|
|
| | | | Checkliste der Tribus Onthophagini Lacordaire, 1856 (Kotkäfer)
für Ferner Osten (Asien)
Klasse
| Insecta
Linnaeus, 1758
Freikiefler, Insekten, Kerbtiere, Kerfe | 1256
36222
255417 | Familien
Gattungen
Arten |
Taxom
|
Unterklasse
| Pterygota
Lang, 1888
Fluginsekten | 1250
36175
255323 | Familien
Gattungen
Arten |
Taxom
|
Teilklasse
| Neoptera
van der Wulp, 1890
Neuflügler | 1148
35562
252662 | Familien
Gattungen
Arten |
Taxom
|
Überordnung
| Holometabola
Heider, 1889
Holometabolen Insekten | 671
26426
204688 | Familien
Gattungen
Arten |
Taxom
|
Ordnung
| Coleoptera
Linnaeus, 1758
Käfer | 186
10806
97430 | Familien
Gattungen
Arten |
Taxom
|
Unterordnung
| Polyphaga
Emery, 1886
| 152
9707
84101 | Familien
Gattungen
Arten |
Taxom
|
Teilordnung
| Scarabaeiformia
Crowson, 1960
| 18
1070
13561 | Familien
Gattungen
Arten |
Taxom
|
Überfamilie
| Scarabaeoidea
Latreille, 1802
Blatthornkäfer | 18
1070
13561 | Familien
Gattungen
Arten |
Taxom
| | Familie | Scarabaeidae
Latreille, 1802
Blatthornkäfer | 795
10477 | Gattungen
Arten |
Taxom
| | Unterfamilie | Scarabaeinae
Latreille, 1802
Pillendreher | 66
1328 | Gattungen
Arten |
Taxom
| | Tribus | Onthophagini
Lacordaire, 1856
Kotkäfer | 14
878 | Gattungen
Arten |
Taxom
|
Gattungen für Ferner Osten
14 Gattungen mit 878 Arten | Anoctus Sharp, 1942 | | 4 | Arten | Taxom | | Caccobius Thomson, 1936 | | 39 | Arten | Taxom | | Cassolus Sharp, 1885 | | 9 | Arten | Taxom | | Cleptocaccobius Cambefort, 0 | | 4 | Arten | Taxom | | Cyobius Sharp, 1875 | | 5 | Arten | Taxom | | Diastellopalpus van Lansberge, 1884 | | 1 | Art | Taxom | | Digitonthophagus Balthasar, 1787 | | 3 | Arten | Taxom | | Euonthophagus Balthasar, 1910 | | 4 | Arten | Taxom | | Haroldius Boucomont, 1993 | | 27 | Arten | Taxom | | Hyalonthophagus Palestrini &..., 1913 | | 1 | Art | Taxom | | Milichus Péringuey, 1871 | | 1 | Art | Taxom | | Onthophagus Latreille, 1963 | Kotfresser | 809 | Arten | Taxom | | Phalops Erichson, 1847 | | 1 | Art | Taxom | | Proagoderus Lansberge, 1912 | | 10 | Arten | Taxom |
Artenliste für Ferner Osten
878 Arten und 37 Unterarten | Anoctus laevis Sharp, 1875 | | | Anoctus rasilis Boucomont, 1921 | | | Anoctus scatonomoides Lansberge, 1885 | | | Anoctus siamensis Balthasar, 1942 | | | Caccobius (Caccobius) chujoi Paulian, 1942 | | | Caccobius (Caccophilus) scrofa (Balthasar, 1942) | ⇒ Cleptocaccobius scrofa Balthasar, 1942 | | Caccobius (Caccophilus) tortus Sharp, 1875 | | | Caccobius tortus matsudai Ochi & Araya, 1996 | | | Caccobius tortus tortus | | | Caccobius (Caccophilus) unicornis Fabricius, 1798 | | | Caccobius bawangensis Ochi & Kon &..., 1997 | | | Caccobius bidentatus Boucomont, 1923 | | | Caccobius binodulus Harold, 1877 | | | Caccobius boucomonti Balthasar, 1935 | | | Caccobius brevis Waterhouse, 1875 | | | Caccobius castaneus (Klug, 1855) | ⇒ Caccocnemus castaneus Klug, 1855 | | Caccobius christophi Harold, 1879 | | | Caccobius demangei Boucomont, 1919 | | | Caccobius dickinsoni Masumoto, Oc..., 2022 | | | Caccobius excavatus Bai, Zhang &..., 2007 | | | Caccobius flavolimbatus Balthasar, 1942 | | | Caccobius fukiensis Balthasar, 1942 | | | Caccobius gilleti Balthasar, 1933 | | | Caccobius gonoderus Fairmaire, 1888 | | | Caccobius