|
|
| | | | Checkliste der Gattung Onthophagus Latreille, 1802 (Kotfresser)
für Ecoregionen
Klasse
| Insecta
Linnaeus, 1758
Freikiefler, Insekten, Kerbtiere, Kerfe | 814
32225
411216 | Familien
Gattungen
Arten |
Taxom
|
Unterklasse
| Pterygota
Lang, 1888
Fluginsekten | 809
32196
411162 | Familien
Gattungen
Arten |
Taxom
|
Teilklasse
| Neoptera
van der Wulp, 1890
Neuflügler | 776
31715
407401 | Familien
Gattungen
Arten |
Taxom
|
Überordnung
| Holometabola
Heider, 1889
Holometabolen Insekten | 508
27635
383586 | Familien
Gattungen
Arten |
Taxom
|
Ordnung
| Coleoptera
Linnaeus, 1758
Käfer | 152
13619
245491 | Familien
Gattungen
Arten |
Taxom
|
Unterordnung
| Polyphaga
Emery, 1886
| 138
11890
200776 | Familien
Gattungen
Arten |
Taxom
|
Teilordnung
| Scarabaeiformia
Crowson, 1960
| 18
2639
43663 | Familien
Gattungen
Arten |
Taxom
|
Überfamilie
| Scarabaeoidea
Latreille, 1802
Blatthornkäfer | 18
2639
43663 | Familien
Gattungen
Arten |
Taxom
| | Familie | Scarabaeidae
Latreille, 1802
Blatthornkäfer | 2300
39341 | Gattungen
Arten |
Taxom
| | Unterfamilie | Scarabaeinae
Latreille, 1802
Pillendreher | 226
6709 | Gattungen
Arten |
Taxom
| | Tribus | Onthophagini
Lacordaire, 1856
Kotkäfer | 33
3020 | Gattungen
Arten |
Taxom
| | Gattung | Onthophagus
Latreille, 1802
Kotfresser | 2402 | Arten |
Taxom
|
Gattungen für Ecoregionen
1 Gattung mit 2402 Arten
Artenliste für Ecoregionen
2402 Arten und 75 Unterarten | Onthophagus (Afrostrandius) adspersus d´Orbigny, 1908 | | | Onthophagus (Afrostrandius) bimetallicus d´Orbigny, 1907 | | | Onthophagus (Afrostrandius) demeyeri Moretto, 2008 | | | Onthophagus (Afrostrandius) granulipennis (Lansberge, 1886) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus (Afrostrandius) loxodontaphilus Moretto, 2008 | | | Onthophagus (Afrostrandius) plebejus Klug, 1855 | | | Onthophagus (Afrostrandius) pseudoplebejus Moretto, 2008 | | | Onthophagus (Afrostrandius) tigrinus (d´Orbigny, 1908) | ⇒ Onthophagus pygmaeus Schaller, 1783 | | Onthophagus tigrinus kageraensis Moretto, 2009 | | | Onthophagus tigrinus tigrinus | | | Onthophagus (Altonthophagus) cervenkai Kabakov, 2008 | | | Onthophagus (Altonthophagus) inelegans Balthasar, 1935 | | | Onthophagus (Altonthophagus) kozlovi Kabakov, 1990 | | | Onthophagus (Altonthophagus) kukunorensis Kabakov, 1990 | | | Onthophagus (Altonthophagus) marmotae Kabakov, 1990 | | | Onthophagus (Altonthophagus) sibiricus Harold, 1877 | | | Onthophagus (Altonthophagus) turpidoides Kabakov, 2008 | | | Onthophagus (Amphionthophagus) falzonii Goidanich, 1926 | | | Onthophagus (Amphionthophagus) melitaeus Fabricius, 1798 | | | Onthophagus (Colobonthophagus) aenescens Wiedemann, 1823 | | | Onthophagus (Colobonthophagus) agnus Gillet, 1925 | | | Onthophagus (Colobonthophagus) arunensis Scheuern, 1995 | | | Onthophagus (Colobonthophagus) caprai Frey, 1956 | | | Onthophagus (Colobonthophagus) dama Fabricius, 1798 | | | Onthophagus (Colobonthophagus) metalliceps Arrow, 1931 | | | Onthophagus (Colobonthophagus) neocolobus Scheuern, 1996 | | | Onthophagus (Colobonthophagus) occipitalis Lansberge, 1885 | | | Onthophagus (Colobonthophagus) paliceps Arrow, 1931 | | | Onthophagus (Colobonthophagus) piceorufulus Kabakov, 1994 | | | Onthophagus (Colobonthophagus) poggii Scheuern, 1996 | | | Onthophagus (Colobonthophagus) quadridentatus (Fabricius, 1798) | ⇒ Onthophagus fissicornis Stevens, 1809 | | Onthophagus (Colobonthophagus) shillongensis Scheuern, 1995 | | | Onthophagus (Colobonthophagus) thai Kabakov, 1994 | | | Onthophagus (Colobonthophagus) tragus Fabricius, 1792 | | | Onthophagus (Colobonthophagus) transquadridentatus Scheuern, 1995 | | | Onthophagus (Euonthophagus) aeneobrunneus Kabakov, 1977 | | | Onthophagus (Euonthophagus) amyntas (Olivier, 1789) | ⇒ Euonthophagus amyntas Olivier, 1789 | | Onthophagus amyntas auchenia Redtenbacher, 1850 | | | Onthophagus (Euonthophagus) carbonarius Klug, 1855 | | | Onthophagus (Euonthophagus) conterminus Petrovitz, 1971 | | | Onthophagus (Furconthophagus) amicus Gillet, 1925 | | | Onthophagus (Furconthophagus) boucomonti Paulian, 1931 | | | Onthophagus (Furconthophagus) furcatus (Fabricius, 1781) | ⇒ Onthophagus furcaticeps Masters, 1886 | | Onthophagus (Furconthophagus) karenensis Masumoto, Oc..., 2008 | | | Onthophagus (Furconthophagus) khonkaenus Masumoto, Oc..., 2008 | | | Onthophagus (Furconthophagus) lilliputanus Lansberge, 1883 | | | Onthophagus (Furconthophagus) monforti Josso & Prév..., 2006 | | | Onthophagus (Furconthophagus) papulatorius Kabakov, 2008 | | | Onthophagus (Furconthophagus) papulatus Boucomont, 1914 | | | Onthophagus (Furconthophagus) promontorii Boucomont, 1924 | | | Onthophagus (Furconthophagus) schoolmeestersi Ochi & Kon, 2007 | | | Onthophagus (Gibbonthophagus) apicetinctus (d´Orbigny, 1900) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus (Gibbonthophagus) atripennis Waterhouse, 1875 | | | Onthophagus (Gibbonthophagus) chineicus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) denticollis Lansberge, 1883 | | | Onthophagus (Gibbonthophagus) duporti Boucomont, 1914 | | | Onthophagus (Gibbonthophagus) euryceros Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) fujiii Ochi & Kon, 1995 | | | Onthophagus (Gibbonthophagus) gracilipes Boucomont, 1914 | | | Onthophagus (Gibbonthophagus) hiabunicus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) kentingensis Nomura, 1973 | | | Onthophagus (Gibbonthophagus) kimioi Ochi, Kon & ..., 2011 | | | Onthophagus (Gibbonthophagus) lenis Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) limbatus Herbst, 1789 | | | Onthophagus (Gibbonthophagus) luridipennis Boheman, 1858 | | | Onthophagus (Gibbonthophagus) nigriobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) palawanicus Ochi & Kon, 2004 | | | Onthophagus (Gibbonthophagus) parviobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) penmani Masumoto, Oc..., 2002 | | | Onthophagus (Gibbonthophagus) privus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) rufiobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) sasajii Ochi & Kon, 2001 | | | Onthophagus (Gibbonthophagus) semipersonatus Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) sunantaae Masumoto, 1989 | | | Onthophagus (Gibbonthophagus) taurinus White, 1844 | | | Onthophagus (Gibbonthophagus) viduus (Harold, 1875) | ⇒ Onthophagus orbus Boucomont, 1924 | | Onthophagus (Gibbonthophagus) viridicervicapra Ochi, Kon & ..., 2009 | | | Onthophagus (Hyalonthophagus) hyalcyon Palestrini &..., 1989 | | | Onthophagus (Indachorius) arai Masumoto, 1989 | | | Onthophagus (Indachorius) baykanus Kabakov, 1994 | | | Onthophagus (Indachorius) bonengus Kabakov, 1994 | | | Onthophagus (Indachorius) chinensis Balthasar, 1952 | | | Onthophagus (Indachorius) danumensis (Ochi, Kon & ..., 2009) | ⇒ Onthophagus danumicus Huijbregts &..., 2012 | | Onthophagus (Indachorius) doisuthepensis Masumoto, 1989 | | | Onthophagus (Indachorius) hsui Masumoto, Ch..., 2004 | | | Onthophagus (Indachorius) koshunensis Balthasar, 1941 | | | Onthophagus (Indachorius) lannamiibun Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) maesaensis Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) masaoi Ochi, 1992 | | | Onthophagus (Indachorius) mendeli Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) mongkhoni Masumoto, Ha..., 2002 | | | Onthophagus (Indachorius) nongkaiensis Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) octonaevus Kabakov, 1994 | | | Onthophagus (Indachorius) paramasaoi Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) phetchabunensis Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) phuquoci Paulian, 1945 | | | Onthophagus (Indachorius) platypus Zhang, 1997 | | | Onthophagus (Indachorius) pseudoworoae Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) scotti Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) semidanumensis Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) subsapaensis Kabakov, 1994 | | | Onthophagus (Indachorius) suginoi Ochi, 1984 | | | Onthophagus (Indachorius) tiamicus Kabakov, 1994 | | | Onthophagus (Indachorius) tongbantumi Masumoto, Oc..., 2002 | | | Onthophagus (Macronthophagus) cludtsi Ochi, 2003 | | | Onthophagus (Macronthophagus) curvicarinatus Boucomont, 1914 | | | Onthophagus (Macronthophagus) diabolicus Harold, 1877 | | | Onthophagus (Macronthophagus) manipurensis Arrow, 1907 | | | Onthophagus (Macronthophagus) menieri Ochi, 2003 | | | Onthophagus (Macronthophagus) nilgirensis Gillet, 1922 | | | Onthophagus (Macronthophagus) rubricollis Hope, 1831 | | | Onthophagus (Macronthophagus) uenoi Ochi, 1995 | | | Onthophagus uenoi borneomontanus Ochi, 2003 | | | Onthophagus uenoi uenoi Ochi, 1995 | | | Onthophagus (Matashia) anguliceps (Boucomont, 1914) | ⇒ Onthophagus avocetta Arrow, 1933 | | Onthophagus (Matashia) yubarinus (Matsumura, 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus (Micronthophagus) falsivigilans Masumoto, 1995 | | | Onthophagus (Micronthophagus) gigantivigilans Masumoto, Ha..., 2002 | | | Onthophagus (Micronthophagus) konoi Matsumura, 1938 | | | Onthophagus (Micronthophagus) quaestus Sharp, 1875 | | | Onthophagus (Micronthophagus) vigilans Boucomont, 1921 | | | Onthophagus (Micronthophagus) wangi Masumoto, Ch..., 2004 | | | Onthophagus (Onthophagiellus) kondaoensis Kabakov, 1994 | | | Onthophagus (Onthophagiellus) opacifalculatus Ochi, Kon & ..., 2009 | | | Onthophagus (Onthophagiellus) parafalculatus Ochi, Kon & ..., 2009 | | | Onthophagus (Onthophagiellus) popovi Kabakov, 1983 | | | Onthophagus (Onthophagiellus) taiyaruensis Masumoto, 1977 | | | Onthophagus (Onthophagiellus) thanwaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagiellus) unguiculatus Kabakov, 1994 | | | Onthophagus (Onthophagus) abacus Boucomont, 1921 | | | Onthophagus (Onthophagus) aciculatulus Blatchley, 1928 | | | Onthophagus (Onthophagus) acuticollis (Gillet, 1927) | ⇒ Onthophagus amphioxus Arrow, 1933 | | Onthophagus (Onthophagus) acuticornis Endrödi, 1973 | | | Onthophagus (Onthophagus) alluvius Howden and C..., 1963 | | | Onthophagus (Onthophagus) alternans (Raffray, 1877) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus (Onthophagus) amirus Kabakov, 1982 | | | Onthophagus (Onthophagus) anguicorius Boucomont, 1914 | | | Onthophagus (Onthophagus) angustatus Boucomont, 1914 | | | Onthophagus (Onthophagus) apilularius Masumoto, 1995 | | | Onthophagus (Onthophagus) arnetti Howden and C..., 1963 | | | Onthophagus (Onthophagus) arnoldii Kabakov, 1982 | | | Onthophagus (Onthophagus) barbieri Paulian, 1978 | | | Onthophagus (Onthophagus) batesi Howden and C..., 1963 | | | Onthophagus (Onthophagus) bokiaunus Masumoto, 1995 | | | Onthophagus (Onthophagus) brevifrons (Horn, 1881) | ⇒ Onthophagus abyssinicus Gillet, 1925 | | Onthophagus (Onthophagus) browni Howden and C..., 1963 | | | Onthophagus (Onthophagus) canelasensis Howden and G..., 2004 | | | Onthophagus (Onthophagus) cartwrighti Howden, 1973 | | | Onthophagus (Onthophagus) cavernicollis Howden and C..., 1963 | | | Onthophagus (Onthophagus) chaiyaphumensis Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) chevrolati Harold, 1869 | | | Onthophagus chevrolati longecarinatus Zunino & Hal..., 1988 | | | Onthophagus chevrolati omiltemius Bates, 1887 | | | Onthophagus chevrolati retusus Harold, 1869 | | | Onthophagus (Onthophagus) coahuilae Zunino and H..., 1988 | | | Onthophagus (Onthophagus) cochisus Brown, 1927 | | | Onthophagus (Onthophagus) concinnus (Laporte, 1840) | ⇒ Onthophagus striatulus Palisot de B..., 1809 | | Onthophagus (Onthophagus) coproides Horn, 1881 | | | Onthophagus (Onthophagus) cuevensis Howden, 1973 | | | Onthophagus (Onthophagus) cyanellus Bates, 1887 | | | Onthophagus (Onthophagus) cynomysi Brown, 1927 | | | Onthophagus (Onthophagus) dapcauensis Boucomont, 1921 | | | Onthophagus (Onthophagus) depressus Harold, 1871 | | | Onthophagus (Onthophagus) doiinthanonensis Masumoto, 1989 | | | Onthophagus (Onthophagus) doipuiensis Masumoto, 1989 | | | Onthophagus (Onthophagus) doitungensis Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) dubernardi Boucomont, 1914 | | | Onthophagus (Onthophagus) embersoni Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) finschi Harold, 1877 | | | Onthophagus (Onthophagus) fissicornis Stevens, 1809 | | | Onthophagus (Onthophagus) formosanus Gillet, 1924 | | | Onthophagus (Onthophagus) fuscus Boucomont, 1932 | | | Onthophagus fuscus canescens Zunino & Hal..., 1988 | | | Onthophagus fuscus mycetorum Zunino & Hal..., 1988 | | | Onthophagus fuscus parafuscus Zunino & Hal..., 2005 | | | Onthophagus (Onthophagus) gibbulus Pallas, 1781 | | | Onthophagus (Onthophagus) gorochovi Kabakov, 1998 | | | Onthophagus (Onthophagus) grandivigilans Masumoto, 1995 | | | Onthophagus (Onthophagus) hajimei Masumoto, 1984 | | | Onthophagus (Onthophagus) hastifer Lansberge, 1885 | | | Onthophagus (Onthophagus) hayashii Masumoto, 1991 | | | Onthophagus (Onthophagus) hecate Panzer, 1794 | | | Onthophagus hecate blatchleyi Brown, 1929 | | | Onthophagus hecate hecate Panzer, 1794 | | | Onthophagus (Onthophagus) hidalgus Zunino and H..., 1988 | | | Onthophagus (Onthophagus) hoepfneri Harold, 1869 | | | Onthophagus (Onthophagus) horii Ochi, Kon & ..., 2009 | | | Onthophagus (Onthophagus) illyricus Scopoli, 1763 | | | Onthophagus (Onthophagus) incensus Say, 1835 | | | Onthophagus (Onthophagus) inermifrons (d´Orbigny, 1902) | ⇒ Onthophagus margaritifer d´Orbigny, 1898 | | Onthophagus (Onthophagus) ishiii Ochi & Kon, 1995 | | | Onthophagus (Onthophagus) ivae Balthasar, 1963 | | | Onthophagus (Onthophagus) javaecola Balthasar, 1959 | | | Onthophagus (Onthophagus) kanyaayonus Masumoto, 1992 | | | Onthophagus (Onthophagus) kaszabi (Balthasar, 1963) | ⇒ Onthophagus vitulus Péringuey, 1901 | | Onthophagus (Onthophagus) khonmiinitnoi Masumoto, 1990 | | | Onthophagus (Onthophagus) kinhthaicus Kabakov, 1998 | | | Onthophagus (Onthophagus) kiuchii Masumoto, 1995 | | | Onthophagus (Onthophagus) knausi Brown, 1927 | | | Onthophagus (Onthophagus) knulli Howden and C..., 1963 | | | Onthophagus (Onthophagus) koryoensis Kim, 1985 | | | Onthophagus (Onthophagus) krakadaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagus) kryzhanovskii (Kabakov, 1982) | ⇒ Onthophagus zuvandi Qaryagdy, 1939 | | Onthophagus (Onthophagus) kuatunensis Balthasar, 1942 | | | Onthophagus (Onthophagus) landolti (Harold, 1880) | ⇒ Onthophagus schaefferi Howden and C..., 1963 | | Onthophagus landolti landolti Harold, 1880 | | | Onthophagus landolti texanus Schaeffer, 1914 | | | Onthophagus (Onthophagus) lecontei (Harold, 1871) | ⇒ Onthophagus subopacus Robinson, 1940 | | Onthophagus (Onthophagus) lindaae Masumoto, 1989 | | | Onthophagus (Onthophagus) lutosopictus Fairmaire, 1897 | | | Onthophagus (Onthophagus) maleengnaafon Masumoto, 1990 | | | Onthophagus (Onthophagus) maleengnoi Masumoto, 1990 | | | Onthophagus (Onthophagus) maniti Masumoto, 1989 | | | Onthophagus (Onthophagus) margaritifer d´Orbigny, 1898 | | | Onthophagus (Onthophagus) masumotoi Ochi, 1985 | | | Onthophagus (Onthophagus) matsudai Ochi & Kon, 2004 | | | Onthophagus (Onthophagus) mcclevei Howden and G..., 2004 | | | Onthophagus (Onthophagus) medorensis Brown, 1929 | | | Onthophagus (Onthophagus) mendicus Gillet, 1924 | | | Onthophagus (Onthophagus) mextexus (Howden and C..., 1970) | ⇒ Onthophagus durangoensis Balthasar, 1939 | | Onthophagus (Onthophagus) mulleri van Lansberge, 1883 | | | Onthophagus (Onthophagus) naaroon Masumoto, 1990 | | | Onthophagus (Onthophagus) namnaoensis Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) nasonis Kabakov, 1998 | | | Onthophagus (Onthophagus) nuchicornis Linnaeus, 1758 | | | Onthophagus (Onthophagus) nurestanicus Kabakov, 1982 | | | Onthophagus (Onthophagus) oklahomensis Brown, 1927 | | | Onthophagus (Onthophagus) orientalis (Harold, 1868) | ⇒ Onthophagus fissinasus Fairmaire, 1895 | | Onthophagus (Onthophagus) orpheus Panzer, 1794 | | | Onthophagus orpheus canadensis Fabricius, 1801 | | | Onthophagus orpheus orpheus Panzer, 1794 | | | Onthophagus orpheus pseudorpheus Howden & Car..., 1963 | | | Onthophagus (Onthophagus) paraprotzenkoi (Tesar, 1967) | ⇒ Onthophagus finschi Harold, 1877 | | Onthophagus (Onthophagus) penicillatus (Harold, 1879) | ⇒ Onthophagus olsoufieffi Boucomont, 1924 | | Onthophagus (Onthophagus) pennsylvanicus Harold, 1871 | | | Onthophagus (Onthophagus) phrutsaphaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagus) poephagus