaire, 1891 | | | Liatongus capricornis Arrow, 1931 | | | Liatongus clypeocornis Scheuern, 1988 | | | Liatongus davidi Boucomont, 1919 | | | Liatongus denticornis (Fairmaire, 1887) | ⇒ Liatongus bucerus Fairmaire, 1891 | | Liatongus endroedii Balthasar, 1956 | | | Liatongus fairmairei Boucomont, 1921 | | | Liatongus femoratus Illiger, 1800 | | | Liatongus gagatinus Hope, 1831 | | | Liatongus hastatus Kral & Rejsek, 1999 | | | Liatongus imitator (Balthasar, 1938) | ⇒ Liatongus medius Fairmaire, 1889 | | Liatongus incurvicornis Fairmaire, 1887 | | | Liatongus indicus Arrow, 1908 | | | Liatongus martialis Harold, 1879 | | | Liatongus medius Fairmaire, 1889 | | | Liatongus mergacerus Hope, 1831 | | | Liatongus militaris Castelnau, 1840 | | | Liatongus minutus Motschulsky, 1860 | | | Liatongus phanaeoides Westwood, 1839 | | | Liatongus phanaeoides fumiroi (Yohena, 1937) | ⇒ Liatongus minutus Motschulsky, 1860 | | Liatongus pugionatus Boheman, 1858 | | | Liatongus rhadamistus Fabricius, 1775 | | | Liatongus rhinoceros Arrow, 1931 | | | Liatongus spathulatus Roth, 1851 | | | Liatongus triacanthus Boucomont, 1920 | | | Liatongus tridentatus Boucomont, 1919 | | | Liatongus venator Fabricius, 1801 | | | Liatongus vertagus Fabricius, 1798 | | | Liatongus vseteckai (Balthasar, 1933) | ⇒ Liatongus fairmairei Boucomont, 1921 | | Lithanomala crassa ♰ A. G. Ponoma..., 1990 | | | Lithanomala oblonga ♰ A. G. Ponoma..., 1990 | | | Lithanomala oblongus Ponomarenko, 1990 | | | Lithanomala sibirica ♰ A. G. Ponoma..., 1990 | | | Litocopris simplex Harold, 1869 | | | Lobateuchus parisii ♰ O. Montreuil, 2010 | | | Macroderes gifboomi Abdalla & De..., 2018 | | | Macroderes nitidus Harold, 1877 | | | Macroderes politulus (Preudhomme d..., 1880) | ⇒ Macroderes bias Olivier, 1789 | | Macroderes tortuosus Abdalla & De..., 2018 | | | Macroderes undulatus Preudhomme d..., 1880 | | | Macropanelus fujitai Ochi, Kon & ..., 2008 | | | Macropanelus karasuyamai Ochi, Kon & ..., 2008 | | | Macropanelus sumatrensis Ochi & Kon &..., 1998 | | | Melanocanthon granulifer Schmidt, 1920 | | | Melanocanthon nigricornis Say, 1823 | | | Metacatharsius dentinum Ferreira, 1964 | | | Metacatharsius exiguiformis Ferreira, 1964 | | | Metacatharsius exiguus Boheman, 1860 | | | Metacatharsius ferreirae Balthasar, 1965 | | | Metacatharsius inermis Castelnau, 1840 | | | Metacatharsius marani Balthasar, 1940 | | | Metacatharsius opacus Waterhouse, 1891 | | | Metacatharsius pseudoopacus Ferreira, 1965 | | | Metacatharsius pumilionis Wallengren, 1881 | | | Metacatharsius pusio Kolbe, 1908 | | | Metacatharsius troglodytes (Boheman, 1857) | ⇒ Metacatharsius anderseni Waterhouse, 1891 | | Milichus apicalis (Fahraeus, 1857) | ⇒ Milichus rhodesianus Péringuey, 1904 | | Milichus picticollis Gerstaecker, 1871 | | | Mimonthophagus ambiguus Péringuey, 1901 | | | Mimonthophagus anomalus Klug, 1855 | | | Mnematidium multidentatum (Klug, 1845) | ⇒ Scarabaeus multidentatum Klug, 1845 | | Mnematidium multidentatus Klug, 1845 | | | Mnematium silenus Gray, 1832 | | | Monoplistes leai Paulian, 1934 | | | Monoplistes occidentalis MacLeay, 1888 | | | Neosisyphus barbarossa Wiedemann, 1823 | | | Neosisyphus bowringi White, 1844 | | | Neosisyphus calcaratus Klug, 1855 | | | Neosisyphus fortuitus Péringuey, 1901 | | | Neosisyphus kuehni Haaf, 1955 | | | Neosisyphus rubrus Paschalidis, 1974 | | | Neosisyphus spinipes Thunberg, 1818 | | | Neosisyphus tarantula Arrow, 1909 | | | Neosisyphus tibialis Raffray, 1877 | | | Ochicanthon besucheti Cuccodoro, 2011 | | | Ochicanthon cambeforti Ochi & Kon &..., 1997 | | | Ochicanthon ceylonicus Cuccodoro, 2011 | | | Ochicanthon cingalensis Arrow, 1931 | | | Ochicanthon crockermontis Krikken & Hu..., 2007 | | | Ochicanthon crypticus Krikken & Hu..., 2007 | | | Ochicanthon danum Krikken & Hu..., 2007 | | | Ochicanthon deplanatus Paulian, 1983 | | | Ochicanthon devagiriensis Sabu & Latha, 2011 | | | Ochicanthon dulitmontis Krikken & Hu..., 2007 | | | Ochicanthon dytiscoides Boucomont, 1914 | | | Ochicanthon edmondsi Krikken & Hu..., 2007 | | | Ochicanthon ernei Cuccodoro, 2011 | | | Ochicanthon fernandoi Sabu & Latha, 2011 | | | Ochicanthon gangkui Ochi & Kon &..., 1997 | | | Ochicanthon gauricola Cuccodoro, 2011 | | | Ochicanthon hanskii Krikken & Hu..., 2007 | | | Ochicanthon hikidai Ochi & Kon &..., 1997 | | | Ochicanthon javanus Krikken & Hu..., 2007 | | | Ochicanthon karasuyamai Ochi, Kon & ..., 2007 | | | Ochicanthon kikutai Ochi, Ueda &..., 2006 | | | Ochicanthon kimanis Krikken & Hu..., 2007 | | | Ochicanthon laetus Arrow, 1931 | | | Ochicanthon loebli Paulian, 1980 | | | Ochicanthon maryatiae Ochi, Ueda &..., 2006 | | | Ochicanthon masumotoi Ochi & Araya, 1996 | | | Ochicanthon mulu Krikken & Hu..., 2007 | | | Ochicanthon murthyi Vinod & Sabu, 2011 | | | Ochicanthon mussardi Cuccodoro, 2011 | | | Ochicanthon neglectus Krikken & Hu..., 2007 | | | Ochicanthon niinoii Ochi & Kon, 2014 | | | Ochicanthon niisatoi Ochi, Kon & ..., 2017 | | | Ochicanthon nitidus Paulian, 1980 | | | Ochicanthon obscurus Boucomont, 1920 | | | Ochicanthon ochii Hanboonsong ..., 2001 | | | Ochicanthon oharai Ochi, Kon, H..., 2008 | | | Ochicanthon okudai Ochi & Kon, 2014 | | | Ochicanthon parantisae Ochi & Kon &..., 1997 | | | Ochicanthon peninsularis Krikken & Hu..., 2007 | | | Ochicanthon philippinense Ochi, 1990 | | | Ochicanthon punctatus Boucomont, 1914 | | | Ochicanthon ranongensis Ochi, Kon & ..., 2020 | | | Ochicanthon rombauti Krikken & Hu..., 2007 | | | Ochicanthon takakui Ochi, Kon, H..., 2008 | | | Ochicanthon takuapaensis Ochi, Kon & ..., 2020 | | | Ochicanthon tambunan Krikken & Hu..., 2007 | | | Ochicanthon thai Paulian, 1987 | | | Ochicanthon thailandicum Masumoto, 1989 | | | Ochicanthon tristis Arrow, 1931 | | | Ochicanthon tristoides Paulian, 1983 | | | Ochicanthon uedai Ochi, Kon, H..., 2008 | | | Ochicanthon vazdemelloi (Latha & Sabu, 2011) | ⇒ Ochicanthon fernandoi Sabu & Latha, 2011 | | Ochicanthon woroae Ochi, Ueda &..., 2006 | | | Odontoloma dentinum Harold, 1868 | | | Odontoloma endroedyi Howden & Sch..., 1987 | | | Odontoloma louwi Howden & Sch..., 1987 | | | Odontoloma pusillum Howden & Sch..., 1987 | | | Odontoloma sculpturatum Harold, 1868 | | | Oniticellus (Paroniticellus) festivus Steven, 1809 | | | Oniticellus amplicollis ♰ (O. Heer, 1862) | ⇒ Helictopleurus neoamplicollis Krell, 2000 | | Oniticellus caelatus (Gerstaecker, 1871) | ⇒ Latodrepanus caelatus Gerstaecker, 1871 | | Oniticellus cinctus Fabricius, 1775 | Chinesische Gürtelkäfer | | Oniticellus egregius Klug, 1855 | | | Oniticellus falsus (Sharp, 1875) | ⇒ Sinodrepanus falsus Sharp, 1875 | | Oniticellus formosus (Chevrolat, 1830) | ⇒ Oniticellus pictus Hausmann, 1807 | | Oniticellus freyi Balthasar, 1941 | | | Oniticellus fulvus (Goeze, 1777) | ⇒ Euoniticellus fulvus Goeze, 1777 | | Oniticellus gayeni Biswas & Cha..., 1985 | | | Oniticellus namdaphaensis Biswas & Cha..., 1985 | | | Oniticellus pallens (Castelnau, 1840) | ⇒ Euoniticellus intermedius Reiche, 1848 | | Oniticellus pallipes (Fairmaire, 1887) | ⇒ Euoniticellus inaequalis Reiche, 1850 | | Oniticellus pictus Hausmann, 1807 | | | Oniticellus planatus Castelnau, 1840 | | | Oniticellus puberulus Y.-W. Zhang, 1988 | | | Oniticellus rhadamistus Fabricius, 1775 | Kleine Tranquebarische breitgrubichte Hornkäfer | | Oniticellus spinipes (Roth, 1851) | ⇒ Tiniocellus spinipes Roth, 1851 | | Oniticellus subhendui Biswas & Cha..., 1985 | | | Oniticellus tessellatus Harold, 1879 | | | Oniticellus yohenai (Matsumura, 1934) | ⇒ Liatongus phanaeoides Westwood, 1839 | | Onitis aeruginosus Klug, 1855 | | | Onitis alexis (Klug, 1835) | ⇒ Cheironitis ungaricus ungaricus Herbst, 1789 | | Onitis alexis alexis | | | Onitis alexis septentrionalis Balthasar, 1942 | | | Onitis anthracinus Felsche, 1907 | | | Onitis assamensis Biswas, 1980 | | | Onitis aygulus Fabricius, 1781 | | | Onitis belial Fabricius, 1798 | | | Onitis bordati Cambefort, 1988 | | | Onitis brahma Lansberge, 1875 | | | Onitis brevidens Bai, Yang & ..., 2006 | | | Onitis caffer Boheman, 1857 | | | Onitis castaneus Redtenbacher, 1848 | | | Onitis confusus Boheman, 1860 | | | Onitis corydon Boisduval, 1835 | | | Onitis crassus Sharp, 1875 | | | Onitis crenatus Reiche, 1847 | | | Onitis cryptodus Gillet, 1918 | | | Onitis curvipes Lansberge, 1875 | | | Onitis damoetas Steven, 1806 | | | Onitis deceptor Péringuey, 1901 | | | Onitis excavatus Arrow, 1931 | | | Onitis ezechias Reiche, 1856 | | | Onitis falcatus Wulfen, 1786 | | | Onitis feae Felsche, 1907 | | | Onitis fulgidus (Klug, 1855) | ⇒ Pleuronitis fulgidus Castelnau, 1840 | | Onitis hageni Lansberge, 1886 | | | Onitis humerosus Pallas, 1771 | | | Onitis intermedius Frivaldsky, 1892 | | | Onitis ion Olivier, 1789 | | | Onitis lama Lansberge, 1875 | | | Onitis lansbergei Sharp, 1875 | | | Onitis ludekingi Lansberge, 1875 | | | Onitis magus ♰ C. von Heyden, 1862 | | | Onitis mendax Gillet, 1918 | | | Onitis minutus Lansberge, 1875 | | | Onitis naviauxi Cambefort, 1988 | | | Onitis niger (Lansberge, 1875) | ⇒ Liatongus femoratus Illiger, 1800 | | Onitis numida Castelnau, 1840 | | | Onitis numida marginicollis Balthasar, 1928 | | | Onitis obscurus Lansberge, 1886 | | | Onitis pecuarius Lansberge, 1875 | | | Onitis pers Gusakov & Kl..., 2009 | | | Onitis phartopus Lansberge, 1875 | | | Onitis philemon (Fabricius, 1801) | ⇒ Onitis subopacus Arrow, 1931 | | Onitis picticollis Boheman, 1857 | | | Onitis pseudosetosus Ferreira, 1976 | | | Onitis punctatostriatus Janssens, 1937 | | | Onitis reichei Lansberge, 1875 | | | Onitis singhalensis Lansberge, 1875 | | | Onitis siva Gillet, 1911 | | | Onitis sphinx Fabricius, 1775 | | | Onitis spinipes Drury, 1770 | | | Onitis subcrenatus Kolbe, 1897 | | | Onitis subopacus Arrow, 1931 | | | Onitis tingaudi Keith & Moretto, 2007 | | | Onitis tortuosus Houston, 1983 | | | Onitis tumidus Gillet, 1918 | | | Onitis uncinatus Klug, 1855 | | | Onitis vanderkelleni Lansberge, 1886 | | | Onitis virens Lansberge, 1875 | | | Onitis viridulus (Boheman, 1857) | ⇒ Onitis pecuarius Lansberge, 1875 | | Onitis westermanni Lansberge, 1886 | | | Ontherus brevicollis Kirsch, 1870 | | | Ontherus carinifrons Luederwaldt, 1930 | | | Ontherus cephalotes Harold, 1869 | | | Ontherus pseudodidymus Génier, 1996 | | | Ontherus pubens Génier, 1996 | | | Ontherus sanctaemartae Génier, 1996 | | | Onthophagus (Afrostrandius) plebejus Klug, 1855 | | | Onthophagus (Altonthophagus) cervenkai Kabakov, 2008 | | | Onthophagus (Altonthophagus) inelegans Balthasar, 1935 | | | Onthophagus (Altonthophagus) kozlovi Kabakov, 1990 | | | Onthophagus (Altonthophagus) kukunorensis Kabakov, 1990 | | | Onthophagus (Altonthophagus) marmotae Kabakov, 1990 | | | Onthophagus (Altonthophagus) sibiricus Harold, 1877 | | | Onthophagus (Altonthophagus) turpidoides Kabakov, 2008 | | | Onthophagus (Amphionthophagus) falzonii Goidanich, 1926 | | | Onthophagus (Amphionthophagus) melitaeus Fabricius, 1798 | | | Onthophagus (Colobonthophagus) aenescens Wiedemann, 1823 | | | Onthophagus (Colobonthophagus) agnus Gillet, 1925 | | | Onthophagus (Colobonthophagus) arunensis Scheuern, 1995 | | | Onthophagus (Colobonthophagus) caprai Frey, 1956 | | | Onthophagus (Colobonthophagus) dama Fabricius, 1798 | | | Onthophagus (Colobonthophagus) metalliceps Arrow, 1931 | | | Onthophagus (Colobonthophagus) neocolobus Scheuern, 1996 | | | Onthophagus (Colobonthophagus) occipitalis Lansberge, 1885 | | | Onthophagus (Colobonthophagus) paliceps Arrow, 1931 | | | Onthophagus (Colobonthophagus) piceorufulus Kabakov, 1994 | | | Onthophagus (Colobonthophagus) poggii Scheuern, 1996 | | | Onthophagus (Colobonthophagus) quadridentatus (Fabricius, 1798) | ⇒ Onthophagus fissicornis Stevens, 1809 | | Onthophagus (Colobonthophagus) shillongensis Scheuern, 1995 | | | Onthophagus (Colobonthophagus) thai Kabakov, 1994 | | | Onthophagus (Colobonthophagus) tragus Fabricius, 1792 | | | Onthophagus (Colobonthophagus) transquadridentatus Scheuern, 1995 | | | Onthophagus (Digitonthophagus) bonasus Fabricius, 1775 | | | Onthophagus (Digitonthophagus) gazella (Fabricius, 