himalayanus Jekel, 1872 | | | Caccobius hirayamai (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccobius imitans Balthasar, 1933 | | | Caccobius jessoensis Harold, 1867 | | | Caccobius jononis (Matsumura, 1936) | ⇒ Gibbonthophagus viduus Harold, 1875 | | Caccobius kelleri Olsoufieff, 1907 | | | Caccobius maruyamai Masumoto, Oc..., 2012 | | | Caccobius masumotoi Cambefort, 1990 | | | Caccobius nikkoensis Lewis, 1895 | | | Caccobius pullus Jekel, 1872 | | | Caccobius sappoensis Matsumura, 1936 | | | Caccobius sapporensis (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccobius schreberi Linnaeus, 1767 | | | Caccobius sordidus Harold, 1886 | | | Caccobius suzukii (Matsumura, 1936) | ⇒ Caccobius boucomonti Balthasar, 1935 | | Caccobius tsunoellus (Matsumara, 1938) | ⇒ Caccophilus tortus Sharp, 1875 | | Caccobius yamauchii (Matsumura, 1936) | ⇒ Caccobius unicornis Fabricius, 1798 | | Caccobius yubariensis (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Cassolus fukiensis Balthasar, 1960 | | | Cassolus gotoi Masumoto, 1986 | | | Cassolus javanus Boucomont, 1914 | | | Cassolus minutus Boucomont, 1914 | | | Cassolus nudus Sharp, 1875 | | | Cassolus peninsularis Arrow, 1907 | | | Cassolus pongchaii Masumoto, 1989 | | | Cassolus sumatranus Lansberge, 1885 | | | Cleptocaccobius scrofa Balthasar, 1942 | | | Cleptocaccobius simplex Boucomont, 1923 | | | Cleptocaccobius simplex meridionalis Cambefort, 1985 | | | Cleptocaccobius simplex simplex | | | Cyobius brendelli Ochi, Kon & ..., 2010 | | | Cyobius cheyi Ochi, Kon & ..., 2006 | | | Cyobius mendeli Ochi, Kon & ..., 2010 | | | Cyobius tsukamotoi Ochi, Kon & ..., 2008 | | | Cyobius wallacei Sharp, 1875 | | | Diastellopalpus lamellicollis Quedenfeldt, 1884 | | | Digitonthophagus bonasus Fabricius, 1775 | Tranquebarische dreygehörnte Kopf | | Digitonthophagus gazella Fabricius, 1787 | | | Euonthophagus amyntas subviolaceus Ménétries, 1832 | | | Euonthophagus gibbosus Scriba, 1790 | | | Euonthophagus koshantschikoffi Reitter, 1891 | | | Euonthophagus schnabeli Splichal, 1910 | | | Haroldius borneensis Paulian, 1993 | | | Haroldius brendelli Krikken & Hu..., 2009 | | | Haroldius cambeforti Krikken & Hu..., 2006 | | | Haroldius celebensis Krikken & Hu..., 2006 | | | Haroldius discoidalis Paulian, 1993 | | | Haroldius fairmairei Boucomont, 1914 | | | Haroldius fleutiauxi Paulian, 1945 | | | Haroldius globosus Boucomont, 1925 | | | Haroldius hwangi Masumoto, Le..., 2005 | | | Haroldius jechaer Král & Hájek, 2012 | | | Haroldius kawadai Masumoto, 1995 | | | Haroldius kolaka Krikken & Hu..., 2006 | | | Haroldius loebli Paulian, 1987 | | | Haroldius maruyamai Utsunomiya &..., 2005 | | | Haroldius oharai Ochi, Kon & ..., 2009 | | | Haroldius pahangensis Kral, 2003 | | | Haroldius pauliani Scheuern, 1995 | | | Haroldius penelopae Krikken & Hu..., 2006 | | | Haroldius perroti Paulian, 1939 | | | Haroldius philippinensis Pereira, 1954 | | | Haroldius rugatulus Boucomont, 1914 | | | Haroldius stevensi Arrow, 1931 | | | Haroldius sumatranus Paulian & Sc..., 1994 | | | Haroldius tangkoko Krikken & Hu..., 2006 | | | Haroldius thailandensis Paulian & Sc..., 1994 | | | Haroldius turnai Kral, 2003 | | | Haroldius uenoi Masumoto & Yin, 