Kabakov, 1979 | | | Onthophagus (Onthophagus) polyphemi Hubbard, 1894 | | | Onthophagus polyphemi polyphemi Hubbard, 1894 | | | Onthophagus polyphemi sparsisetosus Howden & Car..., 1963 | | | Onthophagus (Onthophagus) proletarius Harold, 1875 | | | Onthophagus (Onthophagus) protzenkoi (Tesar, 1967) | ⇒ Onthophagus finschi Harold, 1877 | | Onthophagus (Onthophagus) pseudoarmatus Balthasar, 1944 | | | Onthophagus (Onthophagus) pseudofuscus Zunino and H..., 1988 | | | Onthophagus (Onthophagus) pseudohystrix Masumoto, 1995 | | | Onthophagus (Onthophagus) punctator Reitter, 1893 | | | Onthophagus (Onthophagus) punneeae Masumoto, 1989 | | | Onthophagus (Onthophagus) ratchasimaensis Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) rechingeri (Petrovitz, 1980) | ⇒ Onthophagus zuvandi Qaryagdy, 1939 | | Onthophagus (Onthophagus) roubali Balthasar, 1935 | | | Onthophagus (Onthophagus) rufimanus Kabakov, 1982 | | | Onthophagus (Onthophagus) rugulosus Harold, 1886 | | | Onthophagus (Onthophagus) sakaeratensis Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) sakainoi Masumoto, 1991 | | | Onthophagus (Onthophagus) salvazai Paulian, 1978 | | | Onthophagus (Onthophagus) sauteri Gillet, 1924 | | | Onthophagus (Onthophagus) schaefferi Howden and C..., 1963 | | | Onthophagus (Onthophagus) seabrai Gomes Alves, 1944 | | | Onthophagus (Onthophagus) setchan Masumoto, 1984 | | | Onthophagus (Onthophagus) singhaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagus) sobrius Balthasar, 1959 | | | Onthophagus (Onthophagus) striatulus Palisot de B..., 1809 | | | Onthophagus striatulus floridanus Blatchley, 1928 | | | Onthophagus striatulus striatulus Palisot de B..., 1809 | | | Onthophagus (Onthophagus) subaeneus (Palisot de B..., 1811) | ⇒ Onthophagus striatulus Palisot de B..., 1809 | | Onthophagus (Onthophagus) subalternans d´Orbigny, 1902 | | | Onthophagus (Onthophagus) subopacus Robinson, 1940 | | | Onthophagus (Onthophagus) subtropicus Howden and C..., 1963 | | | Onthophagus (Onthophagus) taayai Masumoto, 1995 | | | Onthophagus (Onthophagus) tatsienluensis Balthasar, 1942 | | | Onthophagus (Onthophagus) taurus Schreber, 1759 | Oechslein | | Onthophagus (Onthophagus) terminatus Eschscholtz, 1822 | | | Onthophagus (Onthophagus) tholaayi Masumoto, 1990 | | | Onthophagus (Onthophagus) tritinctus Boucomont, 1914 | | | Onthophagus (Onthophagus) trituber Wiedemann, 1823 | | | Onthophagus trituber jacobsoni Kabakov, 2006 | | | Onthophagus (Onthophagus) truncaticornis (Schaller, 1783) | ⇒ Onthophagus quadrinodosus Fahraeus, 1857 | | Onthophagus (Onthophagus) tuberculifrons Harold, 1871 | | | Onthophagus (Onthophagus) tumami Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) velutinus Horn, 1875 | | | Onthophagus (Onthophagus) vitulus Péringuey, 1901 | | | Onthophagus (Onthophagus) wangnamkhieoensis Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) yamaokai Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) yangi Masumoto, Ts..., 2006 | | | Onthophagus (Palaeonthophagus) afghanus Petrovitz, 1961 | | | Onthophagus (Palaeonthophagus) albarracinus Baraud, 1979 | | | Onthophagus (Palaeonthophagus) angorensis Petrovitz, 1963 | | | Onthophagus (Palaeonthophagus) baraudi Nicolas, 1964 | | | Onthophagus (Palaeonthophagus) bytinskii Balthasar, 1960 | | | Onthophagus (Palaeonthophagus) clitellifer Reitter, 1894 | | | Onthophagus (Palaeonthophagus) coenobita Herbst, 1783 | Mönchs-Kotkäfer | | Onthophagus (Palaeonthophagus) conspersus Reitter, 1893 | | | Onthophagus (Palaeonthophagus) cruciatus (Ménétriés, 1832) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus (Palaeonthophagus) dacatrai Pittino, 2004 | | | Onthophagus (Palaeonthophagus) dellacasai Pittino & Ma..., 1981 | | | Onthophagus (Palaeonthophagus) eulaminicornis Pittino, 2006 | | | Onthophagus (Palaeonthophagus) excisus Reiche & Saulcy, 1856 | | | Onthophagus (Palaeonthophagus) excubitor Ziani & Gudenzi, 2006 | | | Onthophagus (Palaeonthophagus) finschi Harold, 1877 | | | Onthophagus (Palaeonthophagus) fissicornis Steven, 1809 | | | Onthophagus (Palaeonthophagus) fortigibber Reitter, 1909 | | | Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790) | ⇒ Onthophagus similis Scriba, 1790 | | Onthophagus (Palaeonthophagus) furciceps Marseul, 1869 | | | Onthophagus (Palaeonthophagus) gibbulus Pallas, 1781 | | | Onthophagus gibbulus gibbulus | | | Onthophagus gibbulus rostrifer Reitter, 1892 | | | Onthophagus (Palaeonthophagus) gorodinskii Kabakov, 2008 | | | Onthophagus (Palaeonthophagus) grossepunctatus Reitter, 1905 | | | Onthophagus (Palaeonthophagus) haroldi Ballion, 1870 | | | Onthophagus (Palaeonthophagus) hissariensis Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) isikdagensis Pittino, 2004 | | | Onthophagus (Palaeonthophagus) joannae Goljan, 1953 | | | Onthophagus (Palaeonthophagus) kindermanni Harold, 1877 | | | Onthophagus (Palaeonthophagus) kolenatii Reitter, 1893 | | | Onthophagus (Palaeonthophagus) laticornis Gebler, 1823 | | | Onthophagus (Palaeonthophagus) latigena d´Orbigny, 1897 | | | Onthophagus (Palaeonthophagus) lemur Fabricius, 1781 | | | Onthophagus (Palaeonthophagus) leucostigma Steven, 1811 | | | Onthophagus (Palaeonthophagus) lucidus Illiger, 1800 | | | Onthophagus (Palaeonthophagus) macedonicus Miksic, 1959 | | | Onthophagus (Palaeonthophagus) marginalis Gebler, 1817 | | | Onthophagus marginalis marginalis | | | Onthophagus marginalis przewalski Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) massai Baraud, 1975 | | | Onthophagus (Palaeonthophagus) medius (Kugelann, 1792) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus (Palaeonthophagus) medvedevi Kabakov, 1982 | | | Onthophagus (Palaeonthophagus) merdarius Chevrolat, 1865 | | | Onthophagus (Palaeonthophagus) muelleri (Lansberge, 1883) | ⇒ Onthophagus statzi Krell, 1990 | | Onthophagus (Palaeonthophagus) nikolajevi Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) novaki Boucomont, 1927 | | | Onthophagus (Palaeonthophagus) nuchicornis Linnaeus, 1758 | Kleine Nackenhorn | | Onthophagus (Palaeonthophagus) olsoufieffi Boucomont, 1924 | | | Onthophagus (Palaeonthophagus) opacicollis Reitter, 1893 | | | Onthophagus (Palaeonthophagus) ovatus Linnaeus, 1767 | Eiförmiger Kotkäfer | | Onthophagus (Palaeonthophagus) panici Petrovitz, 1964 | | | Onthophagus (Palaeonthophagus) parmatus Reitter, 1892 | | | Onthophagus (Palaeonthophagus) petrovitzi (Frey, 1975) | ⇒ Onthophagus petrovitzianus Pittino, 2006 | | Onthophagus (Palaeonthophagus) petrovitzianus Pittino, 2006 | | | Onthophagus (Palaeonthophagus) ponticus Harold, 1883 | | | Onthophagus (Palaeonthophagus) pseudocaccobius Reitter, 1889 | | | Onthophagus (Palaeonthophagus) psychopompus Ziani & Ghar..., 2010 | | | Onthophagus (Palaeonthophagus) pygargus Motschulsky, 1845 | | | Onthophagus (Palaeonthophagus) quadrinodus Reitter, 1896 | | | Onthophagus (Palaeonthophagus) rachelis Martín Piera, 1985 | | | Onthophagus (Palaeonthophagus) rakovici Pittino, 2004 | | | Onthophagus (Palaeonthophagus) rodentium Pittino, 2004 | | | Onthophagus (Palaeonthophagus) ruficapillus Brullé, 1832 | | | Onthophagus ruficapillus guilanensis (Pittino, 1982) | ⇒ Onthophagus ruficapillus Brullé, 1832 | | Onthophagus ruficapillus ruficapillus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) samai Ziani, 2011 | | | Onthophagus (Palaeonthophagus) scabriusculus Harold, 1873 | | | Onthophagus (Palaeonthophagus) semicornis Panzer, 1798 | Halbhörniger Kotkäfer | | Onthophagus (Palaeonthophagus) sericatus Reitter, 1893 | | | Onthophagus (Palaeonthophagus) shapovalovi Gusakov, 2012 | | | Onthophagus (Palaeonthophagus) similis (Scriba, 1790) | ⇒ Onthophagus fracticornis Preyssler, 1790 | | Onthophagus (Palaeonthophagus) speculifer Solsky, 1876 | | | Onthophagus (Palaeonthophagus) strabo Reitter, 1892 | | | Onthophagus (Palaeonthophagus) stylocerus Graells, 1851 | | | Onthophagus (Palaeonthophagus) suturellus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) tesquorum Semenov & Me..., 1927 | | | Onthophagus (Palaeonthophagus) trigibber Reitter, 1893 | | | Onthophagus (Palaeonthophagus) trispinus Reitter, 1892 | | | Onthophagus (Palaeonthophagus) truchmenus Kolenati, 1846 | | | Onthophagus truchmenus iranicus Kabakov, 2006 | | | Onthophagus truchmenus truchmenus | | | Onthophagus truchmenus turkmeniacus Kabakov, 2006 | | | Onthophagus truchmenus xerxes Petrovitz, 1965 | | | Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767 | Kühlein | | Onthophagus (Palaeonthophagus) verticicornis (Laicharting, 1781) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus (Palaeonthophagus) viridis Ménétriés, 1832 | | | Onthophagus (Palaeonthophagus) vitulus (Fabricius, 1776) | ⇒ Onthophagus immundus Boheman, 1858 | | Onthophagus (Palaeonthophagus) vlasovi Medvedev, 1958 | | | Onthophagus (Palaeonthophagus) zuninoi Martín Piera, 1985 | | | Onthophagus (Paleonthophagus) andalusicus Waltl, 1835 | | | Onthophagus andalusicus andalusicus | | | Onthophagus andalusicus persianus Olsoufieff, 1900 | | | Onthophagus (Paleonthophagus) fracticornis Preyssler, 1790 | | | Onthophagus (Paleonthophagus) nuchicornis Linnaeus, 1758 | | | Onthophagus (Paleonthophagus) similis Scriba, 1790 | | | Onthophagus (Paleonthophagus) vacca Linnaeus, 1767 | | | Onthophagus (Paraphanaeomorphus) argyropygus Gillet, 1927 | | | Onthophagus (Paraphanaeomorphus) carinensis Boucomont, 1914 | | | Onthophagus (Paraphanaeomorphus) comottoides Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) insignicollis Frey, 1954 | | | Onthophagus (Paraphanaeomorphus) janushevi Kabakov, 1983 | | | Onthophagus (Paraphanaeomorphus) napolovi Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) pseudojavanus Paulian, 1931 | | | Onthophagus (Paraphanaeomorphus) quadricolor Kabakov, 1983 | | | Onthophagus (Paraphanaeomorphus) semipiceus Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) strnadi Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) vaulogeri Boucomont, 1923 | | | Onthophagus (Parascatonomus) carnarius Nomura, 1976 | | | Onthophagus (Parascatonomus) discedens Sharp, 1875 | | | Onthophagus (Parascatonomus) laotianus Boucomont, 1919 | | | Onthophagus (Parascatonomus) miyakei Ochi & Araya, 1992 | | | Onthophagus (Parascatonomus) miyakoinsularis Ochi, Y. Miy..., 1999 | | | Onthophagus (Parascatonomus) murasakianus Nomura, 1976 | | | Onthophagus (Parascatonomus) negrosensis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) panfilovi Kabakov, 1983 | | | Onthophagus (Parascatonomus) politus Fabricius, 1798 | | | Onthophagus (Parascatonomus) tricornis (Wiedemann, 1823) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus (Parentius) atricapillus d´Orbigny, 1908 | | | Onthophagus (Parentius) emarginatus Mulsant, 1842 | | | Onthophagus (Parentius) hispanicus Baraud, 1963 | | | Onthophagus (Parentius) kabakovi Martin-Piera, 1985 | | | Onthophagus (Parentius) punctatus Illiger, 1803 | | | Onthophagus (Paronthophagus) hidakai Ochi & Kon, 1995 | | | Onthophagus (Phalops) tricuspis (Semenov, 1900) | ⇒ Onthophagus fortigibber Reitter, 1909 | | Onthophagus (Phanaeomorphus) ater Waterhouse, 1875 | | | Onthophagus (Phanaeomorphus) cernyi (Balthasar, 1935) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus (Phanaeomorphus) johkii Ochi & Kon, 1994 | | | Onthophagus (Phanaeomorphus) lenzi Harold, 1875 | | | Onthophagus (Phanaeomorphus) potanini Kabakov, 1979 | | | Onthophagus potanini hiurai Ochi, 1984 | | | Onthophagus (Phanaeomorphus) zetteli Masumoto, Oc..., 2002 | | | Onthophagus (Proagoderus) alcedo d´Orbigny, 1913 | | | Onthophagus (Proagoderus) alcyon (Klug, 1855) | ⇒ Onthophagus alcedo d´Orbigny, 1913 | | Onthophagus (Proagoderus) alcyonides d´Orbigny, 1913 | | | Onthophagus (Proagoderus) aureiceps d´Orbigny, 1902 | | | Onthophagus (Proagoderus) fossidorsis d´Orbigny, 1905 | | | Onthophagus (Proagoderus) mixtifrons d´Orbigny, 1910 | | | Onthophagus (Proagoderus) pseudoalcyon d´Orbigny, 1913 | | | Onthophagus (Proagoderus) pseudovirens d´Orbigny, 1913 | | | Onthophagus (Proagoderus) quadriarmatus Fairmaire, 1892 | | | Onthophagus (Proagoderus) viridiceps d´Orbigny, 1910 | | | Onthophagus (Pseudophanaeomorphus) parachandrai Ochi, Kon & ..., 2009 | | | Onthophagus (Relictonthophagus) emarginatus (Mulsant, 1842) | ⇒ Onthophagus rubrimaculatus MacLeay, 1864 | | Onthophagus (Relictonthophagus) nigellus (Illiger, 1803) | ⇒ Caccobius histeroides Ménétriés, 1832 | | Onthophagus (Relictonthophagus) punctatus (Illiger, 1803) | ⇒ Onthophagus emarginatus Mulsant, 1842 | | Onthophagus (Serrophorus) avocettoides Kabakov, 1983 | | | Onthophagus (Serrophorus) laevis (Harold, 1880) | ⇒ Onthophagus laevissimus d´Orbigny, 1905 | | Onthophagus laevis asiaticus Boucomont, 1919 | | | Onthophagus laevis laevis | | | Onthophagus laevis lampromelas Fairmaire, 1891 | | | Onthophagus laevis stevensi Arrow, 1931 | | | Onthophagus (Serrophorus) oblongus Zhang, 1997 | | | Onthophagus (Serrophorus) rectecornutus Lansberge, 1883 | | | Onthophagus (Serrophorus) sagittarius Fabricius, 1775 | | | Onthophagus (Sinonthophagus) nampatensis Tarasov & Ka..., 2010 | | | Onthophagus (Strandius) changshouensis Zhang, 1997 | | | Onthophagus (Strandius) hiurai (Ochi, 1984) | ⇒ Onthophagus potanini hiurai Ochi, 1984 | | Onthophagus (Strandius) japonicus Harold, 1875 | | | Onthophagus (Strandius) kuraruanus (Matsumura, 1938) | ⇒ Onthophagus lenzi Harold, 1875 | | Onthophagus (Strandius) obliquus Olivier, 1789 | | | Onthophagus (Strandius) potanini R. C. Brusca..., 1990 | | | Onthophagus potanini hiurai Ochi, 1984 | | | Onthophagus potanini potanini | | | Onthophagus (Strandius) subansiriensis Biswas, 1979 | | | Onthophagus (Strandius) taiwanus Nomura, 1973 | | | Onthophagus (Strandius) yakushimanus Nakane, 1983 | | | Onthophagus (Sunenaga) streltsovi Tarasov & Ka..., 2010 | | | Onthophagus (Trichonthophagus) dynastoides Arrow, 1931 | | | Onthophagus (Trichonthophagus) hirtus Illiger, 1803 | | | Onthophagus (Trichonthophagus) maki Illiger, 1803 | | | Onthophagus (Trichonthophagus) tarandus Fabricius, 1792 | | | Onthophagus abas Balthasar, 1946 | | | Onthophagus abei Ochi & Kon, 2015 | | | Onthophagus abeillei d´Orbigny, 1897 | | | Onthophagus abreui Arrow, 1931 | | | Onthophagus abruptus d´Orbigny, 1913 | | | Onthophagus absyrtus Balthasar, 1946 | | | Onthophagus abyssinicus Gillet, 1925 | | | Onthophagus academus Balthasar, 1946 | | | Onthophagus acernorus Moctezuma, H..., 2021 | | | Onthophagus aceroides Krikken & Hu..., 2013 | | | Onthophagus aciculatus (Fahraeus, 1857) | ⇒ Onthophagus aciculatulus Blatchley, 1928 | | Onthophagus acornis Geoffroy, 1785 | | | Onthophagus acrisius Balthasar, 1946 | | | Onthophagus actaeon Balthasar, 1946 | | | Onthophagus acuminatus (Harold, 1880) | ⇒ Onthophagus embrikianus Paulian, 1936 | | Onthophagus acutus d´Orbigny, 1908 | | | Onthophagus admetus Balthasar, 1946 | | | Onthophagus adornatus d´Orbigny, 1904 | | | Onthophagus aegrotus Balthasar, 1967 | | | Onthophagus aeneoniger d´Orbigny, 1913 | | | Onthophagus aeneopiceus d´Orbigny, 1902 | | | Onthophagus aeneus A. G. Olivier, 1789 | | | Onthophagus aequatus Péringuey, 1901 | | | Onthophagus aequepubens d´Orbigny, 1905 | | | Onthophagus aequiperus Matsumura & ..., 1937 | | | Onthophagus aerarius Reitter, 1892 | | | Onthophagus aereidorsis d´Orbigny, 1902 | | | Onthophagus aeremicans d´Orbigny, 1904 | | | Onthophagus aereomaculatus Boucomont, 1914 | | | Onthophagus aereopictus Boucomont, 1914 | | | Onthophagus aereopictus aereopictus | | | Onthophagus aereopictus rubropictus Boucomont, 1924 | | | Onthophagus aerestriatus d´Orbigny, 1913 | | | Onthophagus aeruginosus Roth, 1851 | | | Onthophagus aesopus Lansberge, 1882 | | | Onthophagus aethiopicus d´Orbigny, 1902 | | | Onthophagus affinis Gillet, 1930 | | | Onthophagus africanus Lansberge, 1886 | | | Onthophagus agaricophilus Arrow, 1931 | | | Onthophagus agilis (Matsumura, 1938) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus ahenicollis d´Orbigny, 1902 | | | Onthophagus ahenomicans d´Orbigny, 1902 | | | Onthophagus akamineae Moretto & Gé..., 2020 | | | Onthophagus akhaus Masumoto, Oc..., 2007 | | | Onthophagus akinini Koenig, 1889 | | | Onthophagus akirai (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus alainmonforti Josso, 2022 | | | Onthophagus albicomus d´Orbigny, 1908 | | | Onthophagus albicornis Palisot de B..., 1805 | | | Onthophagus albinae (Reitter, 1898) | ⇒ Onthophagus