1787) | ⇒ Digitonthophagus gazella Fabricius, 1787 | | Onthophagus (Euonthophagus) amyntas (Olivier, 1789) | ⇒ Euonthophagus amyntas Olivier, 1789 | | Onthophagus (Euonthophagus) atramentarius Ménétriés, 1832 | | | Onthophagus (Euonthophagus) carbonarius Klug, 1855 | | | Onthophagus (Furconthophagus) amicus Gillet, 1925 | | | Onthophagus (Furconthophagus) boucomonti Paulian, 1931 | | | Onthophagus (Furconthophagus) furcatus (Fabricius, 1781) | ⇒ Onthophagus furcaticeps Masters, 1886 | | Onthophagus (Furconthophagus) karenensis Masumoto, Oc..., 2008 | | | Onthophagus (Furconthophagus) khonkaenus Masumoto, Oc..., 2008 | | | Onthophagus (Furconthophagus) lilliputanus Lansberge, 1883 | | | Onthophagus (Furconthophagus) papulatorius Kabakov, 2008 | | | Onthophagus (Furconthophagus) papulatus Boucomont, 1914 | | | Onthophagus (Furconthophagus) promontorii Boucomont, 1924 | | | Onthophagus (Furconthophagus) schoolmeestersi Ochi & Kon, 2007 | | | Onthophagus (Gibbonthophagus) apicetinctus (d´Orbigny, 1900) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus (Gibbonthophagus) atripennis Waterhouse, 1875 | | | Onthophagus (Gibbonthophagus) chineicus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) denticollis Lansberge, 1883 | | | Onthophagus (Gibbonthophagus) duporti Boucomont, 1914 | | | Onthophagus (Gibbonthophagus) euryceros Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) fujiii Ochi & Kon, 1995 | | | Onthophagus (Gibbonthophagus) gracilipes Boucomont, 1914 | | | Onthophagus (Gibbonthophagus) hiabunicus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) kentingensis Nomura, 1973 | | | Onthophagus (Gibbonthophagus) kimioi Ochi, Kon & ..., 2011 | | | Onthophagus (Gibbonthophagus) lenis Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) limbatus Herbst, 1789 | | | Onthophagus (Gibbonthophagus) luridipennis Boheman, 1858 | | | Onthophagus (Gibbonthophagus) nigriobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) palawanicus Ochi & Kon, 2004 | | | Onthophagus (Gibbonthophagus) parviobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) penmani Masumoto, Oc..., 2002 | | | Onthophagus (Gibbonthophagus) privus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) rufiobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) sasajii Ochi & Kon, 2001 | | | Onthophagus (Gibbonthophagus) semipersonatus Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) sunantaae Masumoto, 1989 | | | Onthophagus (Gibbonthophagus) taurinus White, 1844 | | | Onthophagus (Gibbonthophagus) viduus (Harold, 1875) | ⇒ Onthophagus orbus Boucomont, 1924 | | Onthophagus (Gibbonthophagus) viridicervicapra Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) arai Masumoto, 1989 | | | Onthophagus (Indachorius) baykanus Kabakov, 1994 | | | Onthophagus (Indachorius) bonengus Kabakov, 1994 | | | Onthophagus (Indachorius) chinensis Balthasar, 1952 | | | Onthophagus (Indachorius) danumensis (Ochi, Kon & ..., 2009) | ⇒ Onthophagus danumicus Huijbregts &..., 2012 | | Onthophagus (Indachorius) doisuthepensis Masumoto, 1989 | | | Onthophagus (Indachorius) hsui Masumoto, Ch..., 2004 | | | Onthophagus (Indachorius) koshunensis Balthasar, 1941 | | | Onthophagus (Indachorius) lannamiibun Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) maesaensis Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) masaoi Ochi, 1992 | | | Onthophagus (Indachorius) mendeli Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) mongkhoni Masumoto, Ha..., 2002 | | | Onthophagus (Indachorius) nongkaiensis Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) octonaevus Kabakov, 1994 | | | Onthophagus (Indachorius) paramasaoi Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) phetchabunensis Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) phuquoci Paulian, 1945 | | | Onthophagus (Indachorius) platypus Zhang, 1997 | | | Onthophagus (Indachorius) pseudoworoae Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) scotti Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) semidanumensis Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) subsapaensis Kabakov, 1994 | | | Onthophagus (Indachorius) suginoi Ochi, 1984 | | | Onthophagus (Indachorius) tiamicus Kabakov, 1994 | | | Onthophagus (Indachorius) tongbantumi Masumoto, Oc..., 2002 | | | Onthophagus (Macronthophagus) cludtsi Ochi, 2003 | | | Onthophagus (Macronthophagus) curvicarinatus Boucomont, 1914 | | | Onthophagus (Macronthophagus) diabolicus Harold, 1877 | | | Onthophagus (Macronthophagus) manipurensis Arrow, 1907 | | | Onthophagus (Macronthophagus) menieri Ochi, 2003 | | | Onthophagus (Macronthophagus) nilgirensis Gillet, 1922 | | | Onthophagus (Macronthophagus) rubricollis Hope, 1831 | | | Onthophagus (Macronthophagus) uenoi Ochi, 1995 | | | Onthophagus uenoi borneomontanus Ochi, 2003 | | | Onthophagus uenoi uenoi Ochi, 1995 | | | Onthophagus (Matashia) anguliceps (Boucomont, 1914) | ⇒ Onthophagus avocetta Arrow, 1933 | | Onthophagus (Matashia) yubarinus (Matsumura, 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus (Micronthophagus) falsivigilans Masumoto, 1995 | | | Onthophagus (Micronthophagus) gigantivigilans Masumoto, Ha..., 2002 | | | Onthophagus (Micronthophagus) konoi Matsumura, 1938 | | | Onthophagus (Micronthophagus) quaestus Sharp, 1875 | | | Onthophagus (Micronthophagus) vigilans Boucomont, 1921 | | | Onthophagus (Micronthophagus) wangi Masumoto, Ch..., 2004 | | | Onthophagus (Onthophagiellus) kondaoensis Kabakov, 1994 | | | Onthophagus (Onthophagiellus) opacifalculatus Ochi, Kon & ..., 2009 | | | Onthophagus (Onthophagiellus) parafalculatus Ochi, Kon & ..., 2009 | | | Onthophagus (Onthophagiellus) popovi Kabakov, 1983 | | | Onthophagus (Onthophagiellus) taiyaruensis Masumoto, 1977 | | | Onthophagus (Onthophagiellus) thanwaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagiellus) unguiculatus Kabakov, 1994 | | | Onthophagus (Onthophagus) abacus Boucomont, 1921 | | | Onthophagus (Onthophagus) acuticollis (Gillet, 1927) | ⇒ Onthophagus amphioxus Arrow, 1933 | | Onthophagus (Onthophagus) acuticornis Endrödi, 1973 | | | Onthophagus (Onthophagus) afghanus Petrovitz, 1961 | | | Onthophagus (Onthophagus) amirus Kabakov, 1982 | | | Onthophagus (Onthophagus) anguicorius Boucomont, 1914 | | | Onthophagus (Onthophagus) angustatus Boucomont, 1914 | | | Onthophagus (Onthophagus) apilularius Masumoto, 1995 | | | Onthophagus (Onthophagus) arnoldii Kabakov, 1982 | | | Onthophagus (Onthophagus) barbieri Paulian, 1978 | | | Onthophagus (Onthophagus) batesi Howden and C..., 1963 | | | Onthophagus (Onthophagus) bokiaunus Masumoto, 1995 | | | Onthophagus (Onthophagus) brevifrons (Horn, 1881) | ⇒ Onthophagus abyssinicus Gillet, 1925 | | Onthophagus (Onthophagus) chaiyaphumensis Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) dapcauensis Boucomont, 1921 | | | Onthophagus (Onthophagus) depressus Harold, 1871 | | | Onthophagus (Onthophagus) doiinthanonensis Masumoto, 1989 | | | Onthophagus (Onthophagus) doipuiensis Masumoto, 1989 | | | Onthophagus (Onthophagus) doitungensis Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) dubernardi Boucomont, 1914 | | | Onthophagus (Onthophagus) embersoni Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) fissicornis Stevens, 1809 | | | Onthophagus (Onthophagus) formosanus Gillet, 1924 | | | Onthophagus (Onthophagus) gibbulus Pallas, 1781 | | | Onthophagus (Onthophagus) gorochovi Kabakov, 1998 | | | Onthophagus (Onthophagus) gosoli Masumoto, 1989 | | | Onthophagus (Onthophagus) grandivigilans Masumoto, 1995 | | | Onthophagus (Onthophagus) hajimei Masumoto, 1984 | | | Onthophagus (Onthophagus) hastifer Lansberge, 1885 | | | Onthophagus (Onthophagus) hayashii Masumoto, 1991 | | | Onthophagus (Onthophagus) hecate Panzer, 1794 | | | Onthophagus (Onthophagus) horii Ochi, Kon & ..., 2009 | | | Onthophagus (Onthophagus) illyricus Scopoli, 1763 | | | Onthophagus (Onthophagus) incensus Say, 1835 | | | Onthophagus (Onthophagus) ishiii Ochi & Kon, 1995 | | | Onthophagus (Onthophagus) javaecola Balthasar, 1959 | | | Onthophagus (Onthophagus) kanyaayonus Masumoto, 1992 | | | Onthophagus (Onthophagus) khonmiinitnoi Masumoto, 1990 | | | Onthophagus (Onthophagus) kinhthaicus Kabakov, 1998 | | | Onthophagus (Onthophagus) kiuchii Masumoto, 1995 | | | Onthophagus (Onthophagus) knausi Brown, 1927 | | | Onthophagus (Onthophagus) koryoensis Kim, 1985 | | | Onthophagus (Onthophagus) krakadaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagus) kuatunensis Balthasar, 1942 | | | Onthophagus (Onthophagus) lakyim Masumoto, 1990 | | | Onthophagus (Onthophagus) landolti (Harold, 1880) | ⇒ Onthophagus schaefferi Howden and C..., 1963 | | Onthophagus (Onthophagus) lindaae Masumoto, 1989 | | | Onthophagus (Onthophagus) lutosopictus Fairmaire, 1897 | | | Onthophagus (Onthophagus) maleengnaafon Masumoto, 1990 | | | Onthophagus (Onthophagus) maleengnoi Masumoto, 1990 | | | Onthophagus (Onthophagus) maniti Masumoto, 1989 | | | Onthophagus (Onthophagus) margaritifer d´Orbigny, 1898 | | | Onthophagus (Onthophagus) masumotoi Ochi, 1985 | | | Onthophagus (Onthophagus) matsudai Ochi & Kon, 2004 | | | Onthophagus (Onthophagus) medorensis Brown, 1929 | | | Onthophagus (Onthophagus) mendicus Gillet, 1924 | | | Onthophagus (Onthophagus) mulleri van Lansberge, 1883 | | | Onthophagus (Onthophagus) naaroon Masumoto, 1990 | | | Onthophagus (Onthophagus) namnaoensis Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) nasonis Kabakov, 1998 | | | Onthophagus (Onthophagus) nuchicornis Linnaeus, 1758 | | | Onthophagus (Onthophagus) nurestanicus Kabakov, 1982 | | | Onthophagus (Onthophagus) oklahomensis Brown, 1927 | | | Onthophagus (Onthophagus) orientalis (Harold, 1868) | ⇒ Onthophagus fissinasus Fairmaire, 1895 | | Onthophagus (Onthophagus) orpheus Panzer, 1794 | | | Onthophagus (Onthophagus) pennsylvanicus Harold, 1871 | | | Onthophagus (Onthophagus) phraoensis Masumoto, 1990 | | | Onthophagus (Onthophagus) phrutsaphaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagus) polyphemi Hubbard, 1894 | | | Onthophagus (Onthophagus) proletarius Harold, 1875 | | | Onthophagus (Onthophagus) pseudoarmatus Balthasar, 1944 | | | Onthophagus (Onthophagus) pseudohystrix Masumoto, 1995 | | | Onthophagus (Onthophagus) punctator Reitter, 1893 | | | Onthophagus (Onthophagus) punneeae Masumoto, 1989 | | | Onthophagus (Onthophagus) ratchasimaensis Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) roubali Balthasar, 1935 | | | Onthophagus (Onthophagus) rufimanus Kabakov, 1982 | | | Onthophagus (Onthophagus) rugulosus Harold, 1886 | | | Onthophagus (Onthophagus) sakaeratensis Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) sakainoi Masumoto, 1991 | | | Onthophagus (Onthophagus) salvazai Paulian, 1978 | | | Onthophagus (Onthophagus) sauteri Gillet, 1924 | | | Onthophagus (Onthophagus) setchan Masumoto, 1984 | | | Onthophagus (Onthophagus) singhaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagus) sobrius Balthasar, 1959 | | | Onthophagus (Onthophagus) striatulus Palisot de B..., 1809 | | | Onthophagus (Onthophagus) subaeneus (Palisot de B..., 1811) | ⇒ Onthophagus striatulus Palisot de B..., 1809 | | Onthophagus (Onthophagus) subtropicus Howden and C..., 1963 | | | Onthophagus (Onthophagus) taayai Masumoto, 1995 | | | Onthophagus (Onthophagus) tatsienluensis Balthasar, 1942 | | | Onthophagus (Onthophagus) taurus Schreber, 1759 | Oechslein | | Onthophagus (Onthophagus) terminatus Eschscholtz, 1822 | | | Onthophagus (Onthophagus) tholaayi Masumoto, 1990 | | | Onthophagus (Onthophagus) tritinctus Boucomont, 1914 | | | Onthophagus (Onthophagus) trituber Wiedemann, 1823 | | | Onthophagus trituber jacobsoni Kabakov, 2006 | | | Onthophagus (Onthophagus) truncaticornis (Schaller, 1783) | ⇒ Onthophagus quadrinodosus Fahraeus, 1857 | | Onthophagus (Onthophagus) tuberculifrons Harold, 1871 | | | Onthophagus (Onthophagus) tumami Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) velutinus Horn, 1875 | | | Onthophagus (Onthophagus) vitulus Péringuey, 1901 | | | Onthophagus (Onthophagus) wangnamkhieoensis Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) yamaokai Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) yangi