1993 | | | Hyalonthophagus alcedo Orbigny, 1913 | | | Milichus picticollis Gerstaecker, 1871 | | | Onthophagus (Altonthophagus) cervenkai Kabakov, 2008 | | | Onthophagus (Altonthophagus) inelegans Balthasar, 1935 | | | Onthophagus (Altonthophagus) kozlovi Kabakov, 1990 | | | Onthophagus (Altonthophagus) kukunorensis Kabakov, 1990 | | | Onthophagus (Altonthophagus) sibiricus Harold, 1877 | | | Onthophagus (Altonthophagus) turpidoides Kabakov, 2008 | | | Onthophagus (Colobonthophagus) aenescens Wiedemann, 1823 | | | Onthophagus (Colobonthophagus) caprai Frey, 1956 | | | Onthophagus (Colobonthophagus) dama Fabricius, 1798 | | | Onthophagus (Colobonthophagus) occipitalis Lansberge, 1885 | | | Onthophagus (Colobonthophagus) piceorufulus Kabakov, 1994 | | | Onthophagus (Colobonthophagus) poggii Scheuern, 1996 | | | Onthophagus (Colobonthophagus) quadridentatus (Fabricius, 1798) | ⇒ Onthophagus fissicornis Stevens, 1809 | | Onthophagus (Colobonthophagus) thai Kabakov, 1994 | | | Onthophagus (Colobonthophagus) tragus Fabricius, 1792 | | | Onthophagus (Digitonthophagus) bonasus Fabricius, 1775 | | | Onthophagus (Euonthophagus) carbonarius Klug, 1855 | | | Onthophagus (Furconthophagus) boucomonti Paulian, 1931 | | | Onthophagus (Furconthophagus) furcatus (Fabricius, 1781) | ⇒ Onthophagus furcaticeps Masters, 1886 | | Onthophagus (Furconthophagus) karenensis Masumoto, Oc..., 2008 | | | Onthophagus (Furconthophagus) khonkaenus Masumoto, Oc..., 2008 | | | Onthophagus (Furconthophagus) lilliputanus Lansberge, 1883 | | | Onthophagus (Furconthophagus) papulatorius Kabakov, 2008 | | | Onthophagus (Furconthophagus) papulatus Boucomont, 1914 | | | Onthophagus (Furconthophagus) promontorii Boucomont, 1924 | | | Onthophagus (Gibbonthophagus) apicetinctus (d´Orbigny, 1900) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus (Gibbonthophagus) atripennis Waterhouse, 1875 | | | Onthophagus (Gibbonthophagus) chineicus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) denticollis Lansberge, 1883 | | | Onthophagus (Gibbonthophagus) duporti Boucomont, 1914 | | | Onthophagus (Gibbonthophagus) euryceros Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) fujiii Ochi & Kon, 1995 | | | Onthophagus (Gibbonthophagus) gracilipes Boucomont, 1914 | | | Onthophagus (Gibbonthophagus) hiabunicus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) kentingensis Nomura, 1973 | | | Onthophagus (Gibbonthophagus) kimioi Ochi, Kon & ..., 2011 | | | Onthophagus (Gibbonthophagus) lenis Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) limbatus Herbst, 1789 | | | Onthophagus (Gibbonthophagus) luridipennis Boheman, 1858 | | | Onthophagus (Gibbonthophagus) nigriobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) palawanicus Ochi & Kon, 2004 | | | Onthophagus (Gibbonthophagus) parviobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) penmani Masumoto, Oc..., 2002 | | | Onthophagus (Gibbonthophagus) privus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) rufiobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) sasajii Ochi & Kon, 2001 | | | Onthophagus (Gibbonthophagus) semipersonatus Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) sunantaae Masumoto, 1989 | | | Onthophagus (Gibbonthophagus) taurinus