formaneki Reitter, 1897 | | Onthophagus albipennis Péringuey, 1908 | | | Onthophagus albipodex d´Orbigny, 1902 | | | Onthophagus aleppensis Redtenbacher, 1843 | | | Onthophagus aleppensis aleppensis | | | Onthophagus aleppensis hyrcanicus Kabakov, 1982 | | | Onthophagus alfuricus Huijbregts &..., 2012 | | | Onthophagus aliceae Josso, 2019 | | | Onthophagus alienus Frey, 1958 | | | Onthophagus allojavanus Huijbregts &..., 2011 | | | Onthophagus alluaudi d´Orbigny, 1902 | | | Onthophagus aloysiellus Zunino, 1977 | | | Onthophagus alpinus (Kolenati, 1846) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus alternatus (d´Orbigny, 1902) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus altidorsis d´Orbigny, 1905 | | | Onthophagus altilamina d´Orbigny, 1905 | | | Onthophagus altivagans Howden & Gén..., 2004 | | | Onthophagus amamiensis Nomura, 1965 | | | Onthophagus amamiensis amamiensis | | | Onthophagus amamiensis matsumurai Ochi, Kawaha..., 2019 | | | Onthophagus ambang Krikken & Hu..., 2011 | | | Onthophagus ambang ambang | | | Onthophagus ambang morowali Krikken & Hu..., 2011 | | | Onthophagus amenthus Matsumura, 1937 | | | Onthophagus amoenus d´Orbigny, 1908 | | | Onthophagus amphicoma Boucomont, 1914 | | | Onthophagus amphinasus Arrow, 1931 | | | Onthophagus amphioxus Arrow, 1933 | | | Onthophagus amplexus Sharp, 1875 | | | Onthophagus amplipennis d´Orbigny, 1905 | | | Onthophagus amycoides Kabakov, 2014 | | | Onthophagus analis (Lucas, 1846) | ⇒ Onthophagus sellatus Klug, 1845 | | Onthophagus anatolicus Petrovitz, 1962 | | | Onthophagus andersoni Howden & Gill, 1987 | | | Onthophagus andonarensis Boucomont, 1914 | | | Onthophagus andreji Prokofiev, 2014 | | | Onthophagus andrewesi Arrow, 1931 | | | Onthophagus androgynus d´Orbigny, 1905 | | | Onthophagus anewtoni Howden & Gén..., 2004 | | | Onthophagus angkhanensis Masumoto, Oc..., 2013 | | | Onthophagus angolensis Paulian, 1937 | | | Onthophagus angularis d´Orbigny, 1908 | | | Onthophagus angulatus (Redtenbacher, 1848) | ⇒ Onthophagus gagates Hope, 1831 | | Onthophagus angulicornis (Mulsant, 1842) | ⇒ Onthophagus semicornis Panzer, 1798 | | Onthophagus annoyeri Moretto, 2010 | | | Onthophagus annulopunctatus Krikken & Hu..., 2017 | | | Onthophagus anogeissii Cambefort, 1984 | | | Onthophagus anomalicollis Frey, 1973 | | | Onthophagus anomalipes d´Orbigny, 1902 | | | Onthophagus anonymus Delabie, 1956 | | | Onthophagus antecornutus Josso, 2019 | | | Onthophagus antennalis Frey, 1961 | | | Onthophagus anthracinus (Harold, 1873) | ⇒ Caccobius histeroides Ménétriés, 1832 | | Onthophagus antillarum Arrow, 1903 | | | Onthophagus antilocapra Balthasar, 1937 | | | Onthophagus antilope (Motschulsky, 1845) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus antoinei Walter, 1990 | | | Onthophagus antoineianus Josso, 2022 | | | Onthophagus aokii Nomura, 1976 | | | Onthophagus aphodioides Lansberge, 1883 | | | Onthophagus apiciosus d´Orbigny, 1902 | | | Onthophagus apomontanus Ochi, Kon & ..., 2023 | | | Onthophagus apunneea Masumoto, Oc..., 2007 | | | Onthophagus araxicola (Reitter, 1893) | ⇒ Onthophagus truchmenus Kolenati, 1846 | | Onthophagus arayai Ochi & Kon, 2007 | | | Onthophagus arboreus Arrow, 1931 | | | Onthophagus arcifer d´Orbigny, 1904 | | | Onthophagus ardoini Frey, 1971 | | | Onthophagus areolatus d´Orbigny, 1905 | | | Onthophagus armatus Blanchard, 1853 | | | Onthophagus arunachalensis Biswas & Cha..., 1985 | | | Onthophagus aschenborni Frey, 1975 | | | Onthophagus ashanticola Frey, 1973 | | | Onthophagus asiaticus Endrödi, 1973 | | | Onthophagus asimilis Péringuey, 1901 | | | Onthophagus asper MacLeay, 1864 | | | Onthophagus asperatus d´Orbigny, 1905 | | | Onthophagus aspericeps d´Orbigny, 1908 | | | Onthophagus asperipennis d´Orbigny, 1902 | | | Onthophagus aspernatus d´Orbigny, 1907 | | | Onthophagus asperodorsatus Howden & Gill, 1993 | | | Onthophagus asperrimus d´Orbigny, 1902 | | | Onthophagus asperulus d´Orbigny, 1905 | | | Onthophagus assimilis Hoppe, 1795 | | | Onthophagus astigma d´Orbigny, 1904 | | | Onthophagus athenae (Goidanich, 1925) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus athiensis d´Orbigny, 1913 | | | Onthophagus atricolor d´Orbigny, 1907 | | | Onthophagus atridorsis d´Orbigny, 1902 | | | Onthophagus atriglabrus Howden & Gill, 1987 | | | Onthophagus atroaereus d´Orbigny, 1908 | | | Onthophagus atrofasciatus d´Orbigny, 1905 | | | Onthophagus atronitidus d´Orbigny, 1902 | | | Onthophagus atropolitus d´Orbigny, 1902 | | | Onthophagus atrosericeus Boucomont, 1932 | | | Onthophagus atrosplendens Frey, 1961 | | | Onthophagus atrostriatus d´Orbigny, 1913 | | | Onthophagus atrovirens d´Orbigny, 1905 | | | Onthophagus atrovittatus d´Orbigny, 1908 | | | Onthophagus aureofuscus Bates, 1887 | | | Onthophagus aureopilosus Boucomont, 1914 | | | Onthophagus auriculatus (Klug, 1855) | ⇒ Onthophagus latestriatus d´Orbigny, 1908 | | Onthophagus australis Guérin-Ménev..., 1830 | | | Onthophagus austriacus (Panzer, 1793) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus aviculus Arrow, 1931 | | | Onthophagus avocetta Arrow, 1933 | | | Onthophagus axanicus Ochi, Kon & ..., 2021 | | | Onthophagus axillaris (Boheman, 1860) | ⇒ Onthophagus criniger d´Orbigny, 1904 | | Onthophagus aztecus Zunino & Hal..., 1988 | | | Onthophagus azusae Ochi & Kon, 2006 | | | Onthophagus babaulti d´Orbigny, 1915 | | | Onthophagus babirussa Eschscholtz, 1822 | | | Onthophagus baenzigeri Masumoto, Oc..., 2007 | | | Onthophagus baeri Boucomont, 1914 | | | Onthophagus bajacalifornianus Moctezuma & ..., 2019 | | | Onthophagus bakeri (Boucomont, 1919) | ⇒ Panelus bakeri Boucomont, 1921 | | Onthophagus bakweri Moretto, 2014 | | | Onthophagus balawaicus Scheuern, 1995 | | | Onthophagus baloghi Balthasar, 1967 | | | Onthophagus balthasari Vsetecka, 1939 | | | Onthophagus bandamai Cambefort, 1984 | | | Onthophagus bangueyensis Boucomont, 1914 | | | Onthophagus baolocensis Ochi & Kon, 2015 | | | Onthophagus baoule Cambefort, 1984 | | | Onthophagus baramtagal Ochi, Kon & ..., 2016 | | | Onthophagus barretti Génier & How..., 1999 | | | Onthophagus barriorum Walter, 1990 | | | Onthophagus bartosi Balthasar, 1966 | | | Onthophagus basakata Walter & Cam..., 1977 | | | Onthophagus basalis (Mulsant, 1842) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus basicarinatus Rossini, Vaz..., 2018 | | | Onthophagus basicruentatus Goidanich, 1926 | | | Onthophagus basilewskyi Frey, 1961 | | | Onthophagus basipustulatus Heyden, 1879 | | | Onthophagus bassariscus Zunino & Hal..., 1988 | | | Onthophagus batanensis Ochi, Kawaha..., 2021 | | | Onthophagus bateke Walter & Cam..., 1977 | | | Onthophagus batillifer Harold, 1875 | | | Onthophagus batui Huijbregts &..., 2009 | | | Onthophagus bayeri Balthasar, 1942 | | | Onthophagus bechynei Frey, 1953 | | | Onthophagus beesoni Arrow, 1931 | | | Onthophagus begoniophilus Krikken & Hu..., 2017 | | | Onthophagus beiranus Péringuey, 1908 | | | Onthophagus belinga Walter, 1989 | | | Onthophagus bellus d´Orbigny, 1905 | | | Onthophagus belorhinus Bates, 1887 | | | Onthophagus benedictorum Walter & Cam..., 1977 | | | Onthophagus bengalensis Harold, 1886 | | | Onthophagus bengali Gordon & Opp..., 1977 | | | Onthophagus benguellianus Paulian, 1937 | | | Onthophagus bequaerti d´Orbigny, 1915 | | | Onthophagus bergeri Frey, 1975 | | | Onthophagus bernaudi Cambefort & ..., 1991 | | | Onthophagus betschuanus Frey, 1975 | | | Onthophagus biarcuatus (Lansberge, 1882) | ⇒ Onthophagus sticticus Harold, 1867 | | Onthophagus bicallifrons d´Orbigny, 1902 | | | Onthophagus bicarinatus Gomes Alves, 1944 | | | Onthophagus bicavifrons d´Orbigny, 1902 | | | Onthophagus bicolensis Ochi & Kon, 2006 | | | Onthophagus bicolensis asin Ochi & Kon, 2006 | | | Onthophagus bicolensis bicolensis | | | Onthophagus bicolor (Raffray, 1877) | ⇒ Onthophagus flavolimbatus Klug, 1855 | | Onthophagus biconifer d´Orbigny, 1905 | | | Onthophagus bicornutus Mulsant, 1842 | | | Onthophagus bicristatus d´Orbigny, 1905 | | | Onthophagus bicristiger d´Orbigny, 1913 | | | Onthophagus bicuneus Kabakov, 2008 | | | Onthophagus bidens Olivier, 1789 | | | Onthophagus bidentarus Mulsant, 1842 | | | Onthophagus bidentatus (Drapiez, 1819) | ⇒ Caccobius schreberi Linnaeus, 1767 | | Onthophagus bidentifrons d´Orbigny, 1902 | | | Onthophagus biexcavatus (d´Orbigny, 1898) | ⇒ Onthophagus tarandus Fabricius, 1792 | | Onthophagus bifasciatus Fabricius, 1781 | Zweibindiger Pillenkäfer | | Onthophagus bifenestratus (Reitter, 1898) | ⇒ Onthophagus formaneki Reitter, 1897 | | Onthophagus bifidicornis d´Orbigny, 1902 | | | Onthophagus bifidus Reiche, 1849 | | | Onthophagus bifossus (d´Orbigny, 1902) | ⇒ Onthophagus spinifex Fabricius, 1781 | | Onthophagus bifrons d´Orbigny, 1905 | | | Onthophagus bilingula Balthasar, 1965 | | | Onthophagus bimarginatus d´Orbigny, 1902 | | | Onthophagus binodis Thunberg, 1818 | | | Onthophagus binodosus d´Orbigny, 1908 | | | Onthophagus binodulus d´Orbigny, 1913 | | | Onthophagus biplagiatus Thomson, 1858 | | | Onthophagus birmanicus (Harold, 1879) | ⇒ Onthophagus bifasciatus Fabricius, 1781 | | Onthophagus birugatus d´Orbigny, 1902 | | | Onthophagus birugifer d´Orbigny, 1908 | | | Onthophagus bisbicornis d´Orbigny, 1909 | | | Onthophagus biscarinulatus Huijbregts &..., 2012 | | | Onthophagus bisectus Arrow, 1931 | | | Onthophagus bisignatus d´Orbigny, 1913 | | | Onthophagus bison Boucomont, 1919 | | | Onthophagus bisscrutator Krikken & Hu..., 2017 | | | Onthophagus bistiniocelloides Krikken, 1986 | | | Onthophagus bituber d´Orbigny, 1904 | | | Onthophagus bituberans d´Orbigny, 1905 | | | Onthophagus bituberculatus (Olivier, 1789) | ⇒ Onthophagus acutus d´Orbigny, 1908 | | Onthophagus bituberifrons Josso, 2018 | | | Onthophagus bituberoculus Krikken & Hu..., 2013 | | | Onthophagus bivertex Heyden, 1887 | | | Onthophagus blackburni Shipp, 1895 | | | Onthophagus blanchardi Harold, 1869 | | | Onthophagus blumei (Lansberge, 1883) | ⇒ Onthophagus avocetta Arrow, 1933 | | Onthophagus bocandei d´Orbigny, 1904 | | | Onthophagus bolivari Moctezuma, R..., 2016 | | | Onthophagus bongkudai Krikken & Hu..., 2017 | | | Onthophagus bonorae Zunino, 1976 | | | Onthophagus bonsae Zunino, 1976 | | | Onthophagus boops d´Orbigny, 1905 | | | Onthophagus borassi Cambefort, 1984 | | | Onthophagus bordati Moretto, 2014 | | | Onthophagus borneensis Harold, 1877 | | | Onthophagus borneotagal Ochi, Kon & ..., 2016 | | | Onthophagus bos (Villa, 1833) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus boucomontianus Balthasar, 1935 | | | Onthophagus bourgognei Paulian, 1945 | | | Onthophagus bovillus (Mulsant, 1842) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus bovinus Péringuey, 1892 | | | Onthophagus brachypterus Zunino & Hal..., 1997 | | | Onthophagus brazzavillianus Balthasar, 1974 | | | Onthophagus breviceps d´Orbigny, 1902 | | | Onthophagus brevicollis Arrow, 1907 | | | Onthophagus breviconus Génier & How..., 1999 | | | Onthophagus brevicornis Fahraeus, 1857 | | | Onthophagus brevigena d´Orbigny, 1902 | | | Onthophagus brevipennis d´Orbigny, 1902 | | | Onthophagus brevipes Herbst, 1783 | | | Onthophagus brevisetis d´Orbigny, 1915 | | | Onthophagus brisouti (d´Orbigny, 1897) | ⇒ Onthophagus illyricus Scopoli, 1763 | | Onthophagus brittoni Paulian, 1948 | | | Onthophagus brivioi Frey, 1973 | | | Onthophagus bronzeus Arrow, 1907 | | | Onthophagus brunellii Müller, 1942 | | | Onthophagus brutus Arrow, 1931 | | | Onthophagus buculus Mannerheim, 1829 | | | Onthophagus bufo (Arrow, 1907) | ⇒ Onthophagus semicinctus d´Orbigny, 1897 | | Onthophagus bufonidius Josso, 2019 | | | Onthophagus burchelli d´Orbigny, 1908 | | | Onthophagus caelator Balthasar, 1967 | | | Onthophagus caesariatus Boucomont, 1921 | | | Onthophagus caffrarius d´Orbigny, 1902 | | | Onthophagus calcaratus Boucomont, 1920 | | | Onthophagus calchas Balthasar, 1967 | | | Onthophagus calliger d´Orbigny, 1913 | | | Onthophagus callosipennis Boucomont, 1930 | | | Onthophagus cambrai Delgado & Curoe, 2014 | | | Onthophagus cameloides d´Orbigny, 1900 | | | Onthophagus camelus Fabricius, 1787 | | | Onthophagus camerunicus d´Orbigny, 1905 | | | Onthophagus camilleae Josso, 2022 | | | Onthophagus campestris Masumoto, Ki..., 2018 | | | Onthophagus cancer Lansberge, 1886 | | | Onthophagus canescens Zunino & Hal..., 1988 | | | Onthophagus capella (Kirby, 1818) | ⇒ Onthophagus ramosellus Bates, 1891 | | Onthophagus capilossus d´Orbigny, 1905 | | | Onthophagus capitatus Castelnau, 1840 | | | Onthophagus capitosus Harold, 1867 | | | Onthophagus capra Fabricius, 1787 | | | Onthophagus capreolus (Mulsant, 1842) | ⇒ Onthophagus finschi Harold, 1877 | | Onthophagus carayoni Cambefort, 1986 | | | Onthophagus carbunculus (Boucomont, 1921) | ⇒ Onthophagus lilliputanus Lansberge, 1883 | | Onthophagus carcharias Harold, 1875 | | | Onthophagus carinicollis Raffray, 1877 | | | Onthophagus carinidorsis d´Orbigny, 1908 | | | Onthophagus carinifer d´Orbigny, 1904 | | | Onthophagus carinulatus Harold, 1877 | | | Onthophagus carpanetoi Pittino, 1982 | | | Onthophagus carpophilus Pereira & Ha..., 1961 | | | Onthophagus castanonotus Seabra, 1907 | | | Onthophagus castetsi Lansberge, 1887 | | | Onthophagus catharinensis Paulian, 1936 | | | Onthophagus catherineae Josso, 2022 | | | Onthophagus cavia Boucomont, 1914 | | | Onthophagus caviceps Frey, 1962 | | | Onthophagus cavifrons Harold, 1886 | | | Onthophagus cavivertex d´Orbigny, 1913 | | | Onthophagus centricornis Fabricius, 1798 | | | Onthophagus centromaculatus Redtenbacher, 1843 | | | Onthophagus cephalophi Cambefort, 1984 | | | Onthophagus cervicapra Boucomont, 1914 | | | Onthophagus cervus (Fabricius, 1798) | ⇒ Onthophagus falsus Gillet, 1925 | | Onthophagus ceylonicus Harold, 1877 | | | Onthophagus chacoensis Rossini, Vaz..., 2018 | | | Onthophagus chalcostolus d´Orbigny, 1902 | | | Onthophagus championi Bates, 1887 | | | Onthophagus chandrai Ochi, 2007 | | | Onthophagus chebaicus Boucomont, 1919 | | | Onthophagus chetroiensis Masumoto, Oc..., 2012 | | | Onthophagus cheyi Ochi & Kon, 2006 | | | Onthophagus chiangdaolekus Masumoto, Oc..., 2020 | | | Onthophagus chiapanecus Zunino & Hal..., 1988 | | | Onthophagus chibanus Matsumura & ..., 1937 | | | Onthophagus chilapensis Gasca-Ã�lvar..., 2018 | | | Onthophagus chimalapensis Delgado & Mo..., 2019 | | | Onthophagus chinantecus Moctezuma & ..., 2019 | | | Onthophagus chinbucephalus Ochi & Kon, 2018 | | | Onthophagus chinvigilans Ochi & Kon, 2018 | | | Onthophagus chirindaensis Josso, 2022 | | | Onthophagus chirindanus d´Orbigny, 1913 | | | Onthophagus chitipaensis Josso, 2019 | | | Onthophagus chloroderus d´Orbigny, 1905 | | | Onthophagus chlorophanus d´Orbigny, 1902 | | | Onthophagus choanicus d´Orbigny, 1904 | | | Onthophagus chosensis Matsumura, 1937 | | | Onthophagus chryses Bates, 1887 | | | Onthophagus chrysoderus d´Orbigny, 1905 | | | Onthophagus chrysurus Arrow, 1931 | | | Onthophagus chumphonensis Masumoto, Oc..., 2008 | | | Onthophagus chuzenjianus (Matsumura & ..., 1937) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus cicatricosus (Balthasar, 1935) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus cicer Balthasar, 1964 | | | Onthophagus cincticollis d´Orbigny, 1902 | | | Onthophagus cinctifrons d´Orbigny, 1902 | | | Onthophagus cinctipennis Quedenfeldt, 1884 | | | Onthophagus cinctus (Dallatorre, 1879) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus cineraceus d´Orbigny, 1902 | | | Onthophagus circulator Reitter, 1891 | | | Onthophagus circulifer Arrow, 1931 | | | Onthophagus circumcinctus (Escalera, 1914) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus circumdatus d´Orbigny, 1913 | | | Onthophagus circumscriptus (Faldermann, 1835) | ⇒ Onthophagus marginalis Gebler, 1817 | | Onthophagus citellorum (Petrovitz, 1967) | ⇒ Onthophagus vitulus Péringuey, 1901 | | Onthophagus citreum Boucomont, 1919 | | | Onthophagus civettae Cambefort, 1984 | | | Onthophagus civitasorum Moctezuma & ..., 2020 | | | Onthophagus clavijeroi Moctezuma, R..., 2016 | | | Onthophagus clavisetis d´Orbigny, 1905 | | | Onthophagus clermonti Paulian, 1931 | | | Onthophagus clitellarius d´Orbigny, 1908 | | | Onthophagus clivimerus Huijbregts &..., 2011 | | | Onthophagus clusifrons d´Orbigny, 1905 | | | Onthophagus clypeatus Blanchard, 1846 | | | Onthophagus coeruleicollis Arrow, 1907 | | | Onthophagus cognatus Boucomont, 1921 | | | Onthophagus coiffaiti Frey, 1970 | | | Onthophagus colasi Paulian, 1937 | | | Onthophagus colffsi Lansberge, 1883 | | | Onthophagus collaris (Kabakov, 1983) | ⇒ Proagoderus loricatus Klug, 1855 | | Onthophagus collinsi Krikken & Hu..., 1987 | | | Onthophagus columbianus Boucomont, 1932 | | | Onthophagus comatulus d´Orbigny, 1905 | | | Onthophagus commottoides Kabakov, 1998 | | | Onthophagus comottoi Lansberge, 1885 | | | Onthophagus compactus Arrow, 1933 | | | Onthophagus compressus Guérin-Ménev..., 1855 | | | Onthophagus concavifrons d´Orbigny, 1904 | | | Onthophagus concolor Sharp, 1878 | | | Onthophagus concursator (Balthasar, 1960) | ⇒ Onthophagus sutleinensis Splichal, 1910 | | Onthophagus confertus Péringuey, 1908 | | | Onthophagus confluens (d´Orbigny, 1902) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus confucius (Goidanich, 1926) | ⇒ Onthophagus solivagus Harold, 1886 | | Onthophagus confusus Boucomont, 1932 | | | Onthophagus conjugata (Seabra, 1907) | ⇒ Onthophagus hirtus Illiger, 1803 | | Onthophagus conradsi Balthasar, 1937 | | | Onthophagus consentaneus Harold, 1867 | | | Onthophagus conspurcatus Geoffroy, 1785 | | | Onthophagus contiguicornis d´Orbigny, 1913 | | | Onthophagus contrapositus Génier, 2017 | | | Onthophagus convexicollis Boheman, 1858 | | | Onthophagus convexipennis Dierkens, 2022 | | | Onthophagus convexus d´Orbigny, 1908 | | | Onthophagus coomani Paulian, 1931 | | | Onthophagus coorgensis Arrow, 1931 | | | Onthophagus coprimorphoides Huijbregts &..., 2012 | | | Onthophagus coptorhinodes Péringuey, 1901 | | | Onthophagus coracinoides Kabakov, 1983 | | | Onthophagus coracinus Boucomont, 1914 | | | Onthophagus coriaceoumbrosus Kohlmann & S..., 2001 | | | Onthophagus cornicollis d´Orbigny, 1902 | | | Onthophagus corniculatus (Reiche, 1847) | ⇒ Onthophagus senegalensis d´Orbigny, 1902 | | Onthophagus corniculiger d´Orbigny, 1913 | | | Onthophagus cornifrons Thomson, 1858 | | | Onthophagus corniger Geoffroy, 1785 | | | Onthophagus cornutus Ferreira, 1971 | | | Onthophagus coronatus d´Orbigny, 1902 | | | Onthophagus corrosus Bates, 1887 | | | Onthophagus corvus Fabricius, 1798 | | | Onthophagus coscineus Bates, 1887 | | | Onthophagus costatus d´Orbigny, 1913 | | | Onthophagus costiger d´Orbigny, 1915 | | | Onthophagus costilatus d´Orbigny, 1908 | | | Onthophagus costulicollis Frey, 1975 | | | Onthophagus crantor Balthasar, 1967 | | | Onthophagus crassicollis Boucomont, 1913 | | | Onthophagus crassus ♰ (Heer, 1862) | ⇒ Parascatonomus tricornis Wiedemann, 1823 | | Onthophagus creber d´Orbigny, 1905 | | | Onthophagus cretus (Péringuey, 1901) | ⇒ Onthophagus setosus Fahraeus, 1857 | | Onthophagus cribellum d´Orbigny, 1902 | | | Onthophagus cribratus Lansberge, 1883 | | | Onthophagus cribripennis d´Orbigny, 1902 | | | Onthophagus criniger d´Orbigny, 1904 | | | Onthophagus crinitus (Harold, 1869) | ⇒ Onthophagus criniger d´Orbigny, 1904 | | Onthophagus crinitus crinitus | | | Onthophagus crinitus panamensis Bates, 1887 | | | Onthophagus cristatus d´Orbigny, 1905 | | | Onthophagus croesulus Bates, 1888 | | | Onthophagus crucenotatus d´Orbigny, 1905 | | | Onthophagus cruentatus Klug, 1855 | | | Onthophagus cruoreus Schrank, 1798 | | | Onthophagus cryptodicranius Kohlmann & S..., 2001 | | | Onthophagus cryptogenus Boucomont, 1914 | | | Onthophagus cuboidalis (Bates, 1887) | ⇒ Onthophagus coproides Horn, 1881 | | Onthophagus cultrifer (Harold, 1886) | ⇒ Onthophagus falcifer Harold, 1880 | | Onthophagus cuneus Boucomont, 1924 | | | Onthophagus cupreiceps Arrow, 1907 | | | Onthophagus cupreolus (Reitter, 1894) | ⇒ Onthophagus punctatus Illiger, 1803 | | Onthophagus cupreopastillatus Ochi & Kon, 2006 | | | Onthophagus cupreopinguis Josso, 2022 | | | Onthophagus cupreotagal Ochi, Kon & ..., 2016 | | | Onthophagus cupreovirens d´Orbigny, 1904 | | | Onthophagus cupreus Harold, 1880 | | | Onthophagus cupricollis Péringuey, 1888 | | | Onthophagus curtipilis d´Orbigny, 1905 | | | Onthophagus curtulus Frey, 1961 | | | Onthophagus curvicinctus (Mulsant, 1842) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus curvicornis Latreille, 1811 | | | Onthophagus curvifrons d´Orbigny, 1906 | | | Onthophagus curvilamina Müller, 1942 | | | Onthophagus curvispina Reitter, 1893 | | | Onthophagus cuspidiusculus (Mulsant, 1842) | ⇒ Onthophagus coenobita Herbst, 1783 | | Onthophagus cutai (Roubal, 1933) | ⇒ Onthophagus ruficapillus Brullé, 1832 | | Onthophagus cyaneiceps d´Orbigny, 1913 | | | Onthophagus cyaneoniger d´Orbigny, 1902 | | | Onthophagus cyaneus Kabakov, 2006 | | | Onthophagus cyanochlorus d´Orbigny, 1902 | | | Onthophagus cyclographus Bates, 1887 | | | Onthophagus cynomisi Brown, 1927 | | | Onthophagus cyobioides Boucomont, 1921 | | | Onthophagus danumicus Huijbregts &..., 2012 | | | Onthophagus dapitanensis Boucomont, 1919 | | | Onthophagus davisi Frey, 1975 | | | Onthophagus dayacus Boucomont, 1914 | | | Onthophagus debilis d´Orbigny, 1905 | | | Onthophagus deccanensis Frey, 1962 | | | Onthophagus decedens Péringuey, 1904 | | | Onthophagus decempunctatus Schaller, 1783 | | | Onthophagus decens Balthasar, 1964 | | | Onthophagus decipiens (Mulsant, 1842) | ⇒ Onthophagus semicornis Panzer, 1798 | | Onthophagus declivicollis d´Orbigny, 1902 | | | Onthophagus decolor d´Orbigny, 1902 | | | Onthophagus decoratus d´Orbigny, 1905 | | | Onthophagus decorsei d´Orbigny, 1908 | | | Onthophagus dedecor Wallengren, 1881 | | | Onthophagus deflexicollis Lansberge, 1883 | | | Onthophagus deflexus d´Orbigny, 1908 | | | Onthophagus degener Mulsant, 1842 | | | Onthophagus delahayei Josso, 2011 | | | Onthophagus deletus (Reitter, 1893) | ⇒ Onthophagus sibiricus Harold, 1877 | | Onthophagus delicatulus Raffray, 1877 | | | Onthophagus delicatus d´Orbigny, 1915 | | | Onthophagus deliensis Lansberge, 1885 | | | Onthophagus deliensis bawangicus Ochi, Kon & ..., 2009 | | | Onthophagus deliensis deliensis | | | Onthophagus delkeskampi (Frey, 1954) | ⇒ Onthophagus mirandus Arrow, 1931 | | Onthophagus densatus Frey, 1963 | | | Onthophagus densepunctatus Frey, 1963 | | | Onthophagus densipilis d´Orbigny, 1902 | | | Onthophagus denticornis Boucomont, 1914 | | | Onthophagus denticulatus d´Orbigny, 1902 | | | Onthophagus denudatus d´Orbigny, 1902 | | | Onthophagus denudatus denudatus | | | Onthophagus denudatus endroedii Frey, 1973 | | | Onthophagus depilatus d´Orbigny, 1913 | | | Onthophagus depilis d´Orbigny, 1902 | | | Onthophagus deplanatus Lansberge, 1883 | | | Onthophagus depolii (Goidanich, 1925) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus depressicollis Boucomont, 1914 | | | Onthophagus depressifrons d´Orbigny, 1915 | | | Onthophagus depressipennis Cambefort, 1975 | | | Onthophagus derasus d´Orbigny, 1902 | | | Onthophagus desaegeri Frey, 1961 | | | Onthophagus deterrens Péringuey, 1901 | | | Onthophagus devagiriensis Schoolmeeste..., 2006 | | | Onthophagus dhanjuricus Frey, 1973 | | | Onthophagus diadematus d´Orbigny, 1902 | | | Onthophagus dicax Balthasar, 1966 | | | Onthophagus dicella Bates, 1888 | | | Onthophagus dicranius Bates, 1887 | | | Onthophagus dicranoides Balthasar, 1939 | | | Onthophagus difficilis (Le Guillou, 1844) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus digitatus Arrow, 1931 | | | Onthophagus digitifer (Boucomont, 1932) | ⇒ Onthophagus coscineus Bates, 1887 | | Onthophagus dillwynii (Stephens, 1830) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus dilutus d´Orbigny, 1905 | | | Onthophagus dinoderus d´Orbigny, 1913 | | | Onthophagus discoideus A. G. Olivier, 1789 | | | Onthophagus discovirens d´Orbigny, 1902 | | | Onthophagus discretus Péringuey, 1901 | | | Onthophagus dispar Péringuey, 1901 | | | Onthophagus dissentaneus Balthasar, 1964 | | | Onthophagus distichus Roth, 1851 | | | Onthophagus distinctus (Fuente, 1910) | ⇒ Onthophagus latigena d´Orbigny, 1897 | | Onthophagus distinguendus Mulsant, 1842 | | | Onthophagus ditus (Péringuey, 1901) | ⇒ Onthophagus fungicola d´Orbigny, 1908 | | Onthophagus diversicornis (Kirschenblatt, 1935) | ⇒ Onthophagus aerarius Reitter, 1892 | | Onthophagus diversus Reiche, 1847 | | | Onthophagus dlabolai Balthasar, 1964 | | | Onthophagus dohertyi d´Orbigny, 1902 | | | Onthophagus doriae Harold, 1877 | | | Onthophagus dorsipilulus Howden & Gill, 1987 | | | Onthophagus dorsofasciatus Fairmaire, 1893 | | | Onthophagus dorsosignatus d´Orbigny, 1898 | | | Onthophagus dorsuosus d´Orbigny, 1902 | | | Onthophagus drescheri Paulian, 1939 | | | Onthophagus drumonti Moretto, 1999 | | | Onthophagus dubitabilis Howden & Gén..., 2004 | | | Onthophagus ducorpsi d´Orbigny, 1913 | | | Onthophagus dudleyianus Josso, 2019 | | | Onthophagus durangoensis Balthasar, 1939 | | | Onthophagus duvivieri d´Orbigny, 1904 | | | Onthophagus ebenicolor d´Orbigny, 1902 | | | Onthophagus ebenus Péringuey, 1888 | | | Onthophagus eburneus d´Orbigny, 1913 | | | Onthophagus echinus Boucomont, 1914 | | | Onthophagus ecopas Josso & Prév..., 2006 | | | Onthophagus egenus (Harold, 1877) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus egurianus Matsumura, 1938 | | | Onthophagus elongatus Frey, 1954 | | | Onthophagus embrikianus Paulian, 1936 | | | Onthophagus emeritus Péringuey, 1901 | | | Onthophagus endroedianus Walter & Cam..., 1977 | | | Onthophagus ensifer Boucomont, 1914 | | | Onthophagus enuguensis Frey, 1975 | | | Onthophagus ephippioderus Arrow, 1907 | | | Onthophagus epilamprus Bates, 1888 | | | Onthophagus erectinasus d´Orbigny, 1902 | | | Onthophagus erectus Wiedemann, 1819 | | | Onthophagus escalerai d´Orbigny, 1902 | | | Onthophagus eschscholtzi Boucomont, 1924 | | | Onthophagus estelleae Josso, 2022 | | | Onthophagus etlaensis Kohlmann, Es..., 2019 | | | Onthophagus eulophus Bates, 1887 | | | Onthophagus euthymi (Mulsant & Go..., 1875) | ⇒ Onthophagus sellatus Klug, 1845 | | Onthophagus euzeti Walter & Cam..., 1977 | | | Onthophagus evae Balthasar, 1944 | | | Onthophagus exasperatus (Gerstaecker, 1871) | ⇒ Onthophagus latissimus d´Orbigny, 1902 | | Onthophagus excisiceps d´Orbigny, 1902 | | | Onthophagus exiguus (Raffray, 1877) | ⇒ Onthophagus rubens d´Orbigny, 1902 | | Onthophagus exilis d´Orbigny, 1913 | | | Onthophagus expansicollis (Fairmaire, 1891) | ⇒ Onthophagus rugulosus Harold, 1886 | | Onthophagus expansicornis Bates, 1891 | | | Onthophagus exquisitus Arrow, 1931 | | | Onthophagus extensicollis d´Orbigny, 1907 | | | Onthophagus extraneus Josso, 2021 | | | Onthophagus fabricianus Goidanich, 1926 | | | Onthophagus fabricii Waterhouse, 1894 | | | Onthophagus falcarius Balthasar, 1966 | | | Onthophagus falcifer Harold, 1880 | | | Onthophagus falculatus Boucomont, 1914 | | | Onthophagus fallaciosus Raffray, 1877 | | | Onthophagus fallax d´Orbigny, 1913 | | | Onthophagus falsus Gillet, 1925 | | | Onthophagus fanatus Laporte, 1840 | | | Onthophagus fasciatus (Boucomont, 1914) | ⇒ Onthophagus fasciolatus Boucomont, 1924 | | Onthophagus fasciculiger d´Orbigny, 1902 | | | Onthophagus fasciolatus Boucomont, 1924 | | | Onthophagus favrei Boucomont, 1914 | | | Onthophagus feai d´Orbigny, 1905 | | | Onthophagus felix Arrow, 1931 | | | Onthophagus femineus (Mulsant, 1842) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus filicornis Harold, 1873 | | | Onthophagus fimetarius Roth, 1851 | | | Onthophagus fissinasus Fairmaire, 1895 | | | Onthophagus fitiniensis Cambefort, 1984 | | | Onthophagus flagrans Reitter, 1893 | | | Onthophagus flavescens Seabra, 1907 | | | Onthophagus flavibasis d´Orbigny, 1904 | | | Onthophagus flaviclava d´Orbigny, 1902 | | | Onthophagus flavimargo d´Orbigny, 1902 | | | Onthophagus flavipennis d´Orbigny, 1905 | | | Onthophagus flavipleurus Dierkens, 2022 | | | Onthophagus flavocinctus Frey, 1975 | | | Onthophagus flavofasciatus (d´Orbigny, 1900) | ⇒ Onthophagus dorsosignatus d´Orbigny, 1898 | | Onthophagus flavolimbatus Klug, 1855 | | | Onthophagus flavonotatus d´Orbigny, 1900 | | | Onthophagus flavorufus d´Orbigny, 1904 | | | Onthophagus flexicornis d´Orbigny, 1902 | | | Onthophagus flexifrons d´Orbigny, 1908 | | | Onthophagus fodiens Waterhouse, 1875 | | | Onthophagus foedus Boucomont, 1914 | | | Onthophagus foliaceus Lansberge, 1886 | | | Onthophagus foliiceps Quedenfeldt, 1884 | | | Onthophagus foraminosus d´Orbigny, 1902 | | | Onthophagus formaneki Reitter, 1897 | | | Onthophagus fossatus Castelnau, 1840 | | | Onthophagus fossibasis d´Orbigny, 1908 | | | Onthophagus fossifrons d´Orbigny, 1902 | | | Onthophagus fossor Arrow, 1931 | | | Onthophagus fossulatus d´Orbigny, 1908 | | | Onthophagus foulliouxi Cambefort, 1971 | | | Onthophagus foveatus Frey, 1957 | | | Onthophagus foveicollis Boucomont, 1930 | | | Onthophagus foveolatus (Harold, 1877) | ⇒ Parascatonomus rudis Sharp, 1875 | | Onthophagus fradei Gomes Alves, 1956 | | | Onthophagus fragosus Génier & Ho..., 2014 | | | Onthophagus francoisi Josso, 2013 | | | Onthophagus frankenbergeri Balthasar, 1933 | | | Onthophagus frankrozendaali Huijbregts &..., 2012 | | | Onthophagus fritschi d´Orbigny, 1902 | | | Onthophagus frontalis Raffray, 1877 | | | Onthophagus frugivorus Arrow, 1931 | | | Onthophagus fucatrus (Mulsant, 1842) | ⇒ Onthophagus ovatus Linnaeus, 1767 | | Onthophagus fugitivus Péringuey, 1901 | | | Onthophagus fukiensis Balthasar, 1942 | | | Onthophagus fukuyamai Ochi, Kon & ..., 2018 | | | Onthophagus fulgens Brahm, 1790 | | | Onthophagus fulvicornis (Motschulsky, 1853) | ⇒ Onthophagus tragus Fabricius, 1792 | | Onthophagus fulvipes (Faldermann, 1835) | ⇒ Onthophagus coenobita Herbst, 1783 | | Onthophagus fulvocinctus d´Orbigny, 1902 | | | Onthophagus fulvus Sharp, 1875 | | | Onthophagus fumatus d´Orbigny, 1913 | | | Onthophagus funebris Boucomont, 1919 | | | Onthophagus funestus Moretto & Gé..., 2010 | | | Onthophagus fungicola d´Orbigny, 1908 | | | Onthophagus fungus Josso, 2022 | | | Onthophagus furcaticeps Masters, 1886 | | | Onthophagus furcatoides (Lansberge, 1886) | ⇒ Onthophagus neofurcatus Goidanich, 1926 | | Onthophagus furcicollis (Arrow, 1931) | ⇒ Onthophagus hecate Panzer, 1794 | | Onthophagus furcicornis (Reitter, 1893) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus furcillifer Bates, 1891 | | | Onthophagus furculifer d´Orbigny, 1905 | | | Onthophagus furculus Fabricius, 1798 | | | Onthophagus fuscatus d´Orbigny, 1908 | | | Onthophagus fuscidorsis (d´Orbigny, 1902) | ⇒ Onthophagus fuscatus d´Orbigny, 1908 | | Onthophagus fuscipennis (Mulsant, 1842) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus fuscivestis d´Orbigny, 1902 | | | Onthophagus fuscopunctatus (Fabricius, 1798) | ⇒ Onthophagus luridipennis Boheman, 1858 | | Onthophagus fuscopunctulatus (Boucomont, 1914) | ⇒ Onthophagus luridipennis Boheman, 1858 | | Onthophagus fuscostriatus Boucomont, 1914 | | | Onthophagus futsukaichianus Matsumura, 1937 | | | Onthophagus gabonensis Walter, 1989 | | | Onthophagus gaesatus Boucomont, 1925 | | | Onthophagus gagates Hope, 1831 | | | Onthophagus gagatoides Kabakov, 2006 | | | Onthophagus gaillardi d´Orbigny, 1911 | | | Onthophagus gajo Krikken, 1977 | | | Onthophagus galeatus Boucomont, 1919 | | | Onthophagus ganalensis Gestro, 1895 | | | Onthophagus gangeticus Gillet, 1925 | | | Onthophagus garambae Frey, 1961 | | | Onthophagus gazellinus Bates, 1887 | | | Onthophagus gemellatus Raffray, 1877 | | | Onthophagus geminatus d´Orbigny, 1907 | | | Onthophagus geminifrons d´Orbigny, 1905 | | | Onthophagus gemma Sharp, 1875 | | | Onthophagus genuinus Kohlmann & S..., 2001 | | | Onthophagus georgwerneri Josso, 2022 | | | Onthophagus germanus Gillet, 1927 | | | Onthophagus germari (Depoli, 1917) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus gestroi (Harold, 1877) | ⇒ Onthophagus bellus d´Orbigny, 1905 | | Onthophagus ghanensis Balthasar, 1966 | | | Onthophagus gibber d´Orbigny, 1905 | | | Onthophagus gibberosus d´Orbigny, 1905 | | | Onthophagus gibbicollis Lansberge, 1885 | | | Onthophagus gibbidorsis d´Orbigny, 1902 | | | Onthophagus gibbifrons d´Orbigny, 1902 | | | Onthophagus gibbus d´Orbigny, 1913 | | | Onthophagus gibsoni Howden & Gén..., 2004 | | | Onthophagus gilli Delgado & Ho..., 2000 | | | Onthophagus ginyunensis Vsetecka, 1942 | | | Onthophagus giraffa Hausmann, 1807 | | | Onthophagus girardinae Walter, 1989 | | | Onthophagus giuseppecarpanetoi Tagliaferri ..., 2012 | | | Onthophagus glabra (Seabra, 1907) | ⇒ Onthophagus maki Illiger, 1803 | | Onthophagus glandicolor (Mulsant, 1842) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus glareanus (Gistel, 1857) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus glasunovi Koshantschikov, 1894 | | | Onthophagus glasunowi D. Koshantsc..., 1894 | | | Onthophagus gnu Frey, 1955 | | | Onthophagus gonipa Krikken & Hu..., 2011 | | | Onthophagus gonopygus d´Orbigny, 1902 | | | Onthophagus gonzaloi Moctezuma, H..., 2022 | | | Onthophagus gorhami (Shipp, 1895) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus gracilicornis (Raffray, 1877) | ⇒ Onthophagus filicornis Harold, 1873 | | Onthophagus gradivus Balthasar, 1966 | | | Onthophagus grandidorsis d´Orbigny, 1913 | | | Onthophagus grandifrons d´Orbigny, 1907 | | | Onthophagus graniceps d´Orbigny, 1908 | | | Onthophagus granosus d´Orbigny, 1913 | | | Onthophagus granulifer (Harold, 1886) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus granulifrons Frey, 1961 | | | Onthophagus granulopygus Dierkens, 2022 | | | Onthophagus granulosus (Koshantschikov, 1894) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus granulum d´Orbigny, 1904 | | | Onthophagus graphicus Wallengren, 1881 | | | Onthophagus grassei Walter, 1989 | | | Onthophagus grataehelenae Kohlmann & S..., 2001 | | | Onthophagus gratus Arrow, 1931 | | | Onthophagus gravearmatus Balthasar, 1944 | | | Onthophagus gravis Walker, 1858 | | | Onthophagus gravoti d´Orbigny, 1908 | | | Onthophagus griseoaeneus Lansberge, 1885 | | | Onthophagus griseosetosus Arrow, 1931 | | | Onthophagus guatemalensis (Bates, 1887) | ⇒ Onthophagus medorensis Brown, 1929 | | Onthophagus guilanensis (Pittino, 1982) | ⇒ Onthophagus ruficapillus Brullé, 1832 | | Onthophagus gulo Arrow, 1931 | | | Onthophagus guttatus (Boheman, 1860) | ⇒ Onthophagus quadrilunatus d´Orbigny, 1902 | | Onthophagus guttiger d´Orbigny, 1905 | | | Onthophagus haafi Frey, 1963 | | | Onthophagus haematopus Harold, 1875 | | | Onthophagus hageni Lansberge, 1883 | | | Onthophagus halffteri Zunino, 1981 | | | Onthophagus hamaticeps Arrow, 1931 | | | Onthophagus hanboonsongae Masumoto, Oc..., 2021 | | | Onthophagus hartiniae Ochi & Kon, 2009 | | | Onthophagus helciatus Harold, 1871 | | | Onthophagus hemichalceus d´Orbigny, 1915 | | | Onthophagus hemichlorus d´Orbigny, 1915 | | | Onthophagus hemipygus Frey, 1964 | | | Onthophagus herbsti Brahm, 1790 | | | Onthophagus hericiniformis Moretto, 2010 | | | Onthophagus hericius d´Orbigny, 1902 | | | Onthophagus hermonensis Baraud, 1982 | | | Onthophagus herus Péringuey, 1901 | | | Onthophagus heteroclitus d´Orbigny, 1911 | | | Onthophagus heterorrhinus Lansberge, 1885 | | | Onthophagus heydeni Harold, 1875 | | | Onthophagus heyrovskyi Vsetecka, 1943 | | | Onthophagus hielkemai Krikken & Hu..., 2011 | | | Onthophagus hikidai Ochi & Kon, 2006 | | | Onthophagus hikosanus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus hilaridis Cambefort, 1984 | | | Onthophagus hilaris d´Orbigny, 1905 | | | Onthophagus hildebrandti Harold, 1878 | | | Onthophagus hindu Arrow, 1931 | | | Onthophagus hingstoni Arrow, 1931 | | | Onthophagus hippopotamus Harold, 1869 | | | Onthophagus hiranoyasutoshii Ochi & Kon, 2018 | | | Onthophagus hirculus (Mannerheim, 1829) | ⇒ Onthophagus mutatus Harold, 1859 | | Onthophagus hircus Billberg, 1815 | | | Onthophagus hiroyukii Ochi, 2007 | | | Onthophagus hirsutulus Lansberge, 1883 | | | Onthophagus hirsutus d´Orbigny, 1902 | | | Onthophagus hirtellus Frey, 1957 | | | Onthophagus hirticulus d´Orbigny, 1915 | | | Onthophagus hirtipodex d´Orbigny, 1904 | | | Onthophagus hirtulus (Reitter, 1893) | ⇒ Onthophagus hirtus Illiger, 1803 | | Onthophagus histeriformis Boucomont, 1914 | | | Onthophagus histrio d´Orbigny, 1902 | | | Onthophagus hoberlandti Balthasar, 1966 | | | Onthophagus hollowayi Krikken & Hu..., 2017 | | | Onthophagus holosericeus Harold, 1877 | | | Onthophagus holosericus Harold, 1877 | | | Onthophagus holzi Boucomont, 1914 | | | Onthophagus hoogstraali Saylor, 1943 | | | Onthophagus horni Balthasar, 1935 | | | Onthophagus horrens d´Orbigny, 1908 | | | Onthophagus horridus d´Orbigny, 1908 | | | Onthophagus hosomai Ochi & Kon, 2014 | | | Onthophagus howdeni Zunino & Hal..., 1988 | | | Onthophagus howdenorum Zunino & Hal..., 1988 | | | Onthophagus hrubanti (Balthasar, 1955) | ⇒ Onthophagus ramosellus Bates, 1891 | | Onthophagus hulstaerti Boucomont, 1932 | | | Onthophagus humboldti Kohlmann, So..., 2019 | | | Onthophagus humeralis (Miksic, 1950) | ⇒ Onthophagus vitulus Péringuey, 1901 | | Onthophagus humpatensis Lansberge, 1886 | | | Onthophagus hvangeus Stebnicka, 1973 | | | Onthophagus hyaena Fabricius, 1801 | | | Onthophagus hypertropis (Olsoufieff, 1900) | ⇒ Onthophagus tragus Fabricius, 1792 | | Onthophagus hyrcanius Kabakov, 1982 | | | Onthophagus hystrix Boucomont, 1914 | | | Onthophagus hyuganus Matsumura & ..., 1937 | | | Onthophagus ibex (Fabricius, 1792) | ⇒ Onthophagus tragus Fabricius, 1792 | | Onthophagus ibonus (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus ibrydus O. G. Costa, 1828 | | | Onthophagus ieti Cambefort, 1984 | | | Onthophagus ifugaoensis Ochi & Kon, 2006 | | | Onthophagus igneus Vigors, 1825 | | | Onthophagus igualensis Bates, 1887 | | | Onthophagus ikelengeensis Josso, 2018 | | | Onthophagus illotus Péringuey, 1901 | | | Onthophagus imbellis d´Orbigny, 1905 | | | Onthophagus imberbis d´Orbigny, 1902 | | | Onthophagus imbutus Sharp, 1875 | | | Onthophagus immaculatus (Mulsant, 1842) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus immundus Boheman, 1858 | | | Onthophagus importunus Péringuey, 1901 | | | Onthophagus impressicollis Boheman, 1860 | | | Onthophagus impunctatus d´Orbigny, 1904 | | | Onthophagus impuncticollis (Frey, 1950) | ⇒ Onthophagus atropolitus d´Orbigny, 1902 | | Onthophagus impurus Harold, 1868 | | | Onthophagus inaequalis d´Orbigny, 1902 | | | Onthophagus incantatus Balthasar, 1967 | | | Onthophagus incertus (d´Orbigny, 1897) | ⇒ Onthophagus asperrimus d´Orbigny, 1902 | | Onthophagus incisus Harold, 1877 | | | Onthophagus includens d´Orbigny, 1905 | | | Onthophagus inclusus d´Orbigny, 1905 | | | Onthophagus incollaris Kabakov, 2008 | | | Onthophagus indigus Péringuey, 1901 | | | Onthophagus indistinctus (Mulsant, 1842) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus indosinicus Boucomont, 1921 | | | Onthophagus indutus d´Orbigny, 1902 | | | Onthophagus inecolorum Moctezuma & ..., 2020 | | | Onthophagus inediapterus Kohlmann & S..., 2001 | | | Onthophagus ineptus Harold, 1871 | | | Onthophagus inermiceps d´Orbigny, 1905 | | | Onthophagus inermicollis d´Orbigny, 1908 | | | Onthophagus inermivertex Boucomont, 1921 | | | Onthophagus infaustus Cambefort, 1984 | | | Onthophagus inflaticollis Bates, 1887 | | | Onthophagus inflatus Walter & Cam..., 1977 | | | Onthophagus infucatus Harold, 1877 | | | Onthophagus infuscata (Seabra, 1907) | ⇒ Onthophagus hirtus Illiger, 1803 | | Onthophagus infuscatus Klug, 1845 | | | Onthophagus inquinatus (Goidanich, 1925) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus insignis (Péringuey, 1896) | ⇒ Proagoderus fossidorsis d´Orbigny, 1905 | | Onthophagus insularis Boheman, 1858 | | | Onthophagus insulindicus Boucomont, 1914 | | | Onthophagus insulsus Péringuey, 1901 | | | Onthophagus intercepta (Seabra, 1907) | ⇒ Onthophagus maki Illiger, 1803 | | Onthophagus intermedius (Mulsant, 1842) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus intermixtus d´Orbigny, 1902 | | | Onthophagus interruptus (Raffray, 1877) | ⇒ Onthophagus leanus Goidanich, 1926 | | Onthophagus interstitialis (Fahraeus, 1857) | ⇒ Euonthophagus carbonarius Klug, 1855 | | Onthophagus intonsus d´Orbigny, 1913 | | | Onthophagus intricatus Moretto, 2010 | | | Onthophagus investis d´Orbigny, 1904 | | | Onthophagus iodiellus Bates, 1887 | | | Onthophagus iringaensis Dierkens, 2022 | | | Onthophagus irroratus Faldermann, 1835 | | | Onthophagus isanus Masumoto, Oc..., 2008 | | | Onthophagus istmenus Moctezuma, S..., 2020 | | | Onthophagus italicus (Goidanich, 1925) | ⇒ Onthophagus andalusicus Waltl, 1835 | | Onthophagus itoi Nomura, 1976 | | | Onthophagus itoianus Masumoto, Oc..., 2022 | | | Onthophagus itonoborui Masumoto, Oc..., 2012 | | | Onthophagus iulicola Cambefort, 1984 | | | Onthophagus iumienus Masumoto, Oc..., 2007 | | | Onthophagus ixtepecorum Moctezuma & ..., 2020 | | | Onthophagus ixtlanensis Moctezuma & ..., 2020 | | | Onthophagus iyengari Arrow, 1931 | | | Onthophagus jacksoni d´Orbigny, 1913 | | | Onthophagus jacobeus Boucomont, 1924 | | | Onthophagus jalapensis Balthasar, 1939 | | | Onthophagus jaliscensis Zunino & Hal..., 1988 | | | Onthophagus janssensi Frey, 1958 | | | Onthophagus janthinus (Harold, 1875) | ⇒ Onthophagus aeruginosus Roth, 1851 | | Onthophagus javanensis Balthasar, 1963 | | | Onthophagus javanoides Huijbregts &..., 2011 | | | Onthophagus javanus (Lansberge, 1883) | ⇒ Onthophagus javanensis Balthasar, 1963 | | Onthophagus jeanneli d´Orbigny, 1913 | | | Onthophagus jeanneli diversiceps Frey, 1975 | | | Onthophagus jeannelianus Paulian, 1945 | | | Onthophagus jeannelianus jeannelianus | | | Onthophagus jeannelianus shokhini Kabakov, 2014 | | | Onthophagus jedensis Matsumura, 1937 | | | Onthophagus jingping Masumoto, Oc..., 2007 | | | Onthophagus jiupengensis Masumoto, La..., 2017 | | | Onthophagus jocquei Josso, 2019 | | | Onthophagus jonathani Moretto & Gé..., 2020 | | | Onthophagus jononis Matsumura, 1936 | | | Onthophagus jucundus (Arrow, 1931) | ⇒ Onthophagus cognatus Boucomont, 1921 | | Onthophagus jugicola d´Orbigny, 1902 | | | Onthophagus juheli Josso, 2022 | | | Onthophagus juncticornis d´Orbigny, 1908 | | | Onthophagus juniperensis Josso, 2019 | | | Onthophagus justei Walter, 1989 | | | Onthophagus juvencus Klug, 1835 | | | Onthophagus jwalae Karimbumkara..., 2016 | | | Onthophagus kabulicus (Balthasar, 1971) | ⇒ Onthophagus conspersus Reitter, 1893 | | Onthophagus kachinicus Kabakov, 2006 | | | Onthophagus kaengkrachangus Masumoto, Oc..., 2008 | | | Onthophagus kanarensis Arrow, 1931 | | | Onthophagus kandai (Matsumura, 1937) | ⇒ Onthophagus olsoufieffi Boucomont, 1924 | | Onthophagus kangeanus Paulian, 1936 | | | Onthophagus kaomaensis Josso, 2022 | | | Onthophagus kaosoidowensis Masumoto, Oc..., 2008 | | | Onthophagus kapangaensis Dierkens, 2022 | | | Onthophagus kapitensis Frey, 1971 | | | Onthophagus kapuri Endrödi, 1974 | | | Onthophagus karlwerneri Josso, 2022 | | | Onthophagus kashizakii Kon & Ochi, 2005 | | | Onthophagus kashmirensis Balthasar, 1966 | | | Onthophagus kassaicus d´Orbigny, 1908 | | | Onthophagus katangensis Frey, 1961 | | | Onthophagus katayamai Masumoto, Oc..., 2012 | | | Onthophagus kavirondus d´Orbigny, 1915 | | | Onthophagus kawaharai Ochi & Kon, 2007 | | | Onthophagus kawarinus (Matsumura & ..., 1937) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus kchatriya Boucomont, 1914 | | | Onthophagus keikoae Ochi & Kon, 2014 | | | Onthophagus keiseri Frey, 1956 | | | Onthophagus keralensis Frey, 1975 | | | Onthophagus keralicus Biswas & Cha..., 1986 | | | Onthophagus kindianus Frey, 1953 | | | Onthophagus kingstoni Cambefort, 1980 | | | Onthophagus kirgisicus (Protzenko, 1963) | ⇒ Onthophagus silus Balthasar, 1960 | | Onthophagus kirki Frey, 1975 | | | Onthophagus kirokanus Frey, 1975 | | | Onthophagus kivuensis Walter, 2018 | | | Onthophagus kiyoshii Ochi & Kon, 2007 | | | Onthophagus klapperichi Balthasar, 1952 | | | Onthophagus kleinei Balthasar, 1935 | | | Onthophagus knapperti Krikken, 1981 | | | Onthophagus kochi Frey, 1957 | | | Onthophagus koechlei Masumoto, Oc..., 2011 | | | Onthophagus kogatanus (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus kohlmanni Génier, 2017 | | | Onthophagus kolaka Huijbregts &..., 2009 | | | Onthophagus koma Matsumura, 1937 | | | Onthophagus komabellus Matsumura & ..., 1937 | | | Onthophagus komareki Balthasar, 1935 | | | Onthophagus kongkaewensis Masumoto, Oc..., 2012 | | | Onthophagus koni Ochi, 2007 | | | Onthophagus konsarnensis Masumoto, Oc..., 2008 | | | Onthophagus kontumicus Kabakov, 1983 | | | Onthophagus kouassii Cambefort, 1984 | | | Onthophagus kozunonis (Matsumura, 1937) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus krali (Balthasar, 1963) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus kudrnaianus Josso, 2022 | | | Onthophagus kulti Balthasar, 1952 | | | Onthophagus kuluensis Bates, 1891 | | | Onthophagus kulzeri Frey, 1958 | | | Onthophagus kumaonensis Arrow, 1931 | | | Onthophagus kyleensis Frey, 1975 | | | Onthophagus kyokoae Masumoto & Ochi, 2015 | | | Onthophagus laborans Arrow, 1931 | | | Onthophagus laceratus (Gerstaecker, 1871) | ⇒ Onthophagus depressus Harold, 1871 | | Onthophagus lacustris Harold, 1877 | | | Onthophagus laetus d´Orbigny, 1913 | | | Onthophagus laevatus d´Orbigny, 1902 | | | Onthophagus laeviceps d´Orbigny, 1902 | | | Onthophagus laevicollis (Lansberge, 1883) | ⇒ Parascatonomus laevicollis Lansberge, 1883 | | Onthophagus laevidorsis d´Orbigny, 1907 | | | Onthophagus laevigatus Fabricius, 1798 | | | Onthophagus laevissimus d´Orbigny, 1905 | | | Onthophagus lagnyi Paulian, 1937 | | | Onthophagus lahorensis Kabakov, 2008 | | | Onthophagus laichauensis Ochi, Kon & ..., 2021 | | | Onthophagus lamellatus (Boucomont, 1924) | ⇒ Onthophagus lunatus Harold, 1868 | | Onthophagus lamellicornis Frey, 1953 | | | Onthophagus lamelliger (Gerstaecker, 1871) | ⇒ Onthophagus ochropygus d´Orbigny, 1902 | | Onthophagus laminatus (MacLeay, 1864) | ⇒ Onthophagus lunatus Harold, 1868 | | Onthophagus laminicornis (Lansberge, 1886) | ⇒ Onthophagus eulaminicornis Pittino, 2006 | | Onthophagus laminidorsis d´Orbigny, 1902 | | | Onthophagus laminiger Mulsant, 1842 | | | Onthophagus laminosus d´Orbigny, 1905 | | | Onthophagus lamnifer d´Orbigny, 1902 | | | Onthophagus lamotellus Cambefort, 1980 | | | Onthophagus lamottei Frey, 1962 | | | Onthophagus lampromelas (Fairmaire, 1891) | ⇒ Onthophagus laevis lampromelas Fairmaire, 1891 | | Onthophagus lamtoensis Cambefort, 1984 | | | Onthophagus lamtoi Cambefort, 1980 | | | Onthophagus lamyi Paulian, 1945 | | | Onthophagus langkawiensis Ochi & Kon, 2015 | | | Onthophagus lapillus Arrow, 1931 | | | Onthophagus laratinus Arrow, 1916 | | | Onthophagus lassulus Balthasar, 1964 | | | Onthophagus latepunctatus d´Orbigny, 1913 | | | Onthophagus latestriatus d´Orbigny, 1908 | | | Onthophagus latevittatus d´Orbigny, 1905 | | | Onthophagus latigibber d´Orbigny, 1902 | | | Onthophagus latipennis d´Orbigny, 1897 | | | Onthophagus latiplaga (Reitter, 1897) | ⇒ Onthophagus truchmenus Kolenati, 1846 | | Onthophagus latissimus d´Orbigny, 1902 | | | Onthophagus leanus Goidanich, 1926 | | | Onthophagus lebasi Boucomont, 1932 | | | Onthophagus lefebvrei d´Orbigny, 1913 | | | Onthophagus lefiniensis Balthasar, 1967 | | | Onthophagus legendrei Walter & Cam..., 1977 | | | Onthophagus lekmaengtibanphrao Masumoto & Ochi, 2015 | | | Onthophagus leknamnaous Masumoto & Ochi, 2015 | | | Onthophagus lemagneni d´Orbigny, 1902 | | | Onthophagus lemekensis d´Orbigny, 1915 | | | Onthophagus lemniscatus Gillet, 1924 | | | Onthophagus lemuroides d´Orbigny, 1898 | | | Onthophagus lenzii Harold, 1874 | | | Onthophagus leomontanus Boucomont, 1914 | | | Onthophagus leroyi d´Orbigny, 1902 | | | Onthophagus leucomelas (Solsky, 1876) | ⇒ Onthophagus leucostigma Steven, 1811 | | Onthophagus leucopygus (Harold, 1867) | ⇒ Onthophagus albipodex d´Orbigny, 1902 | | Onthophagus leusermontis Huijbregts &..., 2011 | | | Onthophagus liberianus Lansberge, 1883 | | | Onthophagus lilongweensis Dierkens, 2022 | | | Onthophagus limonensis Kohlmann & S..., 2001 | | | Onthophagus lindu Huijbregts &..., 2009 | | | Onthophagus lineatus (Mulsant, 1842) | ⇒ Onthophagus transcaspicus Koenig, 1888 | | Onthophagus lineolatus (Mulsant, 1842) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus liodermus d´Orbigny, 1913 | | | Onthophagus lioides d´Orbigny, 1902 | | | Onthophagus liopterus Harold, 1880 | | | Onthophagus liothorax Koshantschikov, 1894 | | | Onthophagus lituratus Roth, 1851 | | | Onthophagus liui Masumoto, Oc..., 2014 | | | Onthophagus liwagensis Ochi & Kon, 2006 | | | Onthophagus liwondeensis Josso, 2019 | | | Onthophagus lobi Cambefort, 1984 | | | Onthophagus lojanus Balthasar, 1939 | | | Onthophagus lomii Müller, 1942 | | | Onthophagus longecarinatus Zunino & Hal..., 1988 | | | Onthophagus longegranus Frey, 1957 | | | Onthophagus longiceps d´Orbigny, 1904 | | | Onthophagus longimanus Bates, 1887 | | | Onthophagus longipilis d´Orbigny, 1905 | | | Onthophagus lopeensis Josso, 2022 | | | Onthophagus lore Huijbregts &..., 2009 | | | Onthophagus loricatus Klug, 1855 | | | Onthophagus loroi Balthasar, 1941 | | | Onthophagus loudetiae Cambefort, 1984 | | | Onthophagus loveni Gillet, 1928 | | | Onthophagus loxodontae Cambefort, 1984 | | | Onthophagus luctuosus Boucomont, 1914 | | | Onthophagus ludio Boucomont, 1914 | | | Onthophagus lugens (Fahraeus, 1857) | ⇒ Onthophagus fimetarius Roth, 1851 | | Onthophagus lugubris (Fahraeus, 1857) | ⇒ Onthophagus lituratus Roth, 1851 | | Onthophagus luismargaritorum Delgado, 1995 | | | Onthophagus lumareti Walter & Cam..., 1977 | | | Onthophagus lunatus Harold, 1868 | | | Onthophagus lunulifer Boucomont, 1914 | | | Onthophagus luridus (Paulian, 1933) | ⇒ Onthophagus rectecornutus Lansberge, 1883 | | Onthophagus lusitanica (Seabra, 1907) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus lutaticollis d´Orbigny, 1907 | | | Onthophagus luteipennis Wiedemann, 1823 | | | Onthophagus luteosignatus Lansberge, 1883 | | | Onthophagus luzonicus (Lansberge, 1883) | ⇒ Onthophagus armatus Blanchard, 1853 | | Onthophagus luzonwallacei Ochi, Kawaha..., 2021 | | | Onthophagus mabuensis Daniel, Strü..., 2023 | | | Onthophagus macroliberianus Moretto, 2010 | | | Onthophagus macrothorax d´Orbigny, 1902 | | | Onthophagus maculosus d´Orbigny, 1908 | | | Onthophagus madoqua Arrow, 1931 | | | Onthophagus maephaluangus Masumoto, Oc..., 2007 | | | Onthophagus maesalongensis Masumoto, Oc..., 2013 | | | Onthophagus magnini Paulian, 1933 | | | Onthophagus magnioculus Ochi & Kon, 2006 | | | Onthophagus magnipygus Boucomont, 1914 | | | Onthophagus makokou Walter, 1989 | | | Onthophagus malabarensis Boucomont, 1919 | | | Onthophagus malangensis Boucomont, 1914 | | | Onthophagus malasiacus Gillet, 1927 | | | Onthophagus malevolus Balthasar, 1964 | | | Onthophagus manantlanensis Sánchez-Huer..., 2021 | | | Onthophagus mandarinus (Harold, 1859) | ⇒ Onthophagus tragus Fabricius, 1792 | | Onthophagus manguliensis Boucomont, 1914 | | | Onthophagus mankonoensis (Balthasar, 1966) | ⇒ Onthophagus mocquerysi d´Orbigny, 1902 | | Onthophagus marahouensis Cambefort, 1984 | | | Onthophagus marani (Vsetecka, 1942) | ⇒ Parascatonomus acuticollis Gillet, 1927 | | Onthophagus marginatus (Castelnau, 1840) | ⇒ Onthophagus andalusicus Waltl, 1835 | | Onthophagus marginicollis Harold, 1880 | | | Onthophagus marginifer Frey, 1953 | | | Onthophagus marginipennis Josso, 2018 | | | Onthophagus marginithorax (Vsetecka, 1939) | ⇒ Onthophagus lenzi Harold, 1875 | | Onthophagus mariozuninoi Delgado, Nav..., 1993 | | | Onthophagus marmoratus (Ménétriés, 1832) | ⇒ Onthophagus marginalis Gebler, 1817 | | Onthophagus marshalli d´Orbigny, 1908 | | | Onthophagus martelli Frey, 1972 | | | Onthophagus martellii Frey, 1972 | | | Onthophagus martialis Boucomont, 1914 | | | Onthophagus martinpierai Moctezuma, R..., 2016 | | | Onthophagus maruchanus Masumoto, Oc..., 2012 | | | Onthophagus maruyamai Ochi, Kon & ..., 2014 | | | Onthophagus marxi (Splichal, 1910) | ⇒ Onthophagus sutleinensis Splichal, 1910 | | Onthophagus maryatiae Ochi & Kon, 2005 | | | Onthophagus masaicus d´Orbigny, 1905 | | | Onthophagus matae Cambefort, 1984 | | | Onthophagus matanyo Huijbregts &..., 2009 | | | Onthophagus mateui Moctezuma & ..., 2020 | | | Onthophagus matsuii Ochi & Kon, 2006 | | | Onthophagus matsuokai (Matsumura & ..., 1937) | ⇒ Onthophagus sauteri Gillet, 1924 | | Onthophagus mauritii Boucomont, 1919 | | | Onthophagus maurus (Lucas, 1846) | ⇒ Onthophagus andalusicus Waltl, 1835 | | Onthophagus maxwellianus Moretto, 2013 | | | Onthophagus maya Zunino, 1981 | | | Onthophagus mediofasciatus (d´Orbigny, 1900) | ⇒ Onthophagus nitidulus Klug, 1845 | | Onthophagus mediofuscatus d´Orbigny, 1908 | | | Onthophagus megapacificus Ochi & Kon, 2006 | | | Onthophagus megathorax Gillet, 1921 | | | Onthophagus mekara Huijbregts &..., 2009 | | | Onthophagus melanocephalus Klug, 1845 | | | Onthophagus mendax (Mulsant, 1842) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus menetriesii (Faldermann, 1835) | ⇒ Onthophagus illyricus Scopoli, 1763 | | Onthophagus menkaoensis Walter & Cam..., 1977 | | | Onthophagus mentaveiensis Boucomont, 1914 | | | Onthophagus merdrignaci Walter & Cam..., 1977 | | | Onthophagus meruanus d´Orbigny, 1908 | | | Onthophagus merus Péringuey, 1901 | | | Onthophagus metalliger d´Orbigny, 1913 | | | Onthophagus mexicanus Bates, 1887 | | | Onthophagus micellus Matsumura, 1937 | | | Onthophagus micropterus Zunino & Hal..., 1981 | | | Onthophagus micros d´Orbigny, 1902 | | | Onthophagus midorianus (Matsumura, 1938) | ⇒ Parascatonomus anguicorius Boucomont, 1914 | | Onthophagus miichunus Masumoto, Oc..., 2020 | | | Onthophagus miles d´Orbigny, 1905 | | | Onthophagus militaris Boucomont, 1914 | | | Onthophagus millingeni d´Orbigny, 1900 | | | Onthophagus mimikanus Balthasar, 1969 | | | Onthophagus mimus d´Orbigny, 1913 | | | Onthophagus minans d´Orbigny, 1913 | | | Onthophagus minax d´Orbigny, 1904 | | | Onthophagus mindanaensis Boucomont, 1914 | | | Onthophagus mindanaoensis Frey, 1971 | | | Onthophagus minettianus Josso, 2019 | | | Onthophagus minoi Matsumura & ..., 1937 | | | Onthophagus minokuchianus (Matsumura, 1934) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus minutissimus d´Orbigny, 1908 | | | Onthophagus minutulus Harold, 1875 | | | Onthophagus minutus (Hausmann, 1807) | ⇒ Onthophagus centricornis Fabricius, 1798 | | Onthophagus mirabilis Bates, 1887 | | | Onthophagus mirandus Arrow, 1931 | | | Onthophagus miricollis Frey, 1962 | | | Onthophagus miricornis d´Orbigny, 1902 | | | Onthophagus mirifrons d´Orbigny, 1905 | | | Onthophagus miscellaneus d´Orbigny, 1908 | | | Onthophagus miscellus d´Orbigny, 1905 | | | Onthophagus misellus d´Orbigny, 1907 | | | Onthophagus missor (Balthasar, 1939) | ⇒ Onthophagus carinensis Boucomont, 1914 | | Onthophagus misujianus Matsumura & ..., 1937 | | | Onthophagus mixticeps d´Orbigny, 1905 | | | Onthophagus mixtidorsis d´Orbigny, 1905 | | | Onthophagus miyabei (Matsumura, 1937) | ⇒ Onthophagus punctator Reitter, 1893 | | Onthophagus miyazakianus Matsumura & ..., 1937 | | | Onthophagus mniszekii Hochhuth, 1873 | | | Onthophagus moajat Huijbregts &..., 2009 | | | Onthophagus mocquerysi d´Orbigny, 1902 | | | Onthophagus modestus Harold, 1862 | | | Onthophagus moerens Walker, 1858 | | | Onthophagus mogo Krikken & Hu..., 2008 | | | Onthophagus monardi (Boucomont, 1936) | ⇒ Onthophagus monardiellus Frey, 1963 | | Onthophagus monardiellus Frey, 1963 | | | Onthophagus monodon Fahraeus, 1857 | | | Onthophagus monticolus (Howden & Car..., 1963) | ⇒ Onthophagus durangoensis Balthasar, 1939 | | Onthophagus montishannoniae Krikken & Hu..., 2008 | | | Onthophagus montivagus d´Orbigny, 1902 | | | Onthophagus montreuili Moretto & Gé..., 2010 | | | Onthophagus mopsus Fabricius, 1792 | | | Onthophagus mordindangus Masumoto, Oc..., 2008 | | | Onthophagus moreleti Baraud, 1980 | | | Onthophagus morenoi Moretto, 2013 | | | Onthophagus morettoi Josso, 2022 | | | Onthophagus morimotoi Masumoto, Oc..., 2021 | | | Onthophagus morio (Brullé, 1832) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus moroni Zunino & Hal..., 1988 | | | Onthophagus morosus Gerstaecker, 1871 | | | Onthophagus mpassa Walter, 1982 | | | Onthophagus mucronatus (Thomson, 1858) | ⇒ Pseudosaproecius validicornis Quedenfeldt, 1884 | | Onthophagus mucronifer d´Orbigny, 1905 | | | Onthophagus mugheseensis Josso, 2019 | | | Onthophagus multipunctatus Boucomont, 1914 | | | Onthophagus mulutagal Ochi, Kon & ..., 2016 | | | Onthophagus murphyi Josso, 2019 | | | Onthophagus musculus Frey, 1963 | | | Onthophagus musculus ganganensis Frey, 1963 | | | Onthophagus musculus musculus | | | Onthophagus mushana Matsumura, 1938 | | | Onthophagus mushensis Matsumura, 1938 | | | Onthophagus mutabilis (Mulsant, 1842) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus mutatus Harold, 1859 | | | Onthophagus mwenseensis Dierkens, 2022 | | | Onthophagus mycetorum Zunino & Hal..., 1988 | | | Onthophagus nabeleki Balthasar, 1939 | | | Onthophagus naevius d´Orbigny, 1913 | | | Onthophagus naevuliger d´Orbigny, 1908 | | | Onthophagus nagasawai Matsumura, 1938 | | | Onthophagus nagpurensis Arrow, 1931 | | | Onthophagus nakatomii (Matsumura, 1937) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus namnaolekus Masumoto, Oc..., 2020 | | | Onthophagus namnaonus Masumoto, Oc..., 2020 | | | Onthophagus namnaous Masumoto, Oc..., 2013 | | | Onthophagus nanus (Harold, 1878) | ⇒ Onthophagus waterhousei Gillet, 1927 | | Onthophagus narashinensis Matsumura & ..., 1936 | | | Onthophagus nasalis Arrow, 1931 | | | Onthophagus nasicornis Harold, 1869 | | | Onthophagus naso Fahraeus, 1857 | | | Onthophagus nasutus Guérin-Ménev..., 1855 | | | Onthophagus natronis Cambefort, 1984 | | | Onthophagus navarretorum Delgado & Ca..., 1996 | | | Onthophagus naviculifer Boucomont, 1919 | | | Onthophagus nchilaensis Josso, 2022 | | | Onthophagus neavei d´Orbigny, 1914 | | | Onthophagus nebulosus Reiche, 1864 | | | Onthophagus necessarius Reitter, 1893 | | | Onthophagus necrophagus Arrow, 1931 | | | Onthophagus neervoorti Boucomont, 1914 | | | Onthophagus nefarius Balthasar, 1941 | | | Onthophagus negligens Walker, 1858 | | | Onthophagus nemorivagus Kohlmann & S..., 2001 | | | Onthophagus neofurcatus Goidanich, 1926 | | | Onthophagus neofuscus Moctezuma & ..., 2020 | | | Onthophagus neomirabilis Howden, 1973 | | | Onthophagus neostenocerus Goidanich, 1926 | | | Onthophagus neptunulus (Lansberge, 1883) | ⇒ Onthophagus tragus Fabricius, 1792 | | Onthophagus neyo Moretto, 2013 | | | Onthophagus nicobaricus Biswas, Chat..., 1999 | | | Onthophagus nigerianus d´Orbigny, 1913 | | | Onthophagus nigriceps Raffray, 1877 | | | Onthophagus nigrimargo (Goidanich, 1926) | ⇒ Onthophagus marginalis Gebler, 1817 | | Onthophagus nigrinus Paulian, 1937 | | | Onthophagus nigripennis d´Orbigny, 1908 | | | Onthophagus nigriventris d´Orbigny, 1902 | | | Onthophagus nigrivestis (d´Orbigny, 1902) | ⇒ Onthophagus laeviceps d´Orbigny, 1902 | | Onthophagus nigroflavus Josso, 2022 | | | Onthophagus nigropubens d´Orbigny, 1908 | | | Onthophagus nigroviolaceus d´Orbigny, 1902 | | | Onthophagus nigrovirescens (Mulsant, 1842) | ⇒ Euonthophagus amyntas Olivier, 1789 | | Onthophagus nilicola d´Orbigny, 1902 | | | Onthophagus niloticus Harold, 1879 | | | Onthophagus nimbatus d´Orbigny, 1905 | | | Onthophagus niokolokoba Moretto, 2007 | | | Onthophagus nitefactus Harold, 1877 | | | Onthophagus nitidicollis (Brullé, 1832) | ⇒ Onthophagus lucidus Illiger, 1800 | | Onthophagus nitidifrons d´Orbigny, 1905 | | | Onthophagus nitidior (Bates, 1887) | ⇒ Onthophagus blackburni Shipp, 1895 | | Onthophagus nitidulus Klug, 1845 | | | Onthophagus nitidus Waterhouse, 1875 | | | Onthophagus noctivagus (Harold, 1877) | ⇒ Onthophagus reticulatus d´Orbigny, 1902 | | Onthophagus nocturnus (Reitter, 1893) | ⇒ Onthophagus sibiricus Harold, 1877 | | Onthophagus nodieri d´Orbigny, 1908 | | | Onthophagus nonstriatus d´Orbigny, 1905 | | | Onthophagus notatus d´Orbigny, 1902 | | | Onthophagus notiodes Solis & Kohl..., 2003 | | | Onthophagus novemmaculatus Gmelin, 1788 | | | Onthophagus nubilus Kohlmann & S..., 2001 | | | Onthophagus nuchidens Fabricius, 1798 | | | Onthophagus nudatus d´Orbigny, 1913 | | | Onthophagus nudifrons Balthasar, 1939 | | | Onthophagus nudus d´Orbigny, 1908 | | | Onthophagus numidicus d´Orbigny, 1908 | | | Onthophagus nummelini Moretto & Gé..., 2020 | | | Onthophagus nutans (Roth, 1851) | ⇒ Onthophagus schaufussi Harold, 1867 | | Onthophagus nyctopus Bates, 1887 | | | Onthophagus nyikaensis Josso, 2018 | | | Onthophagus oaxacanus Zunino & Hal..., 1988 | | | Onthophagus obenbergeri Balthasar, 1965 | | | Onthophagus oberthueri d´Orbigny, 1898 | | | Onthophagus oberthuri d´Orbigny, 1898 | | | Onthophagus obesus (Fahraeus, 1857) | ⇒ Onthophagus vinctus Erichson, 1843 | | Onthophagus obliviosus Balthasar, 1935 | | | Onthophagus obscuratus (Heyden, 1890) | ⇒ Onthophagus maki Illiger, 1803 | | Onthophagus obscurior Boucomont, 1914 | | | Onthophagus obsoletus (Fairmaire, 1875) | ⇒ Onthophagus transcaspicus Koenig, 1888 | | Onthophagus obtusicornis (Fahraeus, 1857) | ⇒ Onthophagus marginalis przewalski Kabakov, 2006 | | Onthophagus obtusus Wiedemann, 1819 | | | Onthophagus obtutus Péringuey, 1901 | | | Onthophagus ocellatopunctatus Waterhouse, 1875 | | | Onthophagus ocellidorsis d´Orbigny, 1915 | | | Onthophagus ocellifer d´Orbigny, 1902 | | | Onthophagus ochii Masumoto, 1988 | | | Onthophagus ochreatus d´Orbigny, 1897 | | | Onthophagus ochromerus Harold, 1877 | | | Onthophagus ochropygus d´Orbigny, 1902 | | | Onthophagus octogonus Frey, 1962 | | | Onthophagus oculatus Arrow, 1931 | | | Onthophagus ohbayashii Nomura, 1939 | | | Onthophagus ohkuboi Ochi & Kon, 2006 | | | Onthophagus oishii Matsumura & ..., 1937 | | | Onthophagus okahandjanus Balthasar, 1974 | | | Onthophagus okinawanus Nomura, 1976 | | | Onthophagus okushirianus Matsumura, 1937 | | | Onthophagus olidus Balthasar, 1964 | | | Onthophagus omiltemius (Bates, 1887) | ⇒ Onthophagus chevrolati omiltemius Bates, 1887 | | Onthophagus omostigma d´Orbigny, 1902 | | | Onthophagus oniellus (Matsumura, 1937) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus onorei Zunino & Hal..., 1997 | | | Onthophagus onthochromus Arrow, 1913 | | | Onthophagus ontosatu Huijbregts &..., 2012 | | | Onthophagus opacihartiniae Ochi & Kon, 2015 | | | Onthophagus ophion Erichson, 1847 | | | Onthophagus ophion confusus Boucomont, 1932 | | | Onthophagus ophtalmicus Frey, 1958 | | | Onthophagus orbus Boucomont, 1924 | | | Onthophagus orissanus Arrow, 1931 | | | Onthophagus orizabensis Moctezuma, J..., 2019 | | | Onthophagus ornatulus d´Orbigny, 1908 | | | Onthophagus orphnoides (Bates, 1887) | ⇒ Onthophagus mirabilis Bates, 1887 | | Onthophagus orthocerus Thomson, 1858 | | | Onthophagus oryx Fabricius, 1792 | | | Onthophagus osculatii Guérin-Ménev..., 1855 | | | Onthophagus osellai Pittino, 1982 | | | Onthophagus oshimanus Nakane, 1960 | | | Onthophagus otai Ochi & Kon, 2006 | | | Onthophagus otjivarongus Balthasar, 1967 | | | Onthophagus ovigranosus Frey, 1971 | | | Onthophagus ovulum Gerstaecker, 1871 | | | Onthophagus pacificus Lansberge, 1885 | | | Onthophagus pactolus Fabricius, 1787 | | | Onthophagus padrinoi Delgado, 1999 | | | Onthophagus palamoui Biswas & Cha..., 1986 | | | Onthophagus palatus Boucomont, 1914 | | | Onthophagus palavanus (Huijbregts &..., 2011) | ⇒ Onthophagus javanensis Balthasar, 1963 | | Onthophagus palawantagal Ochi, Kon & ..., 2016 | | | Onthophagus pallens d´Orbigny, 1908 | | | Onthophagus pallidipennis (Fahraeus, 1857) | ⇒ Onthophagus rasipennis d´Orbigny, 1908 | | Onthophagus pallidus d´Orbigny, 1905 | | | Onthophagus pangmaphanus Masumoto, Oc..., 2022 | | | Onthophagus paracenricornis Ochi & Kon, 2014 | | | Onthophagus paracentricornis Ochi & Kon, 2014 | | | Onthophagus parafasciatus Balthasar, 1974 | | | Onthophagus parafuscus Zunino & Hal..., 2005 | | | Onthophagus parapalatus Krikken & Hu..., 1988 | | | Onthophagus paravinctus Tagliaferri ..., 2012 | | | Onthophagus parceguttatus d´Orbigny, 1902 | | | Onthophagus parcenotatus d´Orbigny, 1913 | | | Onthophagus parcepictus d´Orbigny, 1908 | | | Onthophagus parcepilosus Balthasar, 1966 | | | Onthophagus pardalis Fabricius, 1798 | | | Onthophagus parenthesis Boucomont, 1913 | | | Onthophagus parisii (Balthasar, 1941) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus paroculus Krikken & Hu..., 1987 | | | Onthophagus parryi Harold, 1869 | | | Onthophagus parumnotatus Fahraeus, 1857 | | | Onthophagus parvidens (Frey, 1971) | ⇒ Onthophagus clermonti Paulian, 1931 | | Onthophagus parvifrons (d´Orbigny, 1907) | ⇒ Onthophagus semiasper d´Orbigny, 1902 | | Onthophagus parvulus (Fabricius, 1798) | ⇒ Onthophagus frontalis Raffray, 1877 | | Onthophagus pasotagal Ochi, Kon & ..., 2016 | | | Onthophagus pastillatus Boucomont, 1919 | | | Onthophagus paucigranosus Lansberge, 1886 | | | Onthophagus paulianellus Frey, 1958 | | | Onthophagus pauliani (Balthasar, 1937) | ⇒ Onthophagus paulianellus Frey, 1958 | | Onthophagus pauper Boucomont, 1914 | | | Onthophagus pauperatus Mulsant, 1842 | | | Onthophagus pauxillus d´Orbigny, 1902 | | | Onthophagus pavidus Harold, 1877 | | | Onthophagus pedator Sharp, 1875 | | | Onthophagus pedester Howden & Gén..., 2004 | | | Onthophagus peguanus (Boucomont, 1914) | ⇒ Onthophagus pacificus Lansberge, 1885 | | Onthophagus pendjarius Josso & Prév..., 2006 | | | Onthophagus peninsularis Boucomont, 1914 | | | Onthophagus peninsulomerus Huijbregts &..., 2011 | | | Onthophagus perakensis Ochi, Kon & ..., 2014 | | | Onthophagus peringueyi Shipp, 1895 | | | Onthophagus perniger Boucomont, 1930 | | | Onthophagus persianus (Olsoufieff, 1900) | ⇒ Onthophagus andalusicus persianus Olsoufieff, 1900 | | Onthophagus personatus Boucomont, 1914 | | | Onthophagus petenensis Howden & Gill, 1993 | | | Onthophagus phahompokus Masumoto & Ochi, 2015 | | | Onthophagus phakuaiensis Masumoto, Oc..., 2013 | | | Onthophagus phanaeicollis Lansberge, 1883 | | | Onthophagus phanaeides Frey, 1956 | | | Onthophagus phanaeiformis Boucomont, 1914 | | | Onthophagus phanaeomorphus Janssens, 1954 | | | Onthophagus phatoensis Masumoto, Oc..., 2008 | | | Onthophagus philippemorettoi Josso, 2022 | | | Onthophagus philippinensis Boucomont, 1919 | | | Onthophagus phillippsorum Krikken & Hu..., 1987 | | | Onthophagus phukhieoensis Masumoto, Oc..., 2007 | | | Onthophagus picatus d´Orbigny, 1902 | | | Onthophagus piceiceps d´Orbigny, 1904 | | | Onthophagus pictipennis d´Orbigny, 1913 | | | Onthophagus pictipodex d´Orbigny, 1902 | | | Onthophagus picturatus d´Orbigny, 1908 | | | Onthophagus pictus (Reitter, 1893) | ⇒ Oniticellus tessellatus Harold, 1879 | | Onthophagus pierrei Josso, 2022 | | | Onthophagus piffli Petrovitz, 1961 | | | Onthophagus pilauco Tello, Verdú..., 2021 | | | Onthophagus pilicollis d´Orbigny, 1902 | | | Onthophagus piliger (Mulsant, 1842) | ⇒ Onthophagus illyricus Scopoli, 1763 | | Onthophagus pilipodex d´Orbigny, 1913 | | | Onthophagus pilosus (Fahraeus, 1857) | ⇒ Onthophagus echinus Boucomont, 1914 | | Onthophagus pimpasaleei Masumoto, Oc..., 2008 | | | Onthophagus pinguis Gerstaecker, 1871 | | | Onthophagus pisciphagus Walter & Cam..., 1977 | | | Onthophagus pithankithae Karimbumkara..., 2016 | | | Onthophagus placens Péringuey, 1904 | | | Onthophagus planaticeps d´Orbigny, 1913 | | | Onthophagus planiceps (MacLeay, 1886) | ⇒ Onthophagus pinguis Gerstaecker, 1871 | | Onthophagus planicornis Herbst, 1789 | | | Onthophagus planidorsis Dierkens, 2022 | | | Onthophagus planifrons Frey, 1962 | | | Onthophagus pleurogonus d´Orbigny, 1913 | | | Onthophagus plicatifrons d´Orbigny, 1908 | | | Onthophagus pljushtchi Ivanova, 2012 | | | Onthophagus plurimacula (Depoli, 1924) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus poeophagus Kabakov, 1979 | | | Onthophagus politissimus d´Orbigny, 1908 | | | Onthophagus pollicatus Harold, 1879 | | | Onthophagus polyedrus d´Orbigny, 1905 | | | Onthophagus polyodon d´Orbigny, 1913 | | | Onthophagus polystigma d´Orbigny, 1902 | | | Onthophagus pooensis Cambefort & ..., 1991 | | | Onthophagus porcus Arrow, 1931 | | | Onthophagus poringensis Ochi & Kon, 2005 | | | Onthophagus possoi Walter, 1982 | | | Onthophagus posticicornis d´Orbigny, 1913 | | | Onthophagus potosinus Moctezuma & ..., 2020 | | | Onthophagus praecellens Bates, 1887 | | | Onthophagus praedatus Harold, 1862 | | | Onthophagus praelaminatus Frey, 1957 | | | Onthophagus praestans Péringuey, 1901 | | | Onthophagus praetortus Péringuey, 1904 | | | Onthophagus probus Péringuey, 1901 | | | Onthophagus procurvus Balthasar, 1935 | | | Onthophagus producticollis d´Orbigny, 1908 | | | Onthophagus productus Arrow, 1907 | | | Onthophagus profanus Frey, 1958 | | | Onthophagus prolixus (Walker, 1858) | ⇒ Onthophagus unifasciatus Schaller, 1783 | | Onthophagus pronocarinatus Josso, 2019 | | | Onthophagus propinquus (MacLeay, 1888) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus propraecellens Howden & Gill, 1987 | | | Onthophagus proteus d´Orbigny, 1902 | | | Onthophagus protuberans Frey, 1955 | | | Onthophagus proximus d´Orbigny, 1908 | | | Onthophagus pseudoaeneus d´Orbigny, 1908 | | | Onthophagus pseudoarai Ochi & Kon, 2018 | | | Onthophagus pseudobidens d´Orbigny, 1915 | | | Onthophagus pseudobrutus Kabakov, 1983 | | | Onthophagus pseudobufonidius Josso, 2019 | | | Onthophagus pseudoconvexicollis Bai & Yang, 2016 | | | Onthophagus pseudocoracinus Kabakov, 1983 | | | Onthophagus pseudocribripennis Josso, 2019 | | | Onthophagus pseudofimetarius Balthasar, 1946 | | | Onthophagus pseudoguatemalensis Moctezuma & ..., 2021 | | | Onthophagus pseudojaponicus Balthasar, 1941 | | | Onthophagus pseudojaponicus fukiensis Balthasar, 1941 | | | Onthophagus pseudojaponicus pseudojaponicus | | | Onthophagus pseudoliberianus Moretto, 2010 | | | Onthophagus pseudopilosus Frey, 1958 | | | Onthophagus pseudosanguineus Walter & Cam..., 1977 | | | Onthophagus pseudosellatus Balthasar, 1964 | | | Onthophagus pseudoundulans Zunino & Hal..., 1988 | | | Onthophagus pseudoundulans howdeni Zunino & Hal..., 1988 | | | Onthophagus pseudovinctus Tagliaferri ..., 2012 | | | Onthophagus ptox Erichson, 1842 | | | Onthophagus puberulus d´Orbigny, 1902 | | | Onthophagus puetalis Matsumura, 1938 | | | Onthophagus pugionatus Fahraeus, 1857 | | | Onthophagus pulchellus d´Orbigny, 1905 | | | Onthophagus pullatus d´Orbigny, 1905 | | | Onthophagus pullus (Roth, 1851) | ⇒ Onthophagus parenthesis Boucomont, 1913 | | Onthophagus pumilus Kabakov, 2014 | | | Onthophagus punctatominiatus (Goidanich, 1925) | ⇒ Onthophagus lucidus Illiger, 1800 | | Onthophagus pupillatus Kolbe, 1886 | | | Onthophagus purifrons d´Orbigny, 1908 | | | Onthophagus purpurascens Boucomont, 1914 | | | Onthophagus pusillus (Fabricius, 1798) | ⇒ Onthophagus modestus Harold, 1862 | | Onthophagus pusio (Fahraeus, 1857) | ⇒ Onthophagus confertus Péringuey, 1908 | | Onthophagus putealis (Matsumura, 1938) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus pygialis (Thomson, 1858) | ⇒ Onthophagus sellatus Klug, 1845 | | Onthophagus pygidialis Lansberge, 1883 | | | Onthophagus pygmaeus Schaller, 1783 | | | Onthophagus quadraticeps Harold, 1867 | | | Onthophagus quadricallosus d´Orbigny, 1902 | | | Onthophagus quadricornis Fabricius, 1801 | | | Onthophagus quadricuspis d´Orbigny, 1908 | | | Onthophagus quadrilunatus d´Orbigny, 1902 | | | Onthophagus quadrimaculatus Raffray, 1877 | | | Onthophagus quadrinodosus Fahraeus, 1857 | | | Onthophagus quadrinotatus d´Orbigny, 1905 | | | Onthophagus quadripustulatus Fabricius, 1775 | Neuholländische Vierblatter | | Onthophagus quadrituberculatus Laicharting, 1781 | | | Onthophagus quadrum Kugelann, 1792 | | | Onthophagus quangnamensis Ochi, Kon & ..., 2021 | | | Onthophagus quasijohkii Ochi & Kon, 2005 | | | Onthophagus quasitagal Ochi & Kon, 2005 | | | Onthophagus quetzalis Howden & Gill, 1993 | | | Onthophagus quiproquo Moretto & Gé..., 2010 | | | Onthophagus ragazzii d´Orbigny, 1904 | | | Onthophagus ramosellus Bates, 1891 | | | Onthophagus ramosus Wiedemann, 1823 | | | Onthophagus rana Arrow, 1931 | | | Onthophagus ranongensis Masumoto, Oc..., 2008 | | | Onthophagus ranongjohkii Ochi, Kon & ..., 2016 | | | Onthophagus ranunculus Arrow, 1913 | | | Onthophagus rasipennis d´Orbigny, 1908 | | | Onthophagus ratchataniensis Masumoto, Oc..., 2013 | | | Onthophagus rectestriatus d´Orbigny, 1915 | | | Onthophagus recticornis Leske, 1785 | | | Onthophagus rectilamina d´Orbigny, 1902 | | | Onthophagus rectinicornis (Fairmaire, 1878) | ⇒ Parascatonomus tricornis Wiedemann, 1823 | | Onthophagus rectorispauliani Cambefort, 1980 | | | Onthophagus reflexicornis (Redtenbacher, 1867) | ⇒ Onthophagus spinifex Fabricius, 1781 | | Onthophagus refulgens Arrow, 1931 | | | Onthophagus regalis Arrow, 1907 | | | Onthophagus reichardti (Medvedev, 1928) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus remotus Harold, 1879 | | | Onthophagus renejeanneli Dierkens, 2022 | | | Onthophagus reticulatus d´Orbigny, 1902 | | | Onthophagus reticuliger Frey, 1960 | | | Onthophagus reticulundaticeps Josso, 2022 | | | Onthophagus retusus (Harold, 1869) | ⇒ Onthophagus chevrolati retusus Harold, 1869 | | Onthophagus reyesi Zunino & Hal..., 1988 | | | Onthophagus rhinoceros (Melsheimer, 1845) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus rhinocerus (Gomes Alves, 1944) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus rhinolophus Harold, 1869 | | | Onthophagus rhinophyllus Harold, 1868 | | | Onthophagus rhodesianus Frey, 1975 | | | Onthophagus rhynchophorus Péringuey, 1904 | | | Onthophagus ribbei Boucomont, 1914 | | | Onthophagus rizali Ochi, Kawaha..., 2021 | | | Onthophagus robertminettii Josso, 2022 | | | Onthophagus robertopoggii Ochi & Kon, 2006 | | | Onthophagus roessneri Ziani, 2016 | | | Onthophagus rohwedderi Moretto, 2017 | | | Onthophagus rojkoffianus Josso, 2018 | | | Onthophagus rorarius Harold, 1877 | | | Onthophagus rosemarieae Josso, 2022 | | | Onthophagus rosenbergi Krikken & Hu..., 2011 | | | Onthophagus rosenbergi divergens Krikken & Hu..., 2011 | | | Onthophagus rosenbergi rosenbergi | | | Onthophagus rosettae Frey, 1958 | | | Onthophagus rostratus (Harold, 1869) | ⇒ Onthophagus trinominatus Goidanich, 1926 | | Onthophagus rostrifer (Reitter, 1893) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus rotundatus d´Orbigny, 1905 | | | Onthophagus rotundibasis d´Orbigny, 1902 | | | Onthophagus rotundicollis Lansberge, 1883 | | | Onthophagus rougonorum Cambefort, 1984 | | | Onthophagus rouyeri Boucomont, 1914 | | | Onthophagus royi Biswas & Cha..., 1985 | | | Onthophagus ruandanus Frey, 1971 | | | Onthophagus rubefactus (d´Orbigny, 1913) | ⇒ Onthophagus tessulatus Harold, 1871 | | Onthophagus rubellus d´Orbigny, 1908 | | | Onthophagus rubens d´Orbigny, 1902 | | | Onthophagus rubenticollis d´Orbigny, 1902 | | | Onthophagus rubidus d´Orbigny, 1913 | | | Onthophagus rubrescens (Blanchard, 1843) | ⇒ Onthophagus stylocerus Graells, 1851 | | Onthophagus rubricatus d´Orbigny, 1902 | | | Onthophagus rubrimaculatus MacLeay, 1864 | | | Onthophagus rubripennis (Arrow, 1907) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus rubripes (Mulsant, 1842) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus rubromaculatus Kolbe, 1886 | | | Onthophagus rubrotinctus (d´Orbigny, 1900) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus rudis Sharp, 1875 | | | Onthophagus rufescens Bates, 1887 | | | Onthophagus ruficauda Arrow, 1931 | | | Onthophagus rufipes (Ménétriés, 1832) | ⇒ Onthophagus illyricus Scopoli, 1763 | | Onthophagus rufipodex d´Orbigny, 1913 | | | Onthophagus rufobasalis Fairmaire, 1887 | | | Onthophagus rufocastaneus Fairmaire, 1887 | | | Onthophagus rufolimbatus d´Orbigny, 1913 | | | Onthophagus rufonotatus d´Orbigny, 1902 | | | Onthophagus rufopygus Frey, 1957 | | | Onthophagus rufostillans d´Orbigny, 1907 | | | Onthophagus rufovirens d´Orbigny, 1904 | | | Onthophagus rugicollis Harold, 1880 | | | Onthophagus rugidorsis d´Orbigny, 1913 | | | Onthophagus rugosipennis Frey, 1973 | | | Onthophagus rugosissimus d´Orbigny, 1913 | | | Onthophagus rugosus (Kirby, 1818) | ⇒ Onthophagus tenebrosus Harold, 1871 | | Onthophagus rugulipennis Fairmaire, 1887 | | | Onthophagus rutilans (Sharp, 1875) | ⇒ Onthophagus borneensis Harold, 1877 | | Onthophagus rutilans aborneensis Ochi, Kon & ..., 2009 | | | Onthophagus rutilans rutilans | | | Onthophagus rutilipennis Reitter, 1893 | | | Onthophagus sabahensis Ochi & Kon, 2006 | | | Onthophagus sabai Masumoto, Oc..., 2012 | | | Onthophagus sacharovskii Olsoufieff, 1918 | | | Onthophagus sahai Biswas & Cha..., 1986 | | | Onthophagus saharae (Bedel, 1911) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus saiga (Ballion, 1870) | ⇒ Onthophagus pygargus Motschulsky, 1845 | | Onthophagus saigonensis Boucomont, 1924 | | | Onthophagus sakishimanus Nomura, 1976 | | | Onthophagus saksphus Masumoto, Oc..., 2021 | | | Onthophagus saleyeri Lansberge, 1883 | | | Onthophagus salvadorensis Zunino & Hal..., 1988 | | | Onthophagus samarensis Ochi, Kon & ..., 2023 | | | Onthophagus samoengus Masumoto, Oc..., 2008 | | | Onthophagus sancristobalensis Moctezuma & ..., 2020 | | | Onthophagus sanggona Huijbregts &..., 2009 | | | Onthophagus sangirensis Boucomont, 1914 | | | Onthophagus sanguineus d´Orbigny, 1902 | | | Onthophagus sanguinolentus d´Orbigny, 1908 | | | Onthophagus sangwalus Masumoto, Oc..., 2008 | | | Onthophagus sanpabloetlorum Moctezuma & ..., 2020 | | | Onthophagus sansibaricus Harold, 1878 | | | Onthophagus santamariensis Moctezuma, S..., 2020 | | | Onthophagus sarasinorum Krikken & Hu..., 2011 | | | Onthophagus saudiensis Frey, 1962 | | | Onthophagus savanicola Cambefort, 1984 | | | Onthophagus scaber Roth, 1851 | | | Onthophagus scaberrimus d´Orbigny, 1904 | | | Onthophagus scapularis d´Orbigny, 1902 | | | Onthophagus sceptrifer Boucomont, 1924 | | | Onthophagus schaefernai Balthasar, 1935 | | | Onthophagus schatzmayri (Pierotti, 1959) | ⇒ Onthophagus opacicollis Reitter, 1893 | | Onthophagus schaufussi Harold, 1867 | | | Onthophagus schawalleri Scheuern, 1996 | | | Onthophagus schillhammeri Kabakov, 2006 | | | Onthophagus schunckei Paulian, 1936 | | | Onthophagus sciron Balthasar, 1969 | | | Onthophagus scrutator Harold, 1877 | | | Onthophagus sculptilis Gerstaecker, 1871 | | | Onthophagus secundarius Roth, 1851 | | | Onthophagus sellatulus d´Orbigny, 1902 | | | Onthophagus sellatus Klug, 1845 | | | Onthophagus sembeli Krikken & Hu..., 2008 | | | Onthophagus semiaereopictus Ochi, Kawaha..., 2021 | | | Onthophagus semiaratus d´Orbigny, 1902 | | | Onthophagus semiasper d´Orbigny, 1902 | | | Onthophagus semichalcites d´Orbigny, 1902 | | | Onthophagus semicinctus d´Orbigny, 1897 | | | Onthophagus semicroceus d´Orbigny, 1915 | | | Onthophagus semiflavus (Boheman, 1860) | ⇒ Onthophagus flavolimbatus Klug, 1855 | | Onthophagus semigraniger d´Orbigny, 1905 | | | Onthophagus semigranosus Lansberge, 1883 | | | Onthophagus semilaevis d´Orbigny, 1913 | | | Onthophagus seminitens d´Orbigny, 1902 | | | Onthophagus seminitidus d´Orbigny, 1908 | | | Onthophagus semiopacus (Harold, 1869) | ⇒ Onthophagus seminitens d´Orbigny, 1902 | | Onthophagus semipacificus Ochi & Kon, 2006 | | | Onthophagus semiperakensis Ochi, Kon & ..., 2014 | | | Onthophagus semivestitus Paulian, 1937 | | | Onthophagus semivirescens d´Orbigny, 1902 | | | Onthophagus semiviridis d´Orbigny, 1904 | | | Onthophagus semiworoae Ochi, Kon & ..., 2014 | | | Onthophagus senegalensis d´Orbigny, 1902 | | | Onthophagus senescens Péringuey, 1908 | | | Onthophagus senex Boucomont, 1914 | | | Onthophagus seniculus (Fabricius, 1781) | ⇒ Onthophagus senescens Péringuey, 1908 | | Onthophagus sepilokensis Ochi & Kon, 2006 | | | Onthophagus seramicus Huijbregts &..., 2012 | | | Onthophagus serdangensis Lansberge, 1886 | | | Onthophagus sericans Frey, 1975 | | | Onthophagus sericeicollis Boucomont, 1928 | | | Onthophagus serienotatus d´Orbigny, 1902 | | | Onthophagus seseba Krikken & Hu..., 2017 | | | Onthophagus setoculus Krikken & Hu..., 1987 | | | Onthophagus setosus Fahraeus, 1857 | | | Onthophagus sexdentatus Boucomont, 1919 | | | Onthophagus sharpi Harold, 1875 | | | Onthophagus shaykh Montanaro & ..., 2022 | | | Onthophagus shibatai Nakane, 1960 | | | Onthophagus shigeoi Matsumura, 1937 | | | Onthophagus shimba Cambefort, 1980 | | | Onthophagus shinanensis (Matsumura, 1934) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus shinichii Ochi & Kon, 2018 | | | Onthophagus shirakii Nakane, 1960 | | | Onthophagus shoyozanus (Matsumura, 1937) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus shunichii Ochi, Kon & ..., 2021 | | | Onthophagus shurianus Matsumura & ..., 1937 | | | Onthophagus sibela Huijbregts &..., 2012 | | | Onthophagus sibuyanus Boucomont, 1924 | | | Onthophagus sidama Gestro, 1895 | | | Onthophagus sidangeaus Masumoto, Oc..., 2021 | | | Onthophagus sideki Krikken & Hu..., 1987 | | | Onthophagus signatus Fahraeus, 1857 | | | Onthophagus sihkahonoi Huijbregts &..., 2009 | | | Onthophagus sikkimensis Gillet, 1925 | | | Onthophagus silus Balthasar, 1960 | | | Onthophagus simboroni Ochi & Kon, 2006 | | | Onthophagus simillimus d´Orbigny, 1913 | | | Onthophagus simius Reitter, 1893 | | | Onthophagus simoni d´Orbigny, 1902 | | | Onthophagus simplex Raffray, 1877 | | | Onthophagus simpliciceps d´Orbigny, 1905 | | | Onthophagus simplicifrons Reitter, 1893 | | | Onthophagus simulator d´Orbigny, 1905 | | | Onthophagus sinagai Krikken & Hu..., 2008 | | | Onthophagus singulariformis Kohlmann & S..., 2001 | | | Onthophagus sinicus Zhang & Wang, 1997 | | | Onthophagus sinuosicollis d´Orbigny, 1905 | | | Onthophagus sinuosus d´Orbigny, 1913 | | | Onthophagus sipilouensis Cambefort, 1984 | | | Onthophagus sjoestethi d´Orbigny, 1904 | | | Onthophagus skelleyi Sánchez-Huer..., 2018 | | | Onthophagus smeei (Harold, 1875) | ⇒ Proagoderus amplexus Sharp, 1875 | | Onthophagus smeenki Cambefort, 1980 | | | Onthophagus smetanai Balthasar, 1952 | | | Onthophagus snoflaki Balthasar, 1944 | | | Onthophagus sobosanus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus socialis Arrow, 1931 | | | Onthophagus solidus (Gillet, 1927) | ⇒ Onthophagus compactus Arrow, 1933 | | Onthophagus solisi Howden & Gill, 1993 | | | Onthophagus solivagus Harold, 1886 | | | Onthophagus solmani Stebnicka, 1975 | | | Onthophagus somalicola Balthasar, 1941 | | | Onthophagus sonani Miwa, 1930 | | | Onthophagus songsokensis Biswas & Cha..., 1985 | | | Onthophagus sophiae Nicolas, 2006 | | | Onthophagus sopu Krikken & Hu..., 2017 | | | Onthophagus sparsepunctatus Frey, 1956 | | | Onthophagus sparsulus Reitter, 1893 | | | Onthophagus spathatus Boucomont, 1914 | | | Onthophagus spiculatus Boucomont, 1914 | | | Onthophagus spinifer A. G. Olivier, 1789 | | | Onthophagus spinifex Fabricius, 1781 | | | Onthophagus spitiensis (Gillet, 1925) | ⇒ Onthophagus troglodyta Wiedemann, 1823 | | Onthophagus splendidoides Josso, 2013 | | | Onthophagus splendidus Boucomont, 1932 | | | Onthophagus sprecherae Moretto, 2013 | | | Onthophagus spurcatus d´Orbigny, 1905 | | | Onthophagus spurius Matsumura, 1937 | | | Onthophagus stanleyi Moretto & Gé..., 2010 | | | Onthophagus statzi ♰ Krell, 1990 | | | Onthophagus stehliki Balthasar, 1966 | | | Onthophagus steinheili Harold, 1880 | | | Onthophagus stellio Erichson, 1843 | | | Onthophagus stenocerus (Harold, 1867) | ⇒ Onthophagus neostenocerus Goidanich, 1926 | | Onthophagus sternalis Arrow, 1931 | | | Onthophagus sternax Balthasar, 1959 | | | Onthophagus stevensi Arrow, 1931 | | | Onthophagus sticticus Harold, 1867 | | | Onthophagus stictus d´Orbigny, 1913 | | | Onthophagus stigmosus d´Orbigny, 1902 | | | Onthophagus stillatus d´Orbigny, 1904 | | | Onthophagus stockwelli Howden & Young, 1956 | | | Onthophagus stomachosus Krell, 2009 | | | Onthophagus strandi Balthasar, 1935 | | | Onthophagus strictestriatus d´Orbigny, 1913 | | | Onthophagus strigatus (Mulsant, 1842) | ⇒ Onthophagus maki Illiger, 1803 | | Onthophagus stuhlmanni d´Orbigny, 1908 | | | Onthophagus subaenescens (Goidanich, 1926) | ⇒ Onthophagus ovatus Linnaeus, 1767 | | Onthophagus subcancer Howden, 1973 | | | Onthophagus subcinctus d´Orbigny, 1913 | | | Onthophagus subconvexus Seabra, 1907 | | | Onthophagus subcornutus Boucomont, 1914 | | | Onthophagus subcribratus Arrow, 1907 | | | Onthophagus subdivisus d´Orbigny, 1908 | | | Onthophagus subhumeralis d´Orbigny, 1902 | | | Onthophagus sublaminatus Mulsant, 1842 | | | Onthophagus sublineatus (d´Orbigny, 1900) | ⇒ Onthophagus turpidus Reitter, 1887 | | Onthophagus sublineolatus (Mulsant, 1842) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus submarginalis (Sahlberg, 1926) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus subnudus d´Orbigny, 1902 | | | Onthophagus subocellatus d´Orbigny, 1905 | | | Onthophagus subplanus d´Orbigny, 1902 | | | Onthophagus subprominulus (Mulsant, 1842) | ⇒ Onthophagus coenobita Herbst, 1783 | | Onthophagus subrecticornis Mulsant, 1842 | | | Onthophagus subrugosus d´Orbigny, 1905 | | | Onthophagus subsulcatus d´Orbigny, 1908 | | | Onthophagus subulifer d´Orbigny, 1908 | | | Onthophagus suermelii Petrovitz, 1963 | | | Onthophagus suffusus Klug, 1855 | | | Onthophagus sugihartoi Ochi, 2007 | | | Onthophagus sugillatus Klug, 1855 | | | Onthophagus suginokoichii Ochi & Kon, 2008 | | | Onthophagus suillus Arrow, 1931 | | | Onthophagus sulawesiensis Krikken & Hu..., 2011 | | | Onthophagus sulawesijohkii Ochi, Kon & ..., 2016 | | | Onthophagus sulcatulus d´Orbigny, 1907 | | | Onthophagus sulci Balthasar, 1935 | | | Onthophagus sulcipennis d´Orbigny, 1902 | | | Onthophagus sumatramontanus Ochi & Kon, 2008 | | | Onthophagus sumatranus Lansberge, 1883 | | | Onthophagus sumbavensis Boucomont, 1914 | | | Onthophagus sumptuosus Paulian, 1937 | | | Onthophagus sunanicus (Stebnicka, 1973) | ⇒ Onthophagus punctator Reitter, 1893 | | Onthophagus sundanensis Lansberge, 1883 | | | Onthophagus surdus Boucomont, 1925 | | | Onthophagus susterai Balthasar, 1952 | | | Onthophagus sutiliceps d´Orbigny, 1902 | | | Onthophagus sutleinensis Splichal, 1910 | | | Onthophagus suturalis (Péringuey, 1888) | ⇒ Onthophagus trituber Wiedemann, 1823 | | Onthophagus suturatus (Germar, 1824) | ⇒ Onthophagus tarandus Fabricius, 1792 | | Onthophagus sycophanta (Fairmaire, 1887) | ⇒ Euonthophagus gibbosus Scriba, 1790 | | Onthophagus sylvestris Walter & Cam..., 1977 | | | Onthophagus synceri Cambefort, 1984 | | | Onthophagus tabidus Balthasar, 1935 | | | Onthophagus taboranus d´Orbigny, 1902 | | | Onthophagus tacanensis Chamé-Vázque..., 2022 | | | Onthophagus taeniatus Boucomont, 1914 | | | Onthophagus tagal Boucomont, 1924 | | | Onthophagus taiensis Cambefort, 1984 | | | Onthophagus taiheizanus Matsumura, 1937 | | | Onthophagus takabayashii Matsumura, 1937 | | | Onthophagus takaoi Ochi & Kon, 2014 | | | Onthophagus talpa Fahraeus, 1857 | | | Onthophagus tambing Krikken & Hu..., 2011 | | | Onthophagus tanganus d´Orbigny, 1913 | | | Onthophagus taoi Ochi & Kon, 2006 | | | Onthophagus tapirus Sharp, 1877 | | | Onthophagus taprobanus Arrow, 1931 | | | Onthophagus tarascus Zunino & Hal..., 1988 | | | Onthophagus tarascus jaliscensis Zunino & Hal..., 1988 | | | Onthophagus taschenbergi Gillet, 1925 | | | Onthophagus teitanicus d´Orbigny, 1902 | | | Onthophagus temporalis d´Orbigny, 1902 | | | Onthophagus tenax Balthasar, 1964 | | | Onthophagus tenebrosus Harold, 1871 | | | Onthophagus tenuicornis (Klug, 1855) | ⇒ Onthophagus aeruginosus Roth, 1851 | | Onthophagus tenuigraniger d´Orbigny, 1913 | | | Onthophagus tenuistriatus d´Orbigny, 1905 | | | Onthophagus tersicollis Müller, 1947 | | | Onthophagus tersipennis d´Orbigny, 1902 | | | Onthophagus tersus d´Orbigny, 1913 | | | Onthophagus tesseratus d´Orbigny, 1908 | | | Onthophagus tessulatus Harold, 1871 | | | Onthophagus testaceoviolaceus Paulian, 1937 | | | Onthophagus tetraspilus (Fairmaire, 1887) | ⇒ Onthophagus nitidulus Klug, 1845 | | Onthophagus thailaevis Masumoto, Oc..., 2008 | | | Onthophagus thainuaensis Masumoto, Oc..., 2008 | | | Onthophagus tharalithae Karimbumkara..., 2016 | | | Onthophagus thomsoni Bates, 1888 | | | Onthophagus thungphraensis Masumoto, Oc..., 2020 | | | Onthophagus tibetanus Arrow, 1907 | | | Onthophagus timorensis Boucomont, 1914 | | | Onthophagus tiniocelloides Boucomont, 1925 | | | Onthophagus tirapensis Biswas & Cha..., 1985 | | | Onthophagus tnai Nithya & Sabu, 2012 | | | Onthophagus tonkineus Gillet, 1921 | | | Onthophagus tonsus d´Orbigny, 1902 | | | Onthophagus tonywhitteni Krikken & Hu..., 2011 | | | Onthophagus toraut Krikken & Hu..., 2011 | | | Onthophagus totonacus Moctezuma & ..., 2021 | | | Onthophagus totonicapamus Bates, 1887 | | | Onthophagus traginus Frey, 1972 | | | Onthophagus tragoides Boucomont, 1914 | | | Onthophagus transbaicalicus (Csiki, 1901) | ⇒ Onthophagus clitellifer Reitter, 1894 | | Onthophagus transcaspicus Koenig, 1888 | | | Onthophagus transisthmius Howden & Young, 1956 | | | Onthophagus transvestitus Huijbregts &..., 2009 | | | Onthophagus trapezicornis d´Orbigny, 1902 | | | Onthophagus traversii d´Orbigny, 1904 | | | Onthophagus triacanthus Castelnau, 1840 | | | Onthophagus tricariniger d´Orbigny, 1902 | | | Onthophagus triceratops Arrow, 1913 | | | Onthophagus trichopygus d´Orbigny, 1905 | | | Onthophagus tricolor Boucomont, 1914 | | | Onthophagus tricorniger Boheman, 1860 | | | Onthophagus tricuspidus Mulsant, 1842 | | | Onthophagus tridenticeps d´Orbigny, 1902 | | | Onthophagus tridentitibialis Ochi & Kon, 2008 | | | Onthophagus trifidisetis d´Orbigny, 1905 | | | Onthophagus trihastatus (Reitter, 1909) | ⇒ Onthophagus oberthuri d´Orbigny, 1898 | | Onthophagus triimpressus d´Orbigny, 1905 | | | Onthophagus trinodosus Fahraeus, 1857 | | | Onthophagus trinominatus Goidanich, 1926 | | | Onthophagus tripartitus d´Orbigny, 1902 | | | Onthophagus tripolitanus Heyden, 1890 | | | Onthophagus tristis Harold, 1873 | | | Onthophagus trituberculatus Schrank, 1798 | | | Onthophagus triundulatus Balthasar, 1964 | | | Onthophagus trochilus Arrow, 1931 | | | Onthophagus trochiscobius (Kolenati, 1846) | ⇒ Onthophagus suturellus Brullé, 1832 | | Onthophagus troglodyta Wiedemann, 1823 | | | Onthophagus troniceki Balthasar, 1933 | | | Onthophagus tschadensis Balthasar, 1963 | | | Onthophagus tschoffeni d´Orbigny, 1904 | | | Onthophagus tshuapae Balthasar, 1964 | | | Onthophagus tsubakii Ochi & Kon, 2009 | | | Onthophagus tsunoellus Matsumura, 1938 | | | Onthophagus tsutomui Ochi & Kon, 2016 | | | Onthophagus tubericollis (Raffray, 1877) | ⇒ Onthophagus bidens Olivier, 1789 | | Onthophagus tumidulus Gerstaecker, 1871 | | | Onthophagus tungkamangensis Masumoto, Oc..., 2007 | | | Onthophagus tungkamungensis Masumoto, Oc..., 2008 | | | Onthophagus turbatus (Walker, 1858) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus turfanicus Balthasar, 1971 | | | Onthophagus turgidus Kohlmann & S..., 2012 | | | Onthophagus turmalis (Gillet, 1924) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus turneri Boucomont, 1936 | | | Onthophagus turpidus Reitter, 1887 | | | Onthophagus tuzetae Walter & Cam..., 1977 | | | Onthophagus ubaidillahi Krikken & Hu..., 2012 | | | Onthophagus uedai Ochi & Kon, 2006 | | | Onthophagus uedanus (Matsumura, 1937) | ⇒ Onthophagus olsoufieffi Boucomont, 1924 | | Onthophagus uelensis Frey, 1960 | | | Onthophagus ugandaensis Dierkens, 2022 | | | Onthophagus ugandicus Frey, 1975 | | | Onthophagus ugoi Moretto & Gé..., 2020 | | | Onthophagus ukerewensis Balthasar, 1937 | | | Onthophagus ulugombakensis Ochi, Kon & ..., 2014 | | | Onthophagus ulula Balthasar, 1966 | | | Onthophagus umbilicatus d´Orbigny, 1908 | | | Onthophagus umbilicopunctatus (Medvedev, 1959) | ⇒ Onthophagus trispinus Reitter, 1892 | | Onthophagus umbratus d´Orbigny, 1902 | | | Onthophagus umenoi (Matsumura, 1938) | ⇒ Onthophagus trituber Wiedemann, 1823 | | Onthophagus umeonoi Matsumura, 1938 | | | Onthophagus undaticeps d´Orbigny, 1902 | | | Onthophagus undulans Bates, 1887 | | | Onthophagus undulans oaxacanus Zunino & Hal..., 1988 | | | Onthophagus unicarina d´Orbigny, 1904 | | | Onthophagus unidentatus Boucomont, 1927 | | | Onthophagus unifasciatus Schaller, 1783 | | | Onthophagus uniformis Heyden, 1886 | | | Onthophagus unxovi (Bojachew, 1930) | ⇒ Onthophagus fortigibber Reitter, 1909 | | Onthophagus urellus Boucomont, 1919 | | | Onthophagus ursinus d´Orbigny, 1902 | | | Onthophagus ursus Boucomont, 1926 | | | Onthophagus urus (Harold, 1867) | ⇒ Onthophagus urusheeri Krell, 2000 | | Onthophagus urusheeri ♰ Krell, 2000 | | | Onthophagus usambaricus Paulian, 1937 | | | Onthophagus ushiodai (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus usurpator Balthasar, 1959 | | | Onthophagus valentineae Josso, 2019 | | | Onthophagus vanasseni Krikken & Hu..., 2008 | | | Onthophagus vanderblomi Krikken & Hu..., 2008 | | | Onthophagus vaneyeni Frey, 1960 | | | Onthophagus vanofwegeni Krikken & Hu..., 2008 | | | Onthophagus variabilis (Mulsant, 1842) | ⇒ Onthophagus maki Illiger, 1803 | | Onthophagus variatus d´Orbigny, 1902 | | | Onthophagus variegatus (Fabricius, 1798) | ⇒ Onthophagus filicornis Harold, 1873 | | Onthophagus variegranosus d´Orbigny, 1905 | | | Onthophagus variolaris Lansberge, 1883 | | | Onthophagus variolosus d´Orbigny, 1902 | | | Onthophagus varius d´Orbigny, 1913 | | | Onthophagus vassei d´Orbigny, 1908 | | | Onthophagus vatovai Müller, 1942 | | | Onthophagus velliger d´Orbigny, 1907 | | | Onthophagus ventralis Lansberge, 1883 | | | Onthophagus ventrosus d´Orbigny, 1905 | | | Onthophagus venustulus Erichson, 1843 | | | Onthophagus venzoi Balthasar, 1941 | | | Onthophagus veracruzensis Delgado & Pe..., 1998 | | | Onthophagus verae Josso, 2013 | | | Onthophagus verrucosus d´Orbigny, 1902 | | | Onthophagus versutus Péringuey, 1901 | | | Onthophagus verticalis (Fahraeus, 1857) | ⇒ Onthophagus praedatus Harold, 1862 | | Onthophagus vesanus Balthasar, 1967 | | | Onthophagus vespertilio Howden, Cart..., 1956 | | | Onthophagus vespillo Boucomont, 1919 | | | Onthophagus vestitus d´Orbigny, 1902 | | | Onthophagus vethi Krikken, 1977 | | | Onthophagus vicinus (Harold, 1886) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus vigens Péringuey, 1901 | | | Onthophagus villanuevai Delgado & De..., 1990 | | | Onthophagus vinctoides Frey, 1957 | | | Onthophagus vinctus Erichson, 1843 | | | Onthophagus violaceotinctus Gillet, 1925 | | | Onthophagus violaceoviridis Paulian, 1937 | | | Onthophagus violetae Zunino & Hal..., 1997 | | | Onthophagus virens (d´Orbigny, 1913) | ⇒ Onthophagus alcyonides d´Orbigny, 1913 | | Onthophagus virescens Harold, 1867 | | | Onthophagus viridiaereus d´Orbigny, 1908 | | | Onthophagus viridicatus d´Orbigny, 1902 | | | Onthophagus viridichevrolati Moctezuma & ..., 2020 | | | Onthophagus viridilimbatus (Olsoufieff, 1907) | ⇒ Onthophagus haroldi Ballion, 1870 | | Onthophagus viridiperakensis Ochi, Kon & ..., 2014 | | | Onthophagus viriditinctus Reitter, 1893 | | | Onthophagus viridivinosus Kohlmann & S..., 2001 | | | Onthophagus viviensis d´Orbigny, 1905 | | | Onthophagus vladimiri Frey, 1957 | | | Onthophagus volsellatus (Boucomont, 1920) | ⇒ Onthophagus minax d´Orbigny, 1904 | | Onthophagus volucer Balthasar, 1959 | | | Onthophagus vuattouxi Cambefort, 1984 | | | Onthophagus vulneratus (Mulsant, 1842) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus vulpes Harold, 1877 | | | Onthophagus vulpinus Arrow, 1931 | | | Onthophagus vultuosus d´Orbigny, 1902 | | | Onthophagus vultur Arrow, 1931 | | | Onthophagus vylderi d´Orbigny, 1913 | | | Onthophagus wallacei Harold, 1871 | | | Onthophagus wangnamkieous Masumoto, Oc..., 2008 | | | Onthophagus wangtaichieni Masumoto, Oc..., 2020 | | | Onthophagus waterhousei Gillet, 1927 | | | Onthophagus waterloti d´Orbigny, 1908 | | | Onthophagus waterstradti Boucomont, 1914 | | | Onthophagus watuwila Krikken & Hu..., 2008 | | | Onthophagus wayaua Huijbregts &..., 2012 | | | Onthophagus wensis Josso & Prév..., 2006 | | | Onthophagus wiebesi Krikken, 1977 | | | Onthophagus willameorum Walter & Cam..., 1977 | | | Onthophagus witteianus Krell, 2009 | | | Onthophagus woroae Ochi & Kon, 2006 | | | Onthophagus woroae bruneiensis Ochi, Kon & ..., 2009 | | | Onthophagus woroae woroae | | | Onthophagus xanthochlorus Walter & Cam..., 1977 | | | Onthophagus xanthomerus Bates, 1887 | | | Onthophagus xanthopterus d´Orbigny, 1908 | | | Onthophagus xanthopygus d´Orbigny, 1908 | | | Onthophagus xerxes Petrovitz, 1965 | | | Onthophagus xiphias (Solis & Kohl..., 2003) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus yaeyamanus Nomura, 1964 | | | Onthophagus yakuinsulanus Nakane, 1984 | | | Onthophagus yangmunensis Masumoto & Ochi, 2015 | | | Onthophagus yanoi Matsumura, 1938 | | | Onthophagus yaoi Masumoto, Oc..., 2014 | | | Onthophagus yasuhikoi Masumoto, Oc..., 2012 | | | Onthophagus yescaensis Moctezuma, H..., 2022 | | | Onthophagus yifer Krikken & Hu..., 2012 | | | Onthophagus yohenai (Matsumura, 1937) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus yucatanus Delgado-Cast..., 2006 | | | Onthophagus yugianus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus yujii Masumoto, Oc..., 2012 | | | Onthophagus yumigatanus Matsumura, 1937 | | | Onthophagus yumotoi Ochi & Kon, 2006 | | | Onthophagus yunnanus Boucomont, 1912 | | | Onthophagus zagrosicus Kabakov, 2006 | | | Onthophagus zairensis Walter & Cam..., 1977 | | | Onthophagus zambundaticeps Josso, 2022 | | | Onthophagus zapotecus Zunino & Hal..., 1988 | | | Onthophagus zavattarii Müller, 1947 | | | Onthophagus zavreli Balthasar, 1935 | | | Onthophagus zebra Arrow, 1931 | | | Onthophagus zebu Boucomont, 1921 | | | Onthophagus zicsii Balthasar, 1967 | | | Onthophagus zimmermanni Balthasar, 1959 | | | Onthophagus zinovskyi Qaryagdy, 1939 | | | Onthophagus zolotarevi (Olsoufieff, 1907) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus zubaci (Balthasar, 1932) | ⇒ Onthophagus dama Fabricius, 1798 | | Onthophagus zumpti Frey, 1954 | | | Onthophagus zuvandi Qaryagdy, 1939 | | | Onthophagus zymoticus Moxey, 1963 | |
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