Masumoto, Ts..., 2006 | | | Onthophagus (Palaeonthophagus) afghanus Petrovitz, 1961 | | | Onthophagus (Palaeonthophagus) albarracinus Baraud, 1979 | | | Onthophagus (Palaeonthophagus) angorensis Petrovitz, 1963 | | | Onthophagus (Palaeonthophagus) baraudi Nicolas, 1964 | | | Onthophagus (Palaeonthophagus) bytinskii Balthasar, 1960 | | | Onthophagus (Palaeonthophagus) clitellifer Reitter, 1894 | | | Onthophagus (Palaeonthophagus) coenobita Herbst, 1783 | Mönchs-Kotkäfer | | Onthophagus (Palaeonthophagus) conspersus Reitter, 1893 | | | Onthophagus (Palaeonthophagus) cruciatus (Ménétriés, 1832) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus (Palaeonthophagus) dacatrai Pittino, 2004 | | | Onthophagus (Palaeonthophagus) dellacasai Pittino & Ma..., 1981 | | | Onthophagus (Palaeonthophagus) eulaminicornis Pittino, 2006 | | | Onthophagus (Palaeonthophagus) excisus Reiche & Saulcy, 1856 | | | Onthophagus (Palaeonthophagus) excubitor Ziani & Gudenzi, 2006 | | | Onthophagus (Palaeonthophagus) finschi Harold, 1877 | | | Onthophagus (Palaeonthophagus) fissicornis Steven, 1809 | | | Onthophagus (Palaeonthophagus) fortigibber Reitter, 1909 | | | Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790) | ⇒ Onthophagus similis Scriba, 1790 | | Onthophagus (Palaeonthophagus) furciceps Marseul, 1869 | | | Onthophagus (Palaeonthophagus) gibbulus Pallas, 1781 | | | Onthophagus gibbulus gibbulus | | | Onthophagus gibbulus rostrifer Reitter, 1892 | | | Onthophagus (Palaeonthophagus) gorodinskii Kabakov, 2008 | | | Onthophagus (Palaeonthophagus) grossepunctatus Reitter, 1905 | | | Onthophagus (Palaeonthophagus) haroldi Ballion, 1870 | | | Onthophagus (Palaeonthophagus) hissariensis Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) isikdagensis Pittino, 2004 | | | Onthophagus (Palaeonthophagus) joannae Goljan, 1953 | | | Onthophagus (Palaeonthophagus) kindermanni Harold, 1877 | | | Onthophagus (Palaeonthophagus) kolenatii Reitter, 1893 | | | Onthophagus (Palaeonthophagus) laticornis Gebler, 1823 | | | Onthophagus (Palaeonthophagus) latigena d´Orbigny, 1897 | | | Onthophagus (Palaeonthophagus) lemur Fabricius, 1781 | | | Onthophagus (Palaeonthophagus) leucostigma Steven, 1811 | | | Onthophagus (Palaeonthophagus) lucidus Illiger, 1800 | | | Onthophagus (Palaeonthophagus) macedonicus Miksic, 1959 | | | Onthophagus (Palaeonthophagus) marginalis Gebler, 1817 | | | Onthophagus marginalis marginalis | | | Onthophagus marginalis przewalski Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) massai Baraud, 1975 | | | Onthophagus (Palaeonthophagus) medius (Kugelann, 1792) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus (Palaeonthophagus) medvedevi Kabakov, 1982 | | | Onthophagus (Palaeonthophagus) merdarius Chevrolat, 1865 | | | Onthophagus (Palaeonthophagus) muelleri (Lansberge, 1883) | ⇒ Onthophagus statzi Krell, 1990 | | Onthophagus (Palaeonthophagus) nikolajevi Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) novaki Boucomont, 1927 | | | Onthophagus (Palaeonthophagus) nuchicornis Linnaeus, 1758 | Kleine Nackenhorn | | Onthophagus (Palaeonthophagus) olsoufieffi Boucomont, 1924 | | | Onthophagus (Palaeonthophagus) opacicollis Reitter, 1893 | | | Onthophagus (Palaeonthophagus) ovatus Linnaeus, 1767 | Eiförmiger Kotkäfer | | Onthophagus (Palaeonthophagus) panici Petrovitz, 1964 | | | Onthophagus (Palaeonthophagus) parmatus Reitter, 1892 | | | Onthophagus (Palaeonthophagus) petrovitzi (Frey, 1975) | ⇒ Onthophagus petrovitzianus Pittino, 2006 | | Onthophagus (Palaeonthophagus) petrovitzianus Pittino, 2006 | | | Onthophagus (Palaeonthophagus) ponticus Harold, 1883 | | | Onthophagus (Palaeonthophagus) pseudocaccobius Reitter, 1889 | | | Onthophagus (Palaeonthophagus) psychopompus Ziani & Ghar..., 2010 | | | Onthophagus (Palaeonthophagus) pygargus Motschulsky, 1845 | | | Onthophagus (Palaeonthophagus) quadrinodus Reitter, 1896 | | | Onthophagus (Palaeonthophagus) rachelis Martín Piera, 1985 | | | Onthophagus (Palaeonthophagus) rakovici Pittino, 2004 | | | Onthophagus (Palaeonthophagus) rodentium Pittino, 2004 | | | Onthophagus (Palaeonthophagus) ruficapillus Brullé, 1832 | | | Onthophagus ruficapillus guilanensis (Pittino, 1982) | ⇒ Onthophagus ruficapillus Brullé, 1832 | | Onthophagus ruficapillus ruficapillus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) samai Ziani, 2011 | | | Onthophagus (Palaeonthophagus) scabriusculus Harold, 1873 | | | Onthophagus (Palaeonthophagus) semicornis Panzer, 1798 | Halbhörniger Kotkäfer | | Onthophagus (Palaeonthophagus) sericatus Reitter, 1893 | | | Onthophagus (Palaeonthophagus) shapovalovi Gusakov, 2012 | | | Onthophagus (Palaeonthophagus) similis (Scriba, 1790) | ⇒ Onthophagus fracticornis Preyssler, 1790 | | Onthophagus (Palaeonthophagus) speculifer Solsky, 1876 | | | Onthophagus (Palaeonthophagus) strabo Reitter, 1892 | | | Onthophagus (Palaeonthophagus) stylocerus Graells, 1851 | | | Onthophagus (Palaeonthophagus) suturellus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) tesquorum Semenov & Me..., 1927 | | | Onthophagus (Palaeonthophagus) trigibber Reitter, 1893 | | | Onthophagus (Palaeonthophagus) trispinus Reitter, 1892 | | | Onthophagus (Palaeonthophagus) truchmenus Kolenati, 1846 | | | Onthophagus truchmenus iranicus Kabakov, 2006 | | | Onthophagus truchmenus truchmenus | | | Onthophagus truchmenus turkmeniacus Kabakov, 2006 | | | Onthophagus truchmenus xerxes Petrovitz, 1965 | | | Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767 | Kühlein | | Onthophagus (Palaeonthophagus) verticicornis (Laicharting, 1781) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus (Palaeonthophagus) viridis Ménétriés, 1832 | | | Onthophagus (Palaeonthophagus) vitulus (Fabricius, 1776) | ⇒ Onthophagus immundus Boheman, 1858 | | Onthophagus (Palaeonthophagus) vlasovi Medvedev, 1958 | | | Onthophagus (Palaeonthophagus) zuninoi Martín Piera, 1985 | | | Onthophagus (Paleonthophagus) andalusicus Waltl, 1835 | | | Onthophagus andalusicus andalusicus | | | Onthophagus andalusicus persianus Olsoufieff, 1900 | | | Onthophagus (Paleonthophagus) baraudi Nicolas, 1964 | | | Onthophagus (Paleonthophagus) coenobita Herbst, 1783 | | | Onthophagus (Paleonthophagus) fracticornis Preyssler, 1790 | | | Onthophagus (Paleonthophagus) gibbulus Pallas, 1781 | | | Onthophagus (Paleonthophagus) grossepunctatus Reitter, 1905 | | | Onthophagus (Paleonthophagus) joannae Goljan, 1953 | | | Onthophagus (Paleonthophagus) lemur Fabricius, 1781 | | | Onthophagus (Paleonthophagus) massai Baraud, 1975 | | | Onthophagus (Paleonthophagus) nuchicornis Linnaeus, 1758 | | | Onthophagus (Paleonthophagus) opacicollis Pierotti, 1959 | | | Onthophagus (Paleonthophagus) ovatus Linnaeus, 1767 | | | Onthophagus (Paleonthophagus) ruficapillus Brullé, 1832 | | | Onthophagus (Paleonthophagus) semicornis Panzer, 1798 | | | Onthophagus (Paleonthophagus) similis Scriba, 1790 | | | Onthophagus (Paleonthophagus) vacca Linnaeus, 1767 | | | Onthophagus (Paleonthophagus) verticicornis Laicharting, 1781 | | | Onthophagus (Paleonthophagus) vitulus Fabricius, 1776 | | | Onthophagus (Paraphanaeomorphus) argyropygus Gillet, 1927 | | | Onthophagus (Paraphanaeomorphus) carinensis Boucomont, 1914 | | | Onthophagus (Paraphanaeomorphus) comottoides Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) insignicollis Frey, 1954 | | | Onthophagus (Paraphanaeomorphus) janushevi Kabakov, 1983 | | | Onthophagus (Paraphanaeomorphus) napolovi Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) pseudojavanus Paulian, 1931 | | | Onthophagus (Paraphanaeomorphus) quadricolor Kabakov, 1983 | | | Onthophagus (Paraphanaeomorphus) semipiceus Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) strnadi Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) vaulogeri Boucomont, 1923 | | | Onthophagus (Parascatonomus) anceyi Boucomont, 1921 | | | Onthophagus (Parascatonomus) damaki Masumoto, Ha..., 2002 | | | Onthophagus (Parascatonomus) kiuchianus Masumoto, Ya..., 2004 | | | Onthophagus (Parascatonomus) koichii (Ochi & Araya, 1994) | ⇒ Caccobius jessoensis Harold, 1867 | | Onthophagus (Parascatonomus) leytensis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) miyakei Ochi & Araya, 1992 | | | Onthophagus (Parascatonomus) murasakianus Nomura, 1976 | | | Onthophagus (Parascatonomus) negrosensis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) politus Fabricius, 1798 | | | Onthophagus (Parascatonomus) pseudorudis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) quezonensis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) satoi Ochi, Shimad..., 2003 | | | Onthophagus (Parascatonomus) shizumui Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) takedai Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) takeshii Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) tricornis (Wiedemann, 1823) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus (Parascatonomus) yukae Masumoto, Oc..., 2002 | | | Onthophagus (Parentius) emarginatus Mulsant, 1842 | | | Onthophagus (Parentius) hispanicus Baraud, 1963 | | | Onthophagus (Parentius) kabakovi Martin-Piera, 1985 | | | Onthophagus (Parentius) punctatus Illiger, 1803 | | | Onthophagus (Paronthophagus) hidakai Ochi & Kon, 1995 | | | Onthophagus (Phalops) tricuspis (Semenov, 1900) | ⇒ Onthophagus fortigibber Reitter, 1909 | | Onthophagus (Phanaeomorphus) ater Waterhouse, 1875 | | | Onthophagus (Phanaeomorphus) cernyi (Balthasar, 1935) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus (Phanaeomorphus) johkii Ochi & Kon, 1994 | | | Onthophagus (Phanaeomorphus) lenzi Harold, 1875 | | | Onthophagus potanini hiurai Ochi, 1984 | | | Onthophagus (Phanaeomorphus) zetteli Masumoto, Oc..., 2002 | | | Onthophagus brucei cyanopterus (d´Orbigny, 1905) | ⇒ Proagoderus cyanopterus d´Orbigny, 1905 | | Onthophagus (Proagoderus) pegesimallus Marcus, 1917 | | | Onthophagus (Proagoderus) vividus Arrow, 1907 | | | Onthophagus (Pseudophanaeomorphus) parachandrai Ochi, Kon & ..., 2009 | | | Onthophagus (Relictonthophagus) emarginatus (Mulsant, 1842) | ⇒ Onthophagus rubrimaculatus MacLeay, 1864 | | Onthophagus (Relictonthophagus) nigellus (Illiger, 1803) | ⇒ Caccobius histeroides Ménétriés, 1832 | | Onthophagus (Relictonthophagus) punctatus (Illiger, 1803) | ⇒ Onthophagus emarginatus Mulsant, 1842 | | Onthophagus (Serrophorus) avocettoides Kabakov, 1983 | | | Onthophagus (Serrophorus) laevis (Harold, 1880) | ⇒ Onthophagus laevissimus d´Orbigny, 1905 | | Onthophagus laevis asiaticus Boucomont, 1919 | | | Onthophagus laevis laevis | | | Onthophagus laevis lampromelas Fairmaire, 1891 | | | Onthophagus laevis stevensi Arrow, 1931 | | | Onthophagus (Serrophorus) oblongus Zhang, 1997 | | | Onthophagus (Serrophorus) rectecornutus Lansberge, 1883 | | | Onthophagus (Serrophorus) sagittarius Fabricius, 1775 | | | Onthophagus (Sinonthophagus) nampatensis Tarasov & Ka..., 2010 | | | Onthophagus (Strandius) changshouensis Zhang, 1997 | | | Onthophagus (Strandius) japonicus Harold, 1875 | | | Onthophagus (Strandius) kuraruanus (Matsumura, 1938) | ⇒ Onthophagus lenzi Harold, 1875 | | Onthophagus (Strandius) obliquus Olivier, 1789 | | | Onthophagus (Strandius) potanini R. C. Brusca..., 1990 | | | Onthophagus potanini hiurai Ochi, 1984 | | | Onthophagus potanini potanini | | | Onthophagus (Strandius) subansiriensis Biswas, 1979 | | | Onthophagus (Strandius) taiwanus Nomura, 1973 | | | Onthophagus (Sunenaga) streltsovi Tarasov & Ka..., 2010 | | | Onthophagus (Trichonthophagus) dynastoides Arrow, 1931 | | | Onthophagus (Trichonthophagus) hirtus Illiger, 1803 | | | Onthophagus (Trichonthophagus) maki Illiger, 1803 | | | Onthophagus (Trichonthophagus) tarandus Fabricius, 1792 | | | Onthophagus abei Ochi & Kon, 2015 | | | Onthophagus abreui Arrow, 1931 | | | Onthophagus absyrtus Balthasar, 1946 | | | Onthophagus academus Balthasar, 1946 | | | Onthophagus aceroides Krikken & Hu..., 2013 | | | Onthophagus acrisius Balthasar, 1946 | | | Onthophagus aemulus Gillet, 1930 | | | Onthophagus aeneopiceus d´Orbigny, 1902 | | | Onthophagus aequiperus Matsumura & ..., 1937 | | | Onthophagus aerarius Reitter, 1892 | | | Onthophagus aereomaculatus Boucomont, 1914 | | | Onthophagus aereopictus Boucomont, 1914 | | | Onthophagus aereopictus aereopictus | | | Onthophagus aereopictus rubropictus Boucomont, 1924 | | | Onthophagus aeruginosus Roth, 1851 | | | Onthophagus aethiopicus d´Orbigny, 1902 | | | Onthophagus agaricophilus Arrow, 1931 | | | Onthophagus agilis (Matsumura, 1938) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus akhaus Masumoto, Oc..., 2007 | | | Onthophagus akinini Koenig, 1889 | | | Onthophagus akirai (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus albicornis Palisot de B..., 1805 | | | Onthophagus albipennis Péringuey, 1908 | | | Onthophagus aleppensis Redtenbacher, 1843 | | | Onthophagus aleppensis aleppensis | | | Onthophagus aleppensis hyrcanicus Kabakov, 1982 | | | Onthophagus alfuricus Huijbregts &..., 2012 | | | Onthophagus allojavanus Huijbregts &..., 2011 | | | Onthophagus aloysiellus Zunino, 1977 | | | Onthophagus amamiensis Nomura, 1965 | | | Onthophagus amamiensis amamiensis | | | Onthophagus amamiensis matsumurai Ochi, Kawaha..., 2019 | | | Onthophagus ambang Krikken & Hu..., 2011 | | | Onthophagus ambang ambang | | | Onthophagus ambang morowali Krikken & Hu..., 2011 | | | Onthophagus amenthus Matsumura, 1937 | | | Onthophagus amphicoma Boucomont, 1914 | | | Onthophagus amphinasus Arrow, 1931 | | | Onthophagus amphioxus Arrow, 1933 | | | Onthophagus amycoides Kabakov, 2014 | | | Onthophagus anatolicus Petrovitz, 1962 | | | Onthophagus anchommatus Lea, 1923 | | | Onthophagus andonarensis Boucomont, 1914 | | | Onthophagus andreji Prokofiev, 2014 | | | Onthophagus andrewesi Arrow, 1931 | | | Onthophagus angkhanensis Masumoto, Oc..., 2013 | | | Onthophagus anisocerus Erichson, 1842 | | | Onthophagus annulopunctatus Krikken & Hu..., 2017 | | | Onthophagus aphodioides Lansberge, 1883 | | | Onthophagus apomontanus Ochi, Kon & ..., 2023 | | | Onthophagus apunneea Masumoto, Oc..., 2007 | | | Onthophagus arayai Ochi & Kon, 2007 | | | Onthophagus arboreus Arrow, 1931 | | | Onthophagus ardoini Frey, 1971 | | | Onthophagus armatus Blanchard, 1853 | | | Onthophagus arunachalensis Biswas & Cha..., 1985 | | | Onthophagus arvernus ♰ L. Piton, 1939 | | | Onthophagus asiaticus Endrödi, 1973 | | | Onthophagus asper MacLeay, 1864 | | | Onthophagus asperrimus d´Orbigny, 1902 | | | Onthophagus atropolitus d´Orbigny, 1902 | | | Onthophagus atrox Harold, 1867 | | | Onthophagus aureopilosus Boucomont, 1914 | | | Onthophagus auritus Erichson, 1842 | | | Onthophagus australis Guérin-Ménev..., 1830 | | | Onthophagus avocetta Arrow, 1933 | | | Onthophagus axanicus Ochi, Kon & ..., 2021 | | | Onthophagus axillaris (Boheman, 1860) | ⇒ Onthophagus criniger d´Orbigny, 1904 | | Onthophagus azusae Ochi & Kon, 2006 | | | Onthophagus babirussa Eschscholtz, 1822 | | | Onthophagus baenzigeri Masumoto, Oc..., 2007 | | | Onthophagus baeri Boucomont, 1914 | | | Onthophagus bakeri (Boucomont, 1919) | ⇒ Panelus bakeri Boucomont, 1921 | | Onthophagus balawaicus Scheuern, 1995 | | | Onthophagus balthasari Vsetecka, 1939 | | | Onthophagus bangueyensis Boucomont, 1914 | | | Onthophagus baolocensis Ochi & Kon, 2015 | | | Onthophagus baramtagal Ochi, Kon & ..., 2016 | | | Onthophagus basicruentatus Goidanich, 1926 | | | Onthophagus basipustulatus Heyden, 1879 | | | Onthophagus batillifer Harold, 1875 | | | Onthophagus batui Huijbregts &..., 2009 | | | Onthophagus beesoni Arrow, 1931 | | | Onthophagus begoniophilus Krikken & Hu..., 2017 | | | Onthophagus beiranus Péringuey, 1908 | | | Onthophagus bellus d´Orbigny, 1905 | | | Onthophagus bengalensis Harold, 1886 | | | Onthophagus bengali Gordon & Opp..., 1977 | | | Onthophagus bicarinaticeps Lea, 1923 | | | Onthophagus bicolensis Ochi & Kon, 2006 | | | Onthophagus bicolensis asin Ochi & Kon, 2006 | | | Onthophagus bicolensis bicolensis | | | Onthophagus bicolor (Raffray, 1877) | ⇒ Onthophagus flavolimbatus Klug, 1855 | | Onthophagus bicristatus d´Orbigny, 1905 | | | Onthophagus bicuneus Kabakov, 2008 | | | Onthophagus biexcavatus (d´Orbigny, 1898) | ⇒ Onthophagus tarandus Fabricius, 1792 | | Onthophagus bifasciatus Fabricius, 1781 | Zweibindiger Pillenkäfer | | Onthophagus binodis Thunberg, 1818 | | | Onthophagus biscarinulatus Huijbregts &..., 2012 | | | Onthophagus bisectus Arrow, 1931 | | | Onthophagus bison Boucomont, 1919 | | | Onthophagus bisontinus ♰ Heer, 1862 | | | Onthophagus bisscrutator Krikken & Hu..., 2017 | | | Onthophagus bistiniocelloides Krikken, 1986 | | | Onthophagus bituberoculus Krikken & Hu..., 2013 | | | Onthophagus bivertex Heyden, 1887 | | | Onthophagus blackwoodensis Blackburn, 1891 | | | Onthophagus blanchardi Harold, 1869 | | | Onthophagus blumei (Lansberge, 1883) | ⇒ Onthophagus avocetta Arrow, 1933 | | Onthophagus bongkudai Krikken & Hu..., 2017 | | | Onthophagus bonorae Zunino, 1976 | | | Onthophagus bonsae Zunino, 1976 | | | Onthophagus borneensis Harold, 1877 | | | Onthophagus bornemisszai Matthews, 1972 | | | Onthophagus borneotagal Ochi, Kon & ..., 2016 | | | Onthophagus boucomontianus Balthasar, 1935 | | | Onthophagus bourgognei Paulian, 1945 | | | Onthophagus brevicollis Arrow, 1907 | | | Onthophagus brittoni Paulian, 1948 | | | Onthophagus brivioi Frey, 1973 | | | Onthophagus bronzeus Arrow, 1907 | | | Onthophagus brooksi Matthews, 1972 | | | Onthophagus brutus Arrow, 1931 | | | Onthophagus bunamin Matthews, 1972 | | | Onthophagus caesariatus Boucomont, 1921 | | | Onthophagus calcaratus Boucomont, 1920 | | | Onthophagus cameloides d´Orbigny, 1900 | | | Onthophagus camelus Fabricius, 1787 | | | Onthophagus campestris Masumoto, Ki..., 2018 | | | Onthophagus capella (Kirby, 1818) | ⇒ Onthophagus ramosellus Bates, 1891 | | Onthophagus capelliformis Gillet, 1925 | | | Onthophagus capellinus Frey, 1963 | | | Onthophagus capitosus Harold, 1867 | | | Onthophagus carinulatus Harold, 1877 | | | Onthophagus carpanetoi Pittino, 1982 | | | Onthophagus carpophilus Pereira & Ha..., 1961 | | | Onthophagus castetsi Lansberge, 1887 | | | Onthophagus cavia Boucomont, 1914 | | | Onthophagus caviceps Frey, 1962 | | | Onthophagus centricornis Fabricius, 1798 | | | Onthophagus cervicapra Boucomont, 1914 | | | Onthophagus cervus (Fabricius, 1798) | ⇒ Onthophagus falsus Gillet, 1925 | | Onthophagus ceylonicus Harold, 1877 | | | Onthophagus championi Bates, 1887 | | | Onthophagus chandrai Ochi, 2007 | | | Onthophagus chebaicus Boucomont, 1919 | | | Onthophagus chepara Matthews, 1972 | | | Onthophagus chetroiensis Masumoto, Oc..., 2012 | | | Onthophagus cheyi Ochi & Kon, 2006 | | | Onthophagus chiangdaolekus Masumoto, Oc..., 2020 | | | Onthophagus chibanus Matsumura & ..., 1937 | | | Onthophagus chinbucephalus Ochi & Kon, 2018 | | | Onthophagus chinvigilans Ochi & Kon, 2018 | | | Onthophagus chokakurianus Matsumura, 1938 | | | Onthophagus chosensis Matsumura, 1937 | | | Onthophagus chryses Bates, 1887 | | | Onthophagus chrysurus Arrow, 1931 | | | Onthophagus chumphonensis Masumoto, Oc..., 2008 | | | Onthophagus chuzenjianus (Matsumura & ..., 1937) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus circulator Reitter, 1891 | | | Onthophagus circulifer Arrow, 1931 | | | Onthophagus citreum Boucomont, 1919 | | | Onthophagus clermonti Paulian, 1931 | | | Onthophagus clivimerus Huijbregts &..., 2011 | | | Onthophagus coeruleicollis Arrow, 1907 | | | Onthophagus cognatus Boucomont, 1921 | | | Onthophagus colffsi Lansberge, 1883 | | | Onthophagus collaris (Kabakov, 1983) | ⇒ Proagoderus loricatus Klug, 1855 | | Onthophagus collinsi Krikken & Hu..., 1987 | | | Onthophagus commottoides Kabakov, 1998 | | | Onthophagus comottoi Lansberge, 1885 | | | Onthophagus compactus Arrow, 1933 | | | Onthophagus concolor Sharp, 1878 | | | Onthophagus confluens (d´Orbigny, 1902) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus consentaneus Harold, 1867 | | | Onthophagus conspicuus MacLeay, 1864 | | | Onthophagus convexicollis Boheman, 1858 | | | Onthophagus coomani Paulian, 1931 | | | Onthophagus coorgensis Arrow, 1931 | | | Onthophagus coprimorphoides Huijbregts &..., 2012 | | | Onthophagus coracinoides Kabakov, 1983 | | | Onthophagus coracinus Boucomont, 1914 | | | Onthophagus crassicollis Boucomont, 1913 | | | Onthophagus crassus ♰ (Heer, 1862) | ⇒ Parascatonomus tricornis Wiedemann, 1823 | | Onthophagus cribratus Lansberge, 1883 | | | Onthophagus cryptogenus Boucomont, 1914 | | | Onthophagus cuneus Boucomont, 1924 | | | Onthophagus cuniculus MacLeay, 1864 | | | Onthophagus cupreiceps Arrow, 1907 | | | Onthophagus cupreopastillatus Ochi & Kon, 2006 | | | Onthophagus cupreotagal Ochi, Kon & ..., 2016 | | | Onthophagus curvispina Reitter, 1893 | | | Onthophagus cyaneus Kabakov, 2006 | | | Onthophagus cyanochlorus d´Orbigny, 1902 | | | Onthophagus dandalu Matthews, 1972 | | | Onthophagus danumicus Huijbregts &..., 2012 | | | Onthophagus dapitanensis Boucomont, 1919 | | | Onthophagus dayacus Boucomont, 1914 | | | Onthophagus deccanensis Frey, 1962 | | | Onthophagus declivis Harold, 1869 | | | Onthophagus deflexicollis Lansberge, 1883 | | | Onthophagus deliensis Lansberge, 1885 | | | Onthophagus deliensis bawangicus Ochi, Kon & ..., 2009 | | | Onthophagus deliensis deliensis | | | Onthophagus denticornis Boucomont, 1914 | | | Onthophagus depressicollis Boucomont, 1914 | | | Onthophagus desectus MacLeay, 1871 | | | Onthophagus devagiriensis Schoolmeeste..., 2006 | | | Onthophagus dhanjuricus Frey, 1973 | | | Onthophagus difficilis (Le Guillou, 1844) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus digitatus Arrow, 1931 | | | Onthophagus dinoderus d´Orbigny, 1913 | | | Onthophagus discolor (Hope, 1841) | ⇒ Onthophagus impurus Harold, 1868 | | Onthophagus dissentaneus Balthasar, 1964 | | | Onthophagus doriae Harold, 1877 | | | Onthophagus dorsofasciatus Fairmaire, 1893 | | | Onthophagus dorsosignatus d´Orbigny, 1898 | | | Onthophagus drescheri Paulian, 1939 | | | Onthophagus dunningi Harold, 1869 | | | Onthophagus ebenus Péringuey, 1888 | | | Onthophagus echinus Boucomont, 1914 | | | Onthophagus egenus (Harold, 1877) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus egurianus Matsumura, 1938 | | | Onthophagus elongatus Frey, 1954 | | | Onthophagus emeritus Péringuey, 1901 | | | Onthophagus ensifer Boucomont, 1914 | | | Onthophagus ephippioderus Arrow, 1907 | | | Onthophagus eschscholtzi Boucomont, 1924 | | | Onthophagus euplynes Bates, 1888 | | | Onthophagus evanidus Harold, 1869 | | | Onthophagus excavatus (Redtenbacher, 1848) | ⇒ Liatongus phanaeoides Westwood, 1839 | | Onthophagus expansicornis Bates, 1891 | | | Onthophagus exquisitus Arrow, 1931 | | | Onthophagus fabricianus Goidanich, 1926 | | | Onthophagus fabricii Waterhouse, 1894 | | | Onthophagus falcifer Harold, 1880 | | | Onthophagus falculatus Boucomont, 1914 | | | Onthophagus falsus Gillet, 1925 | | | Onthophagus fasciatus (Boucomont, 1914) | ⇒ Onthophagus fasciolatus Boucomont, 1924 | | Onthophagus fasciolatus Boucomont, 1924 | | | Onthophagus favrei Boucomont, 1914 | | | Onthophagus felix Arrow, 1931 | | | Onthophagus ferox Harold, 1867 | | | Onthophagus ferrari Matthews, 1972 | | | Onthophagus filicornis Harold, 1873 | | | Onthophagus fimetarius Roth, 1851 | | | Onthophagus finchi | | | Onthophagus fissiceps MacLeay, 1888 | | | Onthophagus fissinasus Fairmaire, 1895 | | | Onthophagus flagrans Reitter, 1893 | | | Onthophagus flavoapicalis Lea, 1923 | | | Onthophagus flavolimbatus Klug, 1855 | | | Onthophagus fletcheri Blackburn, 1903 | | | Onthophagus fodiens Waterhouse, 1875 | | | Onthophagus foedus Boucomont, 1914 | | | Onthophagus formaneki Reitter, 1897 | | | Onthophagus fossor Arrow, 1931 | | | Onthophagus frankenbergeri Balthasar, 1933 | | | Onthophagus frankrozendaali Huijbregts &..., 2012 | | | Onthophagus frenchi Blackburn, 1903 | | | Onthophagus froggattellus Monteith & R..., 2017 | | | Onthophagus frontalis Raffray, 1877 | | | Onthophagus frugivorus Arrow, 1931 | | | Onthophagus fugitivus Péringuey, 1901 | | | Onthophagus fukuyamai Ochi, Kon & ..., 2018 | | | Onthophagus fuliginosus Erichson, 1842 | | | Onthophagus fulvus Sharp, 1875 | | | Onthophagus furcaticeps Masters, 1886 | | | Onthophagus furcatoides (Lansberge, 1886) | ⇒ Onthophagus neofurcatus Goidanich, 1926 | | Onthophagus furcicollis (Arrow, 1931) | ⇒ Onthophagus hecate Panzer, 1794 | | Onthophagus furcillifer Bates, 1891 | | | Onthophagus furculus Fabricius, 1798 | | | Onthophagus fuscopunctatus (Fabricius, 1798) | ⇒ Onthophagus luridipennis Boheman, 1858 | | Onthophagus fuscostriatus Boucomont, 1914 | | | Onthophagus gaesatus Boucomont, 1925 | | | Onthophagus gagates Hope, 1831 | | | Onthophagus gagatoides Kabakov, 2006 | | | Onthophagus gajo Krikken, 1977 | | | Onthophagus galeatus Boucomont, 1919 | | | Onthophagus gangeticus Gillet, 1925 | | | Onthophagus gangulu Matthews, 1972 | | | Onthophagus geelongensis Blackburn, 1891 | | | Onthophagus gemma Sharp, 1875 | | | Onthophagus germanus Gillet, 1927 | | | Onthophagus gestroi (Harold, 1877) | ⇒ Onthophagus bellus d´Orbigny, 1905 | | Onthophagus gibbicollis Lansberge, 1885 | | | Onthophagus ginyunensis Vsetecka, 1942 | | | Onthophagus giraffa Hausmann, 1807 | | | Onthophagus giuseppecarpanetoi Tagliaferri ..., 2012 | | | Onthophagus glabratus Hope, 1842 | | | Onthophagus glasunovi Koshantschikov, 1894 | | | Onthophagus glasunowi D. Koshantsc..., 1894 | | | Onthophagus gonipa Krikken & Hu..., 2011 | | | Onthophagus granulatus (Boheman, 1858) | ⇒ Onthophagus consentaneus Harold, 1867 | | Onthophagus granulifer (Harold, 1886) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus gratus Arrow, 1931 | | | Onthophagus gravis Walker, 1858 | | | Onthophagus griseoaeneus Lansberge, 1885 | | | Onthophagus griseosetosus Arrow, 1931 | | | Onthophagus guatemalensis (Bates, 1887) | ⇒ Onthophagus medorensis Brown, 1929 | | Onthophagus gulmarri Matthews, 1972 | | | Onthophagus gulo Arrow, 1931 | | | Onthophagus haagi Harold, 1867 | | | Onthophagus hageni Lansberge, 1883 | | | Onthophagus hamaticeps Arrow, 1931 | | | Onthophagus hanboonsongae Masumoto, Oc..., 2021 | | | Onthophagus hartiniae Ochi & Kon, 2009 | | | Onthophagus hemipygus Frey, 1964 | | | Onthophagus hermonensis Baraud, 1982 | | | Onthophagus herus Péringuey, 1901 | | | Onthophagus heterorrhinus Lansberge, 1885 | | | Onthophagus heydeni Harold, 1875 | | | Onthophagus heyrovskyi Vsetecka, 1943 | | | Onthophagus hielkemai Krikken & Hu..., 2011 | | | Onthophagus hikidai Ochi & Kon, 2006 | | | Onthophagus hikosanus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus hindu Arrow, 1931 | | | Onthophagus hingstoni Arrow, 1931 | | | Onthophagus hiranoyasutoshii Ochi & Kon, 2018 | | | Onthophagus hiroyukii Ochi, 2007 | | | Onthophagus hirsutulus Lansberge, 1883 | | | Onthophagus hirtellus Frey, 1957 | | | Onthophagus histeriformis Boucomont, 1914 | | | Onthophagus histrio d´Orbigny, 1902 | | | Onthophagus hoberlandti Balthasar, 1966 | | | Onthophagus hollowayi Krikken & Hu..., 2017 | | | Onthophagus holosericeus Harold, 1877 | | | Onthophagus holosericus Harold, 1877 | | | Onthophagus holzi Boucomont, 1914 | | | Onthophagus hoplocerus Lea, 1923 | | | Onthophagus hosomai Ochi & Kon, 2014 | | | Onthophagus hyaena Fabricius, 1801 | | | Onthophagus hystrix Boucomont, 1914 | | | Onthophagus ibex (Fabricius, 1792) | ⇒ Onthophagus tragus Fabricius, 1792 | | Onthophagus ibonus (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus ifugaoensis Ochi & Kon, 2006 | | | Onthophagus igneus Vigors, 1825 | | | Onthophagus imbutus Sharp, 1875 | | | Onthophagus immundus Boheman, 1858 | | | Onthophagus imperator Castelnau, 1840 | | | Onthophagus incanus MacLeay, 1888 | | | Onthophagus incertus (d´Orbigny, 1897) | ⇒ Onthophagus asperrimus d´Orbigny, 1902 | | Onthophagus incisus Harold, 1877 | | | Onthophagus incollaris Kabakov, 2008 | | | Onthophagus incornutus MacLeay, 1871 | | | Onthophagus indosinicus Boucomont, 1921 | | | Onthophagus indutus d´Orbigny, 1902 | | | Onthophagus inermivertex Boucomont, 1921 | | | Onthophagus infucatus Harold, 1877 | | | Onthophagus infuscatus Klug, 1845 | | | Onthophagus insulindicus Boucomont, 1914 | | | Onthophagus interstitialis (Fahraeus, 1857) | ⇒ Euonthophagus carbonarius Klug, 1855 | | Onthophagus isanus Masumoto, Oc..., 2008 | | | Onthophagus itoi Nomura, 1976 | | | Onthophagus itoianus Masumoto, Oc..., 2022 | | | Onthophagus itonoborui Masumoto, Oc..., 2012 | | | Onthophagus iumienus Masumoto, Oc..., 2007 | | | Onthophagus iyengari Arrow, 1931 | | | Onthophagus jacobeus Boucomont, 1924 | | | Onthophagus jangga Matthews, 1972 | | | Onthophagus javanensis Balthasar, 1963 | | | Onthophagus javanoides Huijbregts &..., 2011 | | | Onthophagus jeannelianus Paulian, 1945 | | | Onthophagus jeannelianus jeannelianus | | | Onthophagus jeannelianus shokhini Kabakov, 2014 | | | Onthophagus jedensis Matsumura, 1937 | | | Onthophagus jingping Masumoto, Oc..., 2007 | | | Onthophagus jiupengensis Masumoto, La..., 2017 | | | Onthophagus jubatus Harold, 1869 | | | Onthophagus jugicola d´Orbigny, 1902 | | | Onthophagus juvencus Klug, 1835 | | | Onthophagus jwalae Karimbumkara..., 2016 | | | Onthophagus kachinicus Kabakov, 2006 | | | Onthophagus kaengkrachangus Masumoto, Oc..., 2008 | | | Onthophagus kanarensis Arrow, 1931 | | | Onthophagus kandai (Matsumura, 1937) | ⇒ Onthophagus olsoufieffi Boucomont, 1924 | | Onthophagus kangeanus Paulian, 1936 | | | Onthophagus kaosoidowensis Masumoto, Oc..., 2008 | | | Onthophagus kapitensis Frey, 1971 | | | Onthophagus kapuri Endrödi, 1974 | | | Onthophagus kashizakii Kon & Ochi, 2005 | | | Onthophagus kashmirensis Balthasar, 1966 | | | Onthophagus katayamai Masumoto, Oc..., 2012 | | | Onthophagus kawaharai Ochi & Kon, 2007 | | | Onthophagus kawarinus (Matsumura & ..., 1937) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus kchatriya Boucomont, 1914 | | | Onthophagus keikoae Ochi & Kon, 2014 | | | Onthophagus keiseri Frey, 1956 | | | Onthophagus keralensis Frey, 1975 | | | Onthophagus keralicus Biswas & Cha..., 1986 | | | Onthophagus kiambram Storey, 1977 | | | Onthophagus kiyoshii Ochi & Kon, 2007 | | | Onthophagus klapperichi Balthasar, 1952 | | | Onthophagus kleinei Balthasar, 1935 | | | Onthophagus knapperti Krikken, 1981 | | | Onthophagus koebelei Blackburn, 1903 | | | Onthophagus koechlei Masumoto, Oc..., 2011 | | | Onthophagus kogatanus (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus kokereka Matthews, 1972 | | | Onthophagus kolaka Huijbregts &..., 2009 | | | Onthophagus koma Matsumura, 1937 | | | Onthophagus komabellus Matsumura & ..., 1937 | | | Onthophagus komareki Balthasar, 1935 | | | Onthophagus kongkaewensis Masumoto, Oc..., 2012 | | | Onthophagus koni Ochi, 2007 | | | Onthophagus konsarnensis Masumoto, Oc..., 2008 | | | Onthophagus kontumicus Kabakov, 1983 | | | Onthophagus kozunonis (Matsumura, 1937) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus kulti Balthasar, 1952 | | | Onthophagus kuluensis Bates, 1891 | | | Onthophagus kumaonensis Arrow, 1931 | | | Onthophagus kyokoae Masumoto & Ochi, 2015 | | | Onthophagus laborans Arrow, 1931 | | | Onthophagus lacustris Harold, 1877 | | | Onthophagus laevigatus Fabricius, 1798 | | | Onthophagus laevissimus d´Orbigny, 1905 | | | Onthophagus lahorensis Kabakov, 2008 | | | Onthophagus laichauensis Ochi, Kon & ..., 2021 | | | Onthophagus lamelliger (Gerstaecker, 1871) | ⇒ Onthophagus ochropygus d´Orbigny, 1902 | | Onthophagus laminatus (MacLeay, 1864) | ⇒ Onthophagus lunatus Harold, 1868 | | Onthophagus lamyi Paulian, 1945 | | | Onthophagus langkawiensis Ochi & Kon, 2015 | | | Onthophagus lapillus Arrow, 1931 | | | Onthophagus laratinus Arrow, 1916 | | | Onthophagus latipennis d´Orbigny, 1897 | | | Onthophagus latro Harold, 1877 | | | Onthophagus leai Blackburn, 1895 | | | Onthophagus lebasi Boucomont, 1932 | | | Onthophagus lekmaengtibanphrao Masumoto & Ochi, 2015 | | | Onthophagus leknamnaous Masumoto & Ochi, 2015 | | | Onthophagus lemniscatus Gillet, 1924 | | | Onthophagus lemuroides d´Orbigny, 1898 | | | Onthophagus lenzii Harold, 1874 | | | Onthophagus leomontanus Boucomont, 1914 | | | Onthophagus leusermontis Huijbregts &..., 2011 | | | Onthophagus lindu Huijbregts &..., 2009 | | | Onthophagus liothorax Koshantschikov, 1894 | | | Onthophagus liui Masumoto, Oc..., 2014 | | | Onthophagus liwagensis Ochi & Kon, 2006 | | | Onthophagus longipes Paulian, 1937 | | | Onthophagus lore Huijbregts &..., 2009 | | | Onthophagus loricatus Klug, 1855 | | | Onthophagus luctuosus Boucomont, 1914 | | | Onthophagus ludio Boucomont, 1914 | | | Onthophagus lugubris (Fahraeus, 1857) | ⇒ Onthophagus lituratus Roth, 1851 | | Onthophagus lunatus Harold, 1868 | | | Onthophagus lunulifer Boucomont, 1914 | | | Onthophagus luteosignatus Lansberge, 1883 | | | Onthophagus luteus ♰ Oustalet, 1874 | | | Onthophagus luzonwallacei Ochi, Kawaha..., 2021 | | | Onthophagus macleayi Blackburn, 1903 | | | Onthophagus macrocephalus Kirby, 1818 | | | Onthophagus madoqua Arrow, 1931 | | | Onthophagus maephaluangus Masumoto, Oc..., 2007 | | | Onthophagus maesalongensis Masumoto, Oc..., 2013 | | | Onthophagus magnini Paulian, 1933 | | | Onthophagus magnioculus Ochi & Kon, 2006 | | | Onthophagus magnipygus Boucomont, 1914 | | | Onthophagus malabarensis Boucomont, 1919 | | | Onthophagus malangensis Boucomont, 1914 | | | Onthophagus malasiacus Gillet, 1927 | | | Onthophagus malthinus Gillet, 1930 | | | Onthophagus mamillatus Lea, 1923 | | | Onthophagus manguliensis Boucomont, 1914 | | | Onthophagus manya Matthews, 1972 | | | Onthophagus marginatus (Castelnau, 1840) | ⇒ Onthophagus andalusicus Waltl, 1835 | | Onthophagus marginicollis Harold, 1880 | | | Onthophagus martialis Boucomont, 1914 | | | Onthophagus maruchanus Masumoto, Oc..., 2012 | | | Onthophagus maruyamai Ochi, Kon & ..., 2014 | | | Onthophagus maryatiae Ochi & Kon, 2005 | | | Onthophagus matanyo Huijbregts &..., 2009 | | | Onthophagus matsuii Ochi & Kon, 2006 | | | Onthophagus matsuokai (Matsumura & ..., 1937) | ⇒ Onthophagus sauteri Gillet, 1924 | | Onthophagus mauritii Boucomont, 1919 | | | Onthophagus megapacificus Ochi & Kon, 2006 | | | Onthophagus megathorax Gillet, 1921 | | | Onthophagus mekara Huijbregts &..., 2009 | | | Onthophagus melanocephalus Klug, 1845 | | | Onthophagus mentaveiensis Boucomont, 1914 | | | Onthophagus mexicanus Bates, 1887 | | | Onthophagus micellus Matsumura, 1937 | | | Onthophagus midorianus (Matsumura, 1938) | ⇒ Parascatonomus anguicorius Boucomont, 1914 | | Onthophagus miichunus Masumoto, Oc..., 2020 | | | Onthophagus militaris Boucomont, 1914 | | | Onthophagus millingeni d´Orbigny, 1900 | | | Onthophagus mimikanus Balthasar, 1969 | | | Onthophagus mindanaensis Boucomont, 1914 | | | Onthophagus mindanaoensis Frey, 1971 | | | Onthophagus minoi Matsumura & ..., 1937 | | | Onthophagus minokuchianus (Matsumura, 1934) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus minutus (Hausmann, 1807) | ⇒ Onthophagus centricornis Fabricius, 1798 | | Onthophagus mirandus Arrow, 1931 | | | Onthophagus misujianus Matsumura & ..., 1937 | | | Onthophagus miyabei (Matsumura, 1937) | ⇒ Onthophagus punctator Reitter, 1893 | | Onthophagus miyazakianus Matsumura & ..., 1937 | | | Onthophagus mjobergi Gillet, 1925 | | | Onthophagus mniszechi Harold, 1869 | | | Onthophagus moajat Huijbregts &..., 2009 | | | Onthophagus modestus Harold, 1862 | | | Onthophagus mogo Krikken & Hu..., 2008 | | | Onthophagus montishannoniae Krikken & Hu..., 2008 | | | Onthophagus mopsus Fabricius, 1792 | | | Onthophagus mordindangus Masumoto, Oc..., 2008 | | | Onthophagus moreleti Baraud, 1980 | | | Onthophagus morimotoi Masumoto, Oc..., 2021 | | | Onthophagus mouhoti Harold, 1875 | | | Onthophagus mulgravei Paulian, 1937 | | | Onthophagus multipunctatus Boucomont, 1914 | | | Onthophagus mulutagal Ochi, Kon & ..., 2016 | | | Onthophagus mushanus Matsumura, 1938 | | | Onthophagus mushensis Matsumura, 1938 | | | Onthophagus mutatus Harold, 1859 | | | Onthophagus muticus MacLeay, 1864 | | | Onthophagus nagasawai Matsumura, 1938 | | | Onthophagus nagpurensis Arrow, 1931 | | | Onthophagus nakatomii (Matsumura, 1937) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus namnaolekus Masumoto, Oc..., 2020 | | | Onthophagus namnaonus Masumoto, Oc..., 2020 | | | Onthophagus namnaous Masumoto, Oc..., 2013 | | | Onthophagus nasalis Arrow, 1931 | | | Onthophagus nebulosus Reiche, 1864 | | | Onthophagus necessarius Reitter, 1893 | | | Onthophagus necrophagus Arrow, 1931 | | | Onthophagus neervoorti Boucomont, 1914 | | | Onthophagus nefarius Balthasar, 1941 | | | Onthophagus negligens Walker, 1858 | | | Onthophagus neofurcatus Goidanich, 1926 | | | Onthophagus neostenocerus Goidanich, 1926 | | | Onthophagus nicobaricus Biswas, Chat..., 1999 | | | Onthophagus nitefactus Harold, 1877 | | | Onthophagus nitidulus Klug, 1845 | | | Onthophagus nitidus Waterhouse, 1875 | | | Onthophagus nodulifer Harold, 1867 | | | Onthophagus nutans (Roth, 1851) | ⇒ Onthophagus schaufussi Harold, 1867 | | Onthophagus oberthueri d´Orbigny, 1898 | | | Onthophagus oberthuri d´Orbigny, 1898 | | | Onthophagus obliviosus Balthasar, 1935 | | | Onthophagus obscurior Boucomont, 1914 | | | Onthophagus obtusicornis (Fahraeus, 1857) | ⇒ Onthophagus marginalis przewalski Kabakov, 2006 | | Onthophagus obtutus Péringuey, 1901 | | | Onthophagus ocellatopunctatus Waterhouse, 1875 | | | Onthophagus ocelliger Harold, 1877 | | | Onthophagus ochii Masumoto, 1988 | | | Onthophagus ochreatus d´Orbigny, 1897 | | | Onthophagus ochromerus Harold, 1877 | | | Onthophagus octogonus Frey, 1962 | | | Onthophagus oculatus Arrow, 1931 | | | Onthophagus ohbayashii Nomura, 1939 | | | Onthophagus ohkuboi Ochi & Kon, 2006 | | | Onthophagus oishii Matsumura & ..., 1937 | | | Onthophagus okushiranus Matsumura & ..., 1937 | | | Onthophagus okushirianus Matsumura, 1937 | | | Onthophagus oniellus (Matsumura, 1937) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus ontosatu Huijbregts &..., 2012 | | | Onthophagus opacihartiniae Ochi & Kon, 2015 | | | Onthophagus orbus Boucomont, 1924 | | | Onthophagus orcas (Ménétriés, 1838) | ⇒ Onthophagus atramentarius Ménétriés, 1832 | | Onthophagus orissanus Arrow, 1931 | | | Onthophagus orphnoides (Bates, 1887) | ⇒ Onthophagus mirabilis Bates, 1887 | | Onthophagus osellai Pittino, 1982 | | | Onthophagus oshimanus Nakane, 1960 | | | Onthophagus otai Ochi & Kon, 2006 | | | Onthophagus ovatulus ♰ Heer, 1847 | | | Onthophagus pacificus Lansberge, 1885 | | | Onthophagus pactolus Fabricius, 1787 | | | Onthophagus palamoui Biswas & Cha..., 1986 | | | Onthophagus palatus Boucomont, 1914 | | | Onthophagus palavanus (Huijbregts &..., 2011) | ⇒ Onthophagus javanensis Balthasar, 1963 | | Onthophagus palawantagal Ochi, Kon & ..., 2016 | | | Onthophagus pallidipennis (Fahraeus, 1857) | ⇒ Onthophagus rasipennis d´Orbigny, 1908 | | Onthophagus paluma Matthews, 1972 | | | Onthophagus pangmaphanus Masumoto, Oc..., 2022 | | | Onthophagus paracenricornis Ochi & Kon, 2014 | | | Onthophagus paracentricornis Ochi & Kon, 2014 | | | Onthophagus parafasciatus Balthasar, 1974 | | | Onthophagus parapalatus Krikken & Hu..., 1988 | | | Onthophagus pardalis Fabricius, 1798 | | | Onthophagus parenthesis Boucomont, 1913 | | | Onthophagus paroculus Krikken & Hu..., 1987 | | | Onthophagus parryi Harold, 1869 | | | Onthophagus parumnotatus Fahraeus, 1857 | | | Onthophagus parvidens (Frey, 1971) | ⇒ Onthophagus clermonti Paulian, 1931 | | Onthophagus parvulus (Fabricius, 1798) | ⇒ Onthophagus frontalis Raffray, 1877 | | Onthophagus parvus Blanchard, 1853 | | | Onthophagus pasotagal Ochi, Kon & ..., 2016 | | | Onthophagus pastillatus Boucomont, 1919 | | | Onthophagus paulianellus Frey, 1958 | | | Onthophagus pauliani (Balthasar, 1937) | ⇒ Onthophagus paulianellus Frey, 1958 | | Onthophagus pauper Boucomont, 1914 | | | Onthophagus pavidus Harold, 1877 | | | Onthophagus pedator Sharp, 1875 | | | Onthophagus peninsularis Boucomont, 1914 | | | Onthophagus peninsulomerus Huijbregts &..., 2011 | | | Onthophagus pentacanthus Harold, 1867 | | | Onthophagus perakensis Ochi, Kon & ..., 2014 | | | Onthophagus peramelinus Lea, 1923 | | | Onthophagus perpilosus MacLeay, 1871 | | | Onthophagus personatus Boucomont, 1914 | | | Onthophagus pexatus Harold, 1869 | | | Onthophagus phahompokus Masumoto & Ochi, 2015 | | | Onthophagus phakuaiensis Masumoto, Oc..., 2013 | | | Onthophagus phanaeicollis Lansberge, 1883 | | | Onthophagus phanaeides Frey, 1956 | | | Onthophagus phanaeiformis Boucomont, 1914 | | | Onthophagus phanaeoides (Westwood, 1839) | ⇒ Proagoderus lanista Castelnau, 1840 | | Onthophagus phatoensis Masumoto, Oc..., 2008 | | | Onthophagus philippinensis Boucomont, 1919 | | | Onthophagus phillippsorum Krikken & Hu..., 1987 | | | Onthophagus phukhieoensis Masumoto, Oc..., 2007 | | | Onthophagus pictus (Reitter, 1893) | ⇒ Oniticellus tessellatus Harold, 1879 | | Onthophagus piffli Petrovitz, 1961 | | | Onthophagus pilosus (Fahraeus, 1857) | ⇒ Onthophagus echinus Boucomont, 1914 | | Onthophagus pimpasaleei Masumoto, Oc..., 2008 | | | Onthophagus pithankithae Karimbumkara..., 2016 | | | Onthophagus planicollis Harold, 1880 | | | Onthophagus planifrons Frey, 1962 | | | Onthophagus pljushtchi Ivanova, 2012 | | | Onthophagus poeophagus Kabakov, 1979 | | | Onthophagus pollicatus Harold, 1879 | | | Onthophagus porcus Arrow, 1931 | | | Onthophagus poringensis Ochi & Kon, 2005 | | | Onthophagus posticus Erichson, 1842 | | | Onthophagus praecellens Bates, 1887 | | | Onthophagus praedatus Harold, 1862 | | | Onthophagus prehensilis Arrow, 1920 | | | Onthophagus probus Péringuey, 1901 | | | Onthophagus procurvus Balthasar, 1935 | | | Onthophagus prodromus ♰ Heer, 1862 | | | Onthophagus productus Arrow, 1907 | | | Onthophagus pronus Erichson, 1842 | | | Onthophagus propinquus (MacLeay, 1888) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus pseudoarai Ochi & Kon, 2018 | | | Onthophagus pseudobrutus Kabakov, 1983 | | | Onthophagus pseudoconvexicollis Bai & Yang, 2016 | | | Onthophagus pseudocoracinus Kabakov, 1983 | | | Onthophagus pseudojaponicus Balthasar, 1941 | | | Onthophagus pseudojaponicus fukiensis Balthasar, 1941 | | | Onthophagus pseudojaponicus pseudojaponicus | | | Onthophagus ptox Erichson, 1842 | | | Onthophagus pugionatus Fahraeus, 1857 | | | Onthophagus pugnacior Blackburn, 1903 | | | Onthophagus pugnax Harold, 1868 | | | Onthophagus pumilus Kabakov, 2014 | | | Onthophagus pupillatus Kolbe, 1886 | | | Onthophagus purpurascens Boucomont, 1914 | | | Onthophagus purpureicollis MacLeay, 1864 | | | Onthophagus pusillus (Fabricius, 1798) | ⇒ Onthophagus modestus Harold, 1862 | | Onthophagus putealis (Matsumura, 1938) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus pygidialis Lansberge, 1883 | | | Onthophagus pygmaeus Schaller, 1783 | | | Onthophagus quadrinodosus Fahraeus, 1857 | | | Onthophagus quadripustulatus Fabricius, 1775 | Neuholländische Vierblatter | | Onthophagus quangnamensis Ochi, Kon & ..., 2021 | | | Onthophagus quasijohkii Ochi & Kon, 2005 | | | Onthophagus quasitagal Ochi & Kon, 2005 | | | Onthophagus queenslandicus Blackburn, 1903 | | | Onthophagus ramosellus Bates, 1891 | | | Onthophagus ramosus Wiedemann, 1823 | | | Onthophagus rana Arrow, 1931 | | | Onthophagus ranongensis Masumoto, Oc..., 2008 | | | Onthophagus ranongjohkii Ochi, Kon & ..., 2016 | | | Onthophagus ratchataniensis Masumoto, Oc..., 2013 | | | Onthophagus refulgens Arrow, 1931 | | | Onthophagus regalis Arrow, 1907 | | | Onthophagus remotus Harold, 1879 | | | Onthophagus rhodesianus Frey, 1975 | | | Onthophagus rhyncophorus | | | Onthophagus ribbei Boucomont, 1914 | | | Onthophagus rizali Ochi, Kawaha..., 2021 | | | Onthophagus robertopoggii Ochi & Kon, 2006 | | | Onthophagus roessneri Ziani, 2016 | | | Onthophagus rorarius Harold, 1877 | | | Onthophagus rosenbergi Krikken & Hu..., 2011 | | | Onthophagus rosenbergi divergens Krikken & Hu..., 2011 | | | Onthophagus rosenbergi rosenbergi | | | Onthophagus rotundicollis Lansberge, 1883 | | | Onthophagus rouyeri Boucomont, 1914 | | | Onthophagus royi Biswas & Cha..., 1985 | | | Onthophagus rubicundulus MacLeay, 1871 | | | Onthophagus rubrimaculatus MacLeay, 1864 | | | Onthophagus rubripennis (Arrow, 1907) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus ruficauda Arrow, 1931 | | | Onthophagus rufulgens | | | Onthophagus rugicollis Harold, 1880 | | | Onthophagus rugulipennis Fairmaire, 1887 | | | Onthophagus rupicapra Waterhouse, 1894 | | | Onthophagus rutilans (Sharp, 1875) | ⇒ Onthophagus borneensis Harold, 1877 | | Onthophagus rutilans aborneensis Ochi, Kon & ..., 2009 | | | Onthophagus rutilans rutilans | | | Onthophagus sabahensis Ochi & Kon, 2006 | | | Onthophagus sabai Masumoto, Oc..., 2012 | | | Onthophagus sacharovskii Olsoufieff, 1918 | | | Onthophagus sahai Biswas & Cha..., 1986 | | | Onthophagus saigonensis Boucomont, 1924 | | | Onthophagus saksphus Masumoto, Oc..., 2021 | | | Onthophagus samarensis Ochi, Kon & ..., 2023 | | | Onthophagus samoengus Masumoto, Oc..., 2008 | | | Onthophagus sanggona Huijbregts &..., 2009 | | | Onthophagus sangirensis Boucomont, 1914 | | | Onthophagus sangwalus Masumoto, Oc..., 2008 | | | Onthophagus sarasinorum Krikken & Hu..., 2011 | | | Onthophagus sceptrifer Boucomont, 1924 | | | Onthophagus schaefernai Balthasar, 1935 | | | Onthophagus schatzmayri (Pierotti, 1959) | ⇒ Onthophagus opacicollis Reitter, 1893 | | Onthophagus schaufussi Harold, 1867 | | | Onthophagus schawalleri Scheuern, 1996 | | | Onthophagus schillhammeri Kabakov, 2006 | | | Onthophagus sciron Balthasar, 1969 | | | Onthophagus scrutator Harold, 1877 | | | Onthophagus sellatus Klug, 1845 | | | Onthophagus sembeli Krikken & Hu..., 2008 | | | Onthophagus semiaereopictus Ochi, Kawaha..., 2021 | | | Onthophagus semicinctus d´Orbigny, 1897 | | | Onthophagus semiflavus (Boheman, 1860) | ⇒ Onthophagus flavolimbatus Klug, 1855 | | Onthophagus semigranosus Lansberge, 1883 | | | Onthophagus semimetallicus Lea, 1923 | | | Onthophagus semipacificus Ochi & Kon, 2006 | | | Onthophagus semiperakensis Ochi, Kon & ..., 2014 | | | Onthophagus semiworoae Ochi, Kon & ..., 2014 | | | Onthophagus senescens Péringuey, 1908 | | | Onthophagus senex Boucomont, 1914 | | | Onthophagus seniculus (Fabricius, 1781) | ⇒ Onthophagus senescens Péringuey, 1908 | | Onthophagus sepilokensis Ochi & Kon, 2006 | | | Onthophagus seramicus Huijbregts &..., 2012 | | | Onthophagus serdangensis Lansberge, 1886 | | | Onthophagus seseba Krikken & Hu..., 2017 | | | Onthophagus setoculus Krikken & Hu..., 1987 | | | Onthophagus sexdentatus Boucomont, 1919 | | | Onthophagus shaykh Montanaro & ..., 2022 | | | Onthophagus shibatai Nakane, 1960 | | | Onthophagus shigeo (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus shigeoi Matsumura, 1937 | | | Onthophagus shinanensis (Matsumura, 1934) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus shinichii Ochi & Kon, 2018 | | | Onthophagus shirakii Nakane, 1960 | | | Onthophagus shoyozanus (Matsumura, 1937) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus shunichii Ochi, Kon & ..., 2021 | | | Onthophagus shurianus Matsumura & ..., 1937 | | | Onthophagus sibela Huijbregts &..., 2012 | | | Onthophagus sibuyanus Boucomont, 1924 | | | Onthophagus sidangeaus Masumoto, Oc..., 2021 | | | Onthophagus sideki Krikken & Hu..., 1987 | | | Onthophagus sihkahonoi Huijbregts &..., 2009 | | | Onthophagus sikkimensis Gillet, 1925 | | | Onthophagus silus Balthasar, 1960 | | | Onthophagus simboroni Ochi & Kon, 2006 | | | Onthophagus simius Reitter, 1893 | | | Onthophagus simplicifrons Reitter, 1893 | | | Onthophagus sinagai Krikken & Hu..., 2008 | | | Onthophagus sinicus Zhang & Wang, 1997 | | | Onthophagus sloanei Blackburn, 1903 | | | Onthophagus smetanai Balthasar, 1952 | | | Onthophagus sobosanus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus socialis Arrow, 1931 | | | Onthophagus solivagus Harold, 1886 | | | Onthophagus solmani Stebnicka, 1975 | | | Onthophagus sonani Miwa, 1930 | | | Onthophagus songsokensis Biswas & Cha..., 1985 | | | Onthophagus sopu Krikken & Hu..., 2017 | | | Onthophagus sparsepunctatus Frey, 1956 | | | Onthophagus sparsulus Reitter, 1893 | | | Onthophagus spathatus Boucomont, 1914 | | | Onthophagus spiculatus Boucomont, 1914 | | | Onthophagus spinifex Fabricius, 1781 | | | Onthophagus spitsbergeniensis Krell, 2010 | | | Onthophagus spitsbergensis ♰ O. Heer, 1870 | | | Onthophagus statzi ♰ Krell, 1990 | | | Onthophagus stellio Erichson, 1843 | | | Onthophagus sternalis Arrow, 1931 | | | Onthophagus sternax Balthasar, 1959 | | | Onthophagus sticticus Harold, 1867 | | | Onthophagus strandi Balthasar, 1935 | | | Onthophagus subcornutus Boucomont, 1914 | | | Onthophagus subocelliger (Blackburn, 1903) | ⇒ Onthophagus waterhousei Gillet, 1927 | | Onthophagus suermelii Petrovitz, 1963 | | | Onthophagus suffusus Klug, 1855 | | | Onthophagus sugihartoi Ochi, 2007 | | | Onthophagus sugillatus Klug, 1855 | | | Onthophagus suginokoichii Ochi & Kon, 2008 | | | Onthophagus suillus Arrow, 1931 | | | Onthophagus sulawesiensis Krikken & Hu..., 2011 | | | Onthophagus sulawesijohkii Ochi, Kon & ..., 2016 | | | Onthophagus sulci Balthasar, 1935 | | | Onthophagus sumatramontanus Ochi & Kon, 2008 | | | Onthophagus sumatranus Lansberge, 1883 | | | Onthophagus sundanensis Lansberge, 1883 | | | Onthophagus surdus Boucomont, 1925 | | | Onthophagus susterai Balthasar, 1952 | | | Onthophagus sutleinensis Splichal, 1910 | | | Onthophagus suturalis (Péringuey, 1888) | ⇒ Onthophagus trituber Wiedemann, 1823 | | Onthophagus sycophanta (Fairmaire, 1887) | ⇒ Euonthophagus gibbosus Scriba, 1790 | | Onthophagus sydneyensis Blackburn, 1903 | | | Onthophagus tabellicornis MacLeay, 1864 | | | Onthophagus tabidus Balthasar, 1935 | | | Onthophagus taeniatus Boucomont, 1914 | | | Onthophagus tagal Boucomont, 1924 | | | Onthophagus takabayashii Matsumura, 1937 | | | Onthophagus takaoi Ochi & Kon, 2014 | | | Onthophagus tambing Krikken & Hu..., 2011 | | | Onthophagus tamworthi Blackburn, 1903 | | | Onthophagus taoi Ochi & Kon, 2006 | | | Onthophagus taprobanus Arrow, 1931 | | | Onthophagus tenebrosus Harold, 1871 | | | Onthophagus tessulatus Harold, 1871 | | | Onthophagus thailaevis Masumoto, Oc..., 2008 | | | Onthophagus thainuaensis Masumoto, Oc..., 2008 | | | Onthophagus tharalithae Karimbumkara..., 2016 | | | Onthophagus thoreyi Harold, 1868 | | | Onthophagus thungphraensis Masumoto, Oc..., 2020 | | | Onthophagus tibetanus Arrow, 1907 | | | Onthophagus timorensis Boucomont, 1914 | | | Onthophagus tiniocelloides Boucomont, 1925 | | | Onthophagus tirapensis Biswas & Cha..., 1985 | | | Onthophagus tnai Nithya & Sabu, 2012 | | | Onthophagus tonkineus Gillet, 1921 | | | Onthophagus tonsus d´Orbigny, 1902 | | | Onthophagus tonywhitteni Krikken & Hu..., 2011 | | | Onthophagus toraut Krikken & Hu..., 2011 | | | Onthophagus traginus Frey, 1972 | | | Onthophagus tragoides Boucomont, 1914 | | | Onthophagus transcaspicus Koenig, 1888 | | | Onthophagus transvestitus Huijbregts &..., 2009 | | | Onthophagus trapezicornis d´Orbigny, 1902 | | | Onthophagus tricavicollis Lea, 1923 | | | Onthophagus triceratops Arrow, 1913 | | | Onthophagus tricolor Boucomont, 1914 | | | Onthophagus tridentitibialis Ochi & Kon, 2008 | | | Onthophagus trinodosus Fahraeus, 1857 | | | Onthophagus trochilus Arrow, 1931 | | | Onthophagus troglodyta Wiedemann, 1823 | | | Onthophagus troniceki Balthasar, 1933 | | | Onthophagus tsubakii Ochi & Kon, 2009 | | | Onthophagus tsutomui Ochi & Kon, 2016 | | | Onthophagus tuckonie Matthews, 1972 | | | Onthophagus tungkamangensis Masumoto, Oc..., 2007 | | | Onthophagus tungkamungensis Masumoto, Oc..., 2008 | | | Onthophagus turbatus (Walker, 1858) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus turfanicus Balthasar, 1971 | | | Onthophagus turpidus Reitter, 1887 | | | Onthophagus tweedensis Blackburn, 1903 | | | Onthophagus ubaidillahi Krikken & Hu..., 2012 | | | Onthophagus uedai Ochi & Kon, 2006 | | | Onthophagus uedanus (Matsumura, 1937) | ⇒ Onthophagus olsoufieffi Boucomont, 1924 | | Onthophagus ulugombakensis Ochi, Kon & ..., 2014 | | | Onthophagus umbilicopunctatus (Medvedev, 1959) | ⇒ Onthophagus trispinus Reitter, 1892 | | Onthophagus unidentatus Boucomont, 1927 | | | Onthophagus unifasciatus Schaller, 1783 | | | Onthophagus uniformis Heyden, 1886 | | | Onthophagus urellus Boucomont, 1919 | | | Onthophagus urus (Harold, 1867) | ⇒ Onthophagus urusheeri Krell, 2000 | | Onthophagus urusheeri ♰ Krell, 2000 | | | Onthophagus ushiodai (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus usurpator Balthasar, 1959 | | | Onthophagus vanasseni Krikken & Hu..., 2008 | | | Onthophagus vanderblomi Krikken & Hu..., 2008 | | | Onthophagus vanofwegeni Krikken & Hu..., 2008 | | | Onthophagus variegatus (Fabricius, 1798) | ⇒ Onthophagus filicornis Harold, 1873 | | Onthophagus variolaris Lansberge, 1883 | | | Onthophagus ventralis Lansberge, 1883 | | | Onthophagus vermiculatus Frey, 1970 | | | Onthophagus verticalis (Fahraeus, 1857) | ⇒ Onthophagus praedatus Harold, 1862 | | Onthophagus vethi Krikken, 1977 | | | Onthophagus vicinus (Harold, 1886) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus vigens Péringuey, 1901 | | | Onthophagus vilis Harold, 1877 | | | Onthophagus vinctus Erichson, 1843 | | | Onthophagus violaceotinctus Gillet, 1925 | | | Onthophagus viridiperakensis Ochi, Kon & ..., 2014 | | | Onthophagus viriditinctus Reitter, 1893 | | | Onthophagus vladimiri Frey, 1957 | | | Onthophagus volucer Balthasar, 1959 | | | Onthophagus vulpes Harold, 1877 | | | Onthophagus vulpinus Arrow, 1931 | | | Onthophagus vultur Arrow, 1931 | | | Onthophagus wakelbura Matthews, 1972 | | | Onthophagus wallacei Harold, 1871 | | | Onthophagus walteri Josso & Prév..., 2000 | | | Onthophagus waminda Matthews, 1972 | | | Onthophagus wangnamkieous Masumoto, Oc..., 2008 | | | Onthophagus wangtaichieni Masumoto, Oc..., 2020 | | | Onthophagus waterstradti Boucomont, 1914 | | | Onthophagus watuwila Krikken & Hu..., 2008 | | | Onthophagus wayaua Huijbregts &..., 2012 | | | Onthophagus weisei (Reitter, 1891) | ⇒ Euonthophagus gibbosus Scriba, 1790 | | Onthophagus wiebesi Krikken, 1977 | | | Onthophagus woroae Ochi & Kon, 2006 | | | Onthophagus woroae bruneiensis Ochi, Kon & ..., 2009 | | | Onthophagus woroae woroae | | | Onthophagus xanthochlorus Walter & Cam..., 1977 | | | Onthophagus yakuinsulanus Nakane, 1984 | | | Onthophagus yangmunensis Masumoto & Ochi, 2015 | | | Onthophagus yanoi Matsumura, 1938 | | | Onthophagus yaoi Masumoto, Oc..., 2014 | | | Onthophagus yaran Storey & Weir, 1990 | | | Onthophagus yasuhikoi Masumoto, Oc..., 2012 | | | Onthophagus yeyeko Matthews, 1972 | | | Onthophagus yifer Krikken & Hu..., 2012 | | | Onthophagus yugianus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus yujii Masumoto, Oc..., 2012 | | | Onthophagus yumigatanus Matsumura, 1937 | | | Onthophagus yumotoi Ochi & Kon, 2006 | | | Onthophagus yungaburra Matthews, 1972 | | | Onthophagus zagrosicus Kabakov, 2006 | | | Onthophagus zavreli Balthasar, 1935 | | | Onthophagus zebra Arrow, 1931 | | | Onthophagus zebu Boucomont, 1921 | | | Onthophagus zimmermanni Balthasar, 1959 | | | Onthophagus zinovskyi Qaryagdy, 1939 | | | Onthophagus zuvandi Qaryagdy, 1939 | | | Onthophagus zymoticus Moxey, 1963 | | | Onychothecus ateuchoides Boucomont, 1912 | | | Onychothecus corniclypeus Zhang, 1980 | | | Onychothecus riekoae Ochi & Kon, 1998 | | | Onychothecus tridentigeris Zelenka, 1992 | | | Pachylomera femoralis Kirby, 1828 | | | Pachysoma aesculapius Olivier, 1789 | | | Pachysoma bennigseni Felsche, 1907 | | | Pachysoma denticolle Péringuey, 1888 | | | Pachysoma gariepinus Ferreira, 1953 | | | Panelus assamensis Arrow, 1907 | | | Panelus bakeri Boucomont, 1921 | | | Panelus besucheti Paulian, 1980 | | | Panelus borneensis Paulian, 1992 | | | Panelus brendelli Ochi, Kon & ..., 2009 | | | Panelus ceylonicus Balthasar, 1972 | | | Panelus crenatus Nomura, 1973 | | | Panelus danumensis Ochi, Kon & ..., 2009 | | | Panelus deuvei Paulian, 1992 | | | Panelus fallax Balthasar, 1972 | | | Panelus fujitai Ochi, Kon & ..., 2008 | | | Panelus horni | | | Panelus imitator Balthasar, 1972 | | | Panelus jaccoudi Paulian, 1980 | | | Panelus kalimantanicus Ochi, Kon & ..., 2009 | | | Panelus karasuyamai Ochi, Kon & ..., 2008 | | | Panelus keralai Paulian, 1980 | | | Panelus kubotai Kawahara, In..., 2007 | | | Panelus maedai Nomura, 1973 | | | Panelus manasi Paulian, 1983 | | | Panelus manmiaoae Masumoto, Oc..., 2011 | | | Panelus mendeli Ochi, Kon & ..., 2009 | | | Panelus minutus Boucomont, 1914 | | | Panelus mussardi Paulian, 1980 | | | Panelus orousseti Paulian, 1981 | | | Panelus ovatus Nomura, 1973 | | | Panelus parvulus Waterhouse, 1874 | | | Panelus pernitidus Balthasar, 1972 | | | Panelus puncticollis Arrow, 1931 | | | Panelus quadridentatus Balthasar, 1952 | | | Panelus rufocuprea | | | Panelus rufulus Nomura, 1973 | | | Panelus sabahi Paulian, 1992 | | | Panelus saigusai Ochi & Kon, 2016 | | | Panelus setosus Arrow, 1931 | | | Panelus sumatranus Lansberge, 1885 | | | Panelus sumatrensis Ochi, Kon & ..., 1998 | | | Panelus tonkinensis Paulian, 1939 | | | Panelus wangi Masumoto, Oc..., 2011 | | | Panelus watanabei Masumoto & Yin, 1993 | | | Panelus whitehousei | | | Panelus wiebesi | | | Parachorius asymmetricus Tarasov, 2017 | | | Parachorius bolavensis Tarasov, 2017 | | | Parachorius fukiensis Balthasar, 1960 | | | Parachorius fungorum Kryzhanovsky..., 1966 | | | Parachorius globosus Arrow, 1931 | | | Parachorius gotoi Masumoto, 1986 | | | Parachorius hookeri Arrow, 1931 | | | Parachorius humeralis Arrow, 1907 | | | Parachorius javanus Boucomont, 1914 | | | Parachorius krali Utsunomiya &..., 2001 | | | Parachorius lannathai Hanboonsong ..., 2001 | | | Parachorius longipenis Tarasov, 2017 | | | Parachorius maruyamai Masumoto, Oc..., 2012 | | | Parachorius matsudai Ochi, Pham &..., 2016 | | | Parachorius newthayerae Tarasov, 2017 | | | Parachorius nudus Sharp, 1875 | | | Parachorius peninsularis Arrow, 1907 | | | Parachorius pseudojavanus Tarasov, 2017 | | | Parachorius schuelkei Tarasov, 2017 | | | Parachorius semsanganus Tarasov & Keith, 2011 | | | Parachorius solodovnikovi Tarasov, 2017 | | | Parachorius thomsoni Harold, 1873 | | | Paragymnopleurus ambiguus Janssens, 1945 | | | Paragymnopleurus brahminus Waterhouse, 1890 | | | Paragymnopleurus martinezi Balthasar, 1955 | | | Paragymnopleurus maurus Sharp, 1875 | | | Paragymnopleurus melanarius Harold, 1867 | | | Paragymnopleurus planus Sharp, 1875 | | | Paragymnopleurus rudis Sharp, 1875 | | | Paragymnopleurus sinuatus Olivier, 1789 | | | Paragymnopleurus sinuatus abax Sharp, 1875 | | | Paragymnopleurus sinuatus assamensis Waterhouse, 1890 | | | Paragymnopleurus sinuatus productus Sharp, 1875 | | | Paragymnopleurus sinuatus sinuatus | | | Paragymnopleurus sinuatus szechouanicus Balthasar, 1934 | | | Paragymnopleurus sparsus Sharp, 1875 | | | Paragymnopleurus sparsus arnoldi Krikken & Hu..., 1987 | | | Paragymnopleurus sparsus javanus Krikken & Hu..., 1987 | | | Paragymnopleurus sparsus palawanicus Ochi & Kon, 1997 | | | Paragymnopleurus sparsus sparsus | | | Paragymnopleurus spinotus Boucomont, 1914 | | | Paragymnopleurus stipes Sharp, 1875 | | | Paragymnopleurus striatus Sharp, 1875 | | | Paragymnopleurus sumatrensis Ochi & Kon, 1997 | | | Paraphytus andamanus Arrow, 1931 | | | Paraphytus andoi Ochi & Kon, 2015 | | | Paraphytus dentifrons Lewis, 1895 | | | Paraphytus doriae Harold, 1877 | | | Paraphytus foveatus Boucomont, 1923 | | | Paraphytus harmandi Boucomont, 1923 | | | Paraphytus hindu Arrow, 1913 | | | Paraphytus ritsemae Harold, 1880 | | | Paraphytus sumatranus Paulian & Ca..., 1981 | | | Paroniticellus festivus Steven, 1809 | | | Peckolus alpinus Howden & Sch..., 1988 | | | Pedaria barrei Josso & Prév..., 2006 | | | Pedaria cylindrica Fahraeus, 1857 | | | Pedaria decorsei (Boucomont, 1922) | ⇒ Pedaria taylori Waterhouse, 1890 | | Pedaria picea Fahraeus, 1857 | | | Pedaria segregis Péringuey, 1901 | | | Phacosoma fallacilaetum (Masumoto, 1989) | ⇒ Ochicanthon thai Paulian, 1987 | | Phacosoma thailandicum (Masumoto, 1989) | ⇒ Ochicanthon thailandicum Masumoto, 1989 | | Phaedotrogus ceylonicus Balthasar, 1972 | | | Phalops ardea Klug, 1855 | | | Phalops boschas Klug, 1855 | | | Phalops bubalus Harold, 1867 | | | Phalops candezei Lansberge, 1883 | | | Phalops chalybeus Klug, 1845 | | | Phalops cyanescens d´Orbigny, 1897 | | | Phalops divisus Wiedemann, 1823 | | | Phalops dregei Harold, 1867 | | | Phalops flavocinctus Klug, 1855 | | | Phalops iphis Olivier, 1789 | | | Phalops olivaceus Lansberge, 1883 | | | Phalops plancus Erichson, 1843 | | | Phalops prasinus (Erichson, 1843) | ⇒ Phalops beccarii Harold, 1875 | | Phalops rufosignatus (Lansberge, 1885) | ⇒ Phalops dregei Harold, 1867 | | Phalops vanellus Lansberge, 1883 | | | Phalops wittei Harold, 1867 | | | Phanaeus (Notiophanaeus) meleagris Blanchard, 1846 | | | Phanaeus (Notiophanaeus) pyrois Bates, 1887 | | | Phanaeus (Phanaeus) amithaon Harold, 1875 | | | Phanaeus (Phanaeus) igneus MacLeay, 1819 | | | Phanaeus (Phanaeus) palliatus Sturm, 1843 | | | Phanaeus (Phanaeus) vindex MacLeay, 1819 | | | Phanaeus daphnis Harold, 1863 | | | Phanaeus demon Castelnau, 1840 | | | Phanaeus menelas (Castelnau, 1840) | ⇒ Sulcophanaeus menelas Castelnau, 1840 | | Phanaeus mexicanus Harold, 1863 | | | Phanaeus minutus (Motschulsky, 1860) | ⇒ Liatongus phanaeoides Westwood, 1839 | | Phanaeus texensis Edmonds, 1994 | | | Ponerotrogus annandalei (Silvestri, 1924) | ⇒ Haroldius annandalai Silvestri, 1924 | | Prionocephale deplanate ♰ Q. B. Lin, 1980 | | | Proagoderus aciculatus Fahraeus, 1857 | | | Proagoderus amplexus Sharp, 1875 | | | Proagoderus badeni Harold, 1875 | | | Proagoderus bicallosus Klug, 1855 | | | Proagoderus gladiator Arrow, 1907 | | | Proagoderus imperator Castelnau, 1840 | | | Proagoderus lanista Castelnau, 1840 | | | Proagoderus loricatus Klug, 1855 | | | Proagoderus mouhoti Harold, 1875 | | | Proagoderus pactolus Fabricius, 1787 | | | Proagoderus panoplus Bates, 1888 | | | Proagoderus quadriarmatus Fairmaire, 1892 | | | Proagoderus rangifer Klug, 1855 | | | Proagoderus rectefurcatus Fairmaire, 1891 | | | Proagoderus sapphirinus (Péringuey, 1901) | ⇒ Onthophagus aureiceps d´Orbigny, 1902 | | Proagoderus schwaneri Snellen van ..., 1864 | | | Proagoderus victor Sharp, 1875 | | | Proagoderus watanabei Ochi & Kon, 2002 | | | Proagoderus worsissa Roth, 1851 | | | Proagoderus yunnanus Boucomont, 1912 | | | Proteroscarabaeus crassus Ponomarenko, 1990 | | | Proteroscarabaeus magnus ♰ G. V. Nikolajev, 1996 | | | Proteroscarabaeus nikritini ♰ G. V. Nikolajev, 1996 | | | Proteroscarabaeus robustus ♰ H. C. Zhang, 1997 | | | Proteroscarabaeus sibiricus Ponomarenko, 1990 | | | Proteroscarabaeus yeni ♰ Grabau, 1923 | | | Pseudocanthon perplexus LeConte, 1847 | | | Pseudocanthon vitraci Fleutiaux & ..., 1889 | | | Pseudocanthon xanthurus Blanchard, 1846 | | | Pycnopanelus rotundus Arrow, 1931 | | | Sarophorus costatus Fahraeus, 1857 | | | Sarophorus tuberculatus Castelnau, 1840 | | | Sauvagesinella becki Paulian, 1934 | | | Sauvagesinella loftyensis Matthews & Weir, 2002 | | | Scarabaeus (Ateuchetus) armeniacus Ménétriés, 1832 | | | Scarabaeus (Ateuchetus) cicatricosus (Lucas, 1846) | ⇒ Scarabaeus palemo Olivier, 1789 | | Scarabaeus (Ateuchetus) laticollis Linnaeus, 1767 | | | Scarabaeus (Ateuchetus) puncticollis (Latreille, 1819) | ⇒ Geotrupes spiniger Marsham, 1802 | | Scarabaeus (Ateuchetus) semipunctatus Fabricius, 1792 | | | Scarabaeus (Ateuchetus) variolosus (Fabricius, 1787) | ⇒ Ateuchetus semipunctatus Fabricius, 1792 | | Scarabaeus (Scarabaeus) babori Balthasar, 1934 | | | Scarabaeus (Scarabaeus) bannuensis Janssens, 1940 | | | Scarabaeus (Scarabaeus) carinatus (Gebler, 1841) | ⇒ Calcodes carinatus Linnaeus, 1758 | | Scarabaeus (Scarabaeus) pius Illiger, 1803 | | | Scarabaeus (Scarabaeus) sacer (Linnaeus, 1758) | ⇒ Scarabaeus pius Illiger, 1803 | | Scarabaeus (Scarabaeus) typhon (Fischer von ..., 1823) | ⇒ Megasoma gyas Herbst, 1785 | | Scarabaeus (Scarabaeus) winkleri Stolfa, 1938 | | | Scarabaeus abbreviatus (Herbst, 1789) | ⇒ Catharsius molossus Linnaeus, 1758 | | Scarabaeus acuticollis Motschulsky, 1849 | | | Scarabaeus aegaeon Drury, 1773 | | | Scarabaeus aegyptiacus Stolfa, 1938 | | | Scarabaeus aegyptiorum Latreille, 1827 | | | Scarabaeus affinis (Sturm, 1800) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Scarabaeus ambiguus (Boheman, 1857) | ⇒ Gnorimus variabilis Linnaeus, 1758 | | Scarabaeus andrewesi Felsche, 1907 | | | Scarabaeus bellicosus (Olivier, 1789) | ⇒ Coprophanaeus bellicosus Olivier, 1789 | | Scarabaeus berbiceus (Herbst, 1789) | ⇒ Catharsius molossus Linnaeus, 1758 | | Scarabaeus caffer Boheman, 1857 | | | Scarabaeus cineta Drury, 1782 | | | Scarabaeus convexus Hausmann, 1807 | | | Scarabaeus cristatus villiersi Baraud, 1973 | | | Scarabaeus deludens zur Strassen, 1961 | | | Scarabaeus devotus Redtenbacher, 1848 | | | Scarabaeus erichsoni Harold, 1867 | | | Scarabaeus galenus Westwood, 1847 | | | Scarabaeus gangeticus Castelnau, 1840 | | | Scarabaeus goryi Castelnau, 1840 | | | Scarabaeus indus (Linnaeus, 1758) | ⇒ Euphoria inda Linnaeus, 1758 | | Scarabaeus irakensis Stolfa, 1938 | | | Scarabaeus isidis Castelnau, 1840 | | | Scarabaeus janus (Olivier, 1789) | ⇒ Catharsius molossus Linnaeus, 1758 | | Scarabaeus krutai Maly & Montr..., 2011 | | | Scarabaeus leei (Donovan, 1798) | ⇒ Gymnopleurus leei Fabricius, 1792 | | Scarabaeus molossus (Linnaeus, 1758) | ⇒ Catharsius molossus Linnaeus, 1758 | | Scarabaeus multidentatum Klug, 1845 | | | Scarabaeus palemo Olivier, 1789 | | | Scarabaeus proboscideus Guérin-Ménev..., 1844 | | | Scarabaeus proserpinae O. Heer, 1862 | | | Scarabaeus pseudogangeticus Moretto, 2023 | | | Scarabaeus rufus (DeGeer, 1778) | ⇒ Rhysothorax rufus Fabricius, 1792 | | Scarabaeus rugosus (Hausmann, 1807) | ⇒ Onthophagus taurus Schreber, 1759 | | Scarabaeus sanctus Fabricius, 1798 | | | Scarabaeus schaefferi (Linnaeus, 1758) | ⇒ Sisyphus schaefferi schaefferi Linnaeus, 1758 | | Scarabaeus sennaariensis Castelnau, 1840 | | | Scarabaeus sinuatus (Olivier, 1789) | ⇒ Paragymnopleurus sinuatus Olivier, 1789 | | Scarabaeus spretus zur Strassen, 1962 | | | Scarabaeus suri Hausmann, 1807 | | | Scarabaeus transcaspicus Stolfa, 1938 | | | Scarabaeus viator Péringuey, 1901 | | | Scarabaeus wilsoni Waterhouse, 1890 | | | Scarabaeus zagrosensis Montreuil, 2006 | | | Scarabaeus zambesianus Péringuey, 1901 | | | Sceliages granulatus Forgie & Gre..., 2002 | | | Scelocopris enertheus ♰ J. F. Zhang, 1989 | | | Scybalocanthon nigriceps Harold, 1868 | | | Scybalocanthon sexspilotus Guérin-Ménev..., 1855 | | | Sinapisoma minuta Boucomont, 1928 | | | Sinodrepanus besucheti Simonis, 1985 | | | Sinodrepanus exsul Sharp, 1875 | | | Sinodrepanus falsus Sharp, 1875 | | | Sinodrepanus rex Boucomont, 1912 | | | Sinodrepanus rosannae Simonis, 1985 | | | Sinodrepanus similis Simonis, 1985 | | | Sinodrepanus thailandicus Ochi, Kon & ..., 2004 | | | Sinodrepanus tsaii Masumoto, Ya..., 2004 | | | Sinodrepanus uenoi Ochi, Kon & ..., 2004 | | | Sisyphus aages Moxey, 1962 | | | Sisyphus araneolus Arrow, 1927 | | | Sisyphus bowringi White, 1844 | | | Sisyphus caffer Boheman, 1857 | | | Sisyphus caffer goryi Harold, 1859 | | | Sisyphus costatus Thunberg, 1818 | | | Sisyphus crispatus Gory, 1833 | | | Sisyphus crispatus hirtus Wiedemann, 1823 | | | Sisyphus denticrus (Fairmaire, 1886) | ⇒ Sisyphus neglectus Gory, 1833 | | Sisyphus fasciculatus Boheman, 1857 | | | Sisyphus fortuitus Péringuey, 1900 | | | Sisyphus gazanus Arrow, 1909 | | | Sisyphus goryi (Harold, 1859) | ⇒ Sisyphus caffer goryi Harold, 1859 | | Sisyphus hirtus (Gory, 1833) | ⇒ Sisyphus crispatus hirtus Wiedemann, 1823 | | Sisyphus impressipennis Lansberge, 1886 | | | Sisyphus indicus Hope, 1831 | | | Sisyphus laoticus Arrow, 1927 | | | Sisyphus longipes Olivier, 1789 | | | Sisyphus maniti Masumoto, 1988 | | | Sisyphus manni Montreuil, 2015 | | | Sisyphus mendicus Arrow, 1931 | | | Sisyphus moltaensis Vazirani, 1966 | | | Sisyphus neglectus Gory, 1833 | | | Sisyphus popovi Kryzhanovski..., 1966 | | | Sisyphus schaefferi Linnaeus, 1758 | Lanbeiniger Pillendreher | | Sisyphus schaefferi boschniaki Fischer von ..., 1823 | | | Sisyphus schaefferi morio Arrow, 1909 | | | Sisyphus schaefferi schaefferi Linnaeus, 1758 | | | Sisyphus seminulum (Gerstaecker, 1871) | ⇒ Sisyphus costatus Thunberg, 1818 | | Sisyphus sordidus (Boheman, 1857) | ⇒ Sisyphus caffer Boheman, 1857 | | Sisyphus spinipes (Thunberg, 1818) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Sisyphus tarantula (Arrow, 1909) | ⇒ Neosisyphus tarantula Arrow, 1909 | | Sisyphus thoracicus Sharp, 1875 | | | Sisyphus thoracicus chaiyaphumensis Hanboonsong ..., 2000 | | | Sisyphus thoracicus thoracicus | | | Stiptopodius patrizii Boucomont, 1923 | | | Sulcophanaeus menelas Castelnau, 1840 | | | Sulcophanaeus velutinus Murray, 1856 | | | Synapsis aquisgranensis Forster, 1869 | | | Synapsis birmanicus Gillet, 1907 | | | Synapsis boonlongi Hanboonsong ..., 1999 | | | Synapsis brahminus Hope, 1831 | | | Synapsis cambeforti (Ochi, Kon & ..., 2008) | ⇒ Synapsis ritsemae Lansberge, 1874 | | Synapsis davidi (Fairmaire, 1878) | ⇒ Synapsis davidis Fairmaire, 1878 | | Synapsis davidis Fairmaire, 1878 | | | Synapsis dickinsoni Hanboonsong ..., 1999 | | | Synapsis gilleti Arrow, 1931 | | | Synapsis horaki Zidek & Pokorny, 2010 | | | Synapsis kiuchii Hanboonsong ..., 1999 | | | Synapsis masumotoi Ochi, 1992 | | | Synapsis naxiorum Kral & Rejsek, 2000 | | | Synapsis ochii Masumoto, 1995 | | | Synapsis ovalis Boucomont, 1920 | | | Synapsis puluongensis Bui & Bonkowski, 2018 | | | Synapsis punctatus Ochi, Kon & ..., 2008 | | | Synapsis ritsemae Lansberge, 1874 | | | Synapsis roslihashimi Ochi, Kon & ..., 2008 | | | Synapsis satoi Ochi & Kon, 2007 | | | Synapsis simplex Sharp, 1875 | | | Synapsis strnadi Kral, 2002 | | | Synapsis thoas (Sharp, 1875) | ⇒ Synapsis ritsemae Lansberge, 1874 | | Synapsis tmolus Fischer von ..., 1821 | | | Synapsis tridens Sharp, 1881 | | | Synapsis vietnamicus Ochi, Kon & ..., 2020 | | | Synapsis yama Gillet, 1911 | | | Synapsis yunnanus Arrow, 1933 | | | Temnoplectron aeneopiceum Matthews, 1974 | | | Temnoplectron arfakensis Ochi, Kon & ..., 2017 | | | Temnoplectron cooki Reid & Storey, 2000 | | | Temnoplectron politulum MacLeay, 1887 | | | Temnoplectron rotundum Westwood, 1841 | | | Temnoplectron waigeoensis Ochi, Kon & ..., 2017 | | | Tesserodon novaehollandiae Fabricius, 1775 | | | Tesserodon seramicum Krikken & Hu..., 2009 | | | Tesserodon waigeoensis Ochi, Kon & ..., 2017 | | | Tibiodrepanus hircus Wiedemann, 1823 | | | Tibiodrepanus kazirangensis Biswas, 1980 | | | Tibiodrepanus setosus Wiedemann, 1823 | | | Tibiodrepanus simplex Kabakov, 2006 | | | Tibiodrepanus sinicus Harold, 1868 | | | Tibiodrepanus sulcicollis Castelnau, 1840 | | | Tiniocellus averyanovi Kabakov, 2014 | | | Tiniocellus dolosus Branco, 2010 | | | Tiniocellus eurypygus Branco, 2010 | | | Tiniocellus imbellis Bates, 1891 | | | Tiniocellus setifer Kraatz, 1895 | | | Tiniocellus spinipes Roth, 1851 | | | Uroxys boneti Pereira & Ha..., 1961 | | | Uroxys micros Bates, 1887 | | | Uroxys pauliani Balthasar, 1940 | | | Uroxys tuberculatus Lansberge, 1874 | | | Xinidium dentilabris Harold, 1869 | | | Yvescambefortius sarawacus Gillet, 1926 | |
Unterknoten von Scarabaeinae mit Vorkommen in Eurasien | Tribus | Ateuchini
Montreuil, 1998
| 11
37 | Gattungen
Arten |
Taxom
| | Tribus | Canthonini
Péringuey, 1900
| 31
158 | Gattungen
Arten |
Taxom
| | Tribus | Coprini
Kolbe, 1805
| 14
328 | Gattungen
Arten |
Taxom
| | Tribus | Deltochilini
Lacordaire, 1856
| 2
12 | Gattungen
Arten |
Taxom
| | Tribus | Gymnopleurini
Lacordaire, 1856
| 4
81 | Gattungen
Arten |
Taxom
| | Tribus | Oniticellini
d´Orbigny, 1916
| 15
113 | Gattungen
Arten |
Taxom
| | Tribus | Onitini
Hope, 1837
| 4
84 | Gattungen
Arten |
Taxom
| | Tribus | Onthophagini
Lacordaire, 1856
Kotkäfer | 17
1469 | Gattungen
Arten |
Taxom
| | Tribus | Phanaeini
MacLeay, 1819
| 5
23 | Gattungen
Arten |
Taxom
| | Tribus | Scarabaeini
Latreille, 1802
| 7
75 | Gattungen
Arten |
Taxom
| | Tribus | Sisyphini
Mulsant, 1842
| 2
37 | Gattungen
Arten |
Taxom
|
Links, Quellen und weitere Informationen Scarabaeinae bei artfakta.se
Scarabaeinae bei dyntaxa.se
Scarabaeinae bei itis.gov
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