White, 1844 | | | Onthophagus (Gibbonthophagus) viduus (Harold, 1875) | ⇒ Onthophagus orbus Boucomont, 1924 | | Onthophagus (Gibbonthophagus) viridicervicapra Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) arai Masumoto, 1989 | | | Onthophagus (Indachorius) baykanus Kabakov, 1994 | | | Onthophagus (Indachorius) bonengus Kabakov, 1994 | | | Onthophagus (Indachorius) chinensis Balthasar, 1952 | | | Onthophagus (Indachorius) danumensis (Ochi, Kon & ..., 2009) | ⇒ Onthophagus danumicus Huijbregts &..., 2012 | | Onthophagus (Indachorius) doisuthepensis Masumoto, 1989 | | | Onthophagus (Indachorius) hsui Masumoto, Ch..., 2004 | | | Onthophagus (Indachorius) koshunensis Balthasar, 1941 | | | Onthophagus (Indachorius) lannamiibun Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) maesaensis Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) masaoi Ochi, 1992 | | | Onthophagus (Indachorius) mendeli Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) mongkhoni Masumoto, Ha..., 2002 | | | Onthophagus (Indachorius) nongkaiensis Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) octonaevus Kabakov, 1994 | | | Onthophagus (Indachorius) paramasaoi Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) phetchabunensis Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) phuquoci Paulian, 1945 | | | Onthophagus (Indachorius) platypus Zhang, 1997 | | | Onthophagus (Indachorius) pseudoworoae Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) scotti Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) semidanumensis Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) subsapaensis Kabakov, 1994 | | | Onthophagus (Indachorius) suginoi Ochi, 1984 | | | Onthophagus (Indachorius) tiamicus Kabakov, 1994 | | | Onthophagus (Indachorius) tongbantumi Masumoto, Oc..., 2002 | | | Onthophagus (Macronthophagus) cludtsi Ochi, 2003 | | | Onthophagus (Macronthophagus) curvicarinatus Boucomont, 1914 | | | Onthophagus (Macronthophagus) diabolicus Harold, 1877 | | | Onthophagus (Macronthophagus) manipurensis Arrow, 1907 | | | Onthophagus (Macronthophagus) menieri Ochi, 2003 | | | Onthophagus (Macronthophagus) rubricollis Hope, 1831 | | | Onthophagus (Macronthophagus) uenoi Ochi, 1995 | | | Onthophagus uenoi borneomontanus Ochi, 2003 | | | Onthophagus uenoi uenoi Ochi, 1995 | | | Onthophagus (Matashia) anguliceps (Boucomont, 1914) | ⇒ Onthophagus avocetta Arrow, 1933 | | Onthophagus (Matashia) yubarinus (Matsumura, 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus (Micronthophagus) falsivigilans Masumoto, 1995 | | | Onthophagus (Micronthophagus) gigantivigilans Masumoto, Ha..., 2002 | | | Onthophagus (Micronthophagus) konoi Matsumura, 1938 | | | Onthophagus (Micronthophagus) vigilans Boucomont, 1921 | | | Onthophagus (Micronthophagus) wangi Masumoto, Ch..., 2004 | | | Onthophagus (Onthophagiellus) kondaoensis Kabakov, 1994 | | | Onthophagus (Onthophagiellus) opacifalculatus Ochi, Kon & ..., 2009 | | | Onthophagus (Onthophagiellus) parafalculatus Ochi, Kon & ..., 2009 | | | Onthophagus (Onthophagiellus) popovi Kabakov, 1983 | | | Onthophagus (Onthophagiellus) taiyaruensis Masumoto, 1977 | | | Onthophagus (Onthophagiellus) thanwaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagiellus) unguiculatus Kabakov, 1994 | | | Onthophagus (Onthophagus) acuticollis (Gillet, 1927) | ⇒ Onthophagus amphioxus Arrow, 1933 | | Onthophagus (Onthophagus) acuticornis Endrödi, 1973 | | | Onthophagus (Onthophagus) anguicorius Boucomont, 1914 | | | Onthophagus (Onthophagus) angustatus Boucomont, 1914 | | | Onthophagus (Onthophagus) apilularius Masumoto, 1995 | | | Onthophagus (Onthophagus) barbieri Paulian, 1978 | | | Onthophagus (Onthophagus) batesi Howden and C..., 1963 | | | Onthophagus (Onthophagus) bokiaunus Masumoto, 1995 | | | Onthophagus (Onthophagus) chaiyaphumensis Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) dapcauensis Boucomont, 1921 | | | Onthophagus (Onthophagus) doiinthanonensis Masumoto, 1989 | | | Onthophagus (Onthophagus) doipuiensis Masumoto, 1989 | | | Onthophagus (Onthophagus) doitungensis Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) dubernardi Boucomont, 1914 | | | Onthophagus (Onthophagus) embersoni Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) fissicornis Stevens, 1809 | | | Onthophagus (Onthophagus) formosanus Gillet, 1924 | | | Onthophagus (Onthophagus) gibbulus Pallas, 1781 | | | Onthophagus (Onthophagus) gorochovi Kabakov, 1998 | | | Onthophagus (Onthophagus) gosoli Masumoto, 1989 | | | Onthophagus (Onthophagus) grandivigilans Masumoto, 1995 | | | Onthophagus (Onthophagus) hajimei Masumoto, 1984 | | | Onthophagus (Onthophagus) hastifer Lansberge, 1885 | | | Onthophagus (Onthophagus) hayashii Masumoto, 1991 | | | Onthophagus (Onthophagus) horii Ochi, Kon & ..., 2009 | | | Onthophagus (Onthophagus) illyricus Scopoli, 1763 | | | Onthophagus (Onthophagus) ishiii Ochi & Kon, 1995 | | | Onthophagus (Onthophagus) javaecola Balthasar, 1959 | | | Onthophagus (Onthophagus) kanyaayonus Masumoto, 1992 | | | Onthophagus (Onthophagus) khonmiinitnoi Masumoto, 1990 | | | Onthophagus (Onthophagus) kinhthaicus Kabakov, 1998 | | | Onthophagus (Onthophagus) kiuchii Masumoto, 1995 | | | Onthophagus (Onthophagus) koryoensis Kim, 1985 | | | Onthophagus (Onthophagus) krakadaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagus) kuatunensis Balthasar, 1942 | | | Onthophagus (Onthophagus) lakyim Masumoto, 1990 | | | Onthophagus (Onthophagus) lindaae Masumoto, 1989 | | | Onthophagus (Onthophagus) lutosopictus Fairmaire, 1897 | | | Onthophagus (Onthophagus) maleengnaafon Masumoto, 1990 | | | Onthophagus (Onthophagus) maleengnoi Masumoto, 1990 | | | Onthophagus (Onthophagus) maniti Masumoto, 1989 | | | Onthophagus (Onthophagus) masumotoi Ochi, 1985 | | | Onthophagus (Onthophagus) matsudai Ochi & Kon, 2004 | | | Onthophagus (Onthophagus) mendicus Gillet, 1924 | | | Onthophagus (Onthophagus) mulleri van Lansberge, 1883 | | | Onthophagus (Onthophagus) naaroon Masumoto, 1990 | | | Onthophagus (Onthophagus) namnaoensis Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) nasonis Kabakov, 1998 | | | Onthophagus (Onthophagus) orientalis (Harold, 1868) | ⇒ Onthophagus fissinasus Fairmaire, 1895 | | Onthophagus (Onthophagus) phraoensis Masumoto, 1990 | | | Onthophagus (Onthophagus) phrutsaphaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagus) proletarius Harold, 1875 | | | Onthophagus (Onthophagus) pseudoarmatus Balthasar, 1944 | | | Onthophagus (Onthophagus) pseudohystrix Masumoto, 1995 | | | Onthophagus (Onthophagus) punctator Reitter, 1893 | | | Onthophagus (Onthophagus) punneeae Masumoto, 1989 | | | Onthophagus (Onthophagus) ratchasimaensis Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) roubali Balthasar, 1935 | | | Onthophagus (Onthophagus) rugulosus Harold, 1886 | | | Onthophagus (Onthophagus) sakaeratensis Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) sakainoi Masumoto, 1991 | | | Onthophagus (Onthophagus) salvazai Paulian, 1978 | | | Onthophagus (Onthophagus) sauteri Gillet, 1924 | | | Onthophagus (Onthophagus) setchan Masumoto, 1984 | | | Onthophagus (Onthophagus) singhaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagus) sobrius Balthasar, 1959 | | | Onthophagus (Onthophagus) striatulus Palisot de B..., 1809 | | | Onthophagus (Onthophagus) taayai Masumoto, 1995 | | | Onthophagus (Onthophagus) tatsienluensis Balthasar, 1942 | | | Onthophagus (Onthophagus) taurus Schreber, 1759 | Oechslein | | Onthophagus (Onthophagus) terminatus Eschscholtz, 1822 | | | Onthophagus (Onthophagus) tholaayi Masumoto, 1990 | | | Onthophagus (Onthophagus) tritinctus Boucomont, 1914 | | | Onthophagus (Onthophagus) trituber Wiedemann, 1823 | | | Onthophagus trituber jacobsoni Kabakov, 2006 | | | Onthophagus (Onthophagus) tumami Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) wangnamkhieoensis Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) yamaokai Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) yangi Masumoto, Ts..., 2006 | | | Onthophagus (Palaeonthophagus) baraudi Nicolas, 1964 | | | Onthophagus (Palaeonthophagus) clitellifer Reitter, 1894 | | | Onthophagus (Palaeonthophagus) coenobita Herbst, 1783 | Mönchs-Kotkäfer | | Onthophagus (Palaeonthophagus) finschi Harold, 1877 | | | Onthophagus (Palaeonthophagus) fissicornis Steven, 1809 | | | Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790) | ⇒ Onthophagus similis Scriba, 1790 | | Onthophagus (Palaeonthophagus) gibbulus Pallas, 1781 | | | Onthophagus gibbulus gibbulus | | | Onthophagus (Palaeonthophagus) gorodinskii Kabakov, 2008 | | | Onthophagus (Palaeonthophagus) grossepunctatus Reitter, 1905 | | | Onthophagus (Palaeonthophagus) haroldi Ballion, 1870 | | | Onthophagus (Palaeonthophagus) joannae Goljan, 1953 | | | Onthophagus (Palaeonthophagus) laticornis Gebler, 1823 | | | Onthophagus (Palaeonthophagus) lemur Fabricius, 1781 | | | Onthophagus (Palaeonthophagus) marginalis Gebler, 1817 | | | Onthophagus marginalis marginalis | | | Onthophagus marginalis przewalski Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) medius (Kugelann, 1792) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus (Palaeonthophagus) muelleri (Lansberge, 1883) | ⇒ Onthophagus statzi Krell, 1990 | | Onthophagus (Palaeonthophagus) nikolajevi Kabakov, 2006 | | | Onthophagus (Palaeonthophagus) novaki Boucomont, 1927 | | | Onthophagus (Palaeonthophagus) nuchicornis Linnaeus, 1758 | Kleine Nackenhorn | | Onthophagus (Palaeonthophagus) olsoufieffi Boucomont, 1924 | | | Onthophagus (Palaeonthophagus) opacicollis Reitter, 1893 | | | Onthophagus (Palaeonthophagus) ovatus Linnaeus, 1767 | Eiförmiger Kotkäfer | | Onthophagus (Palaeonthophagus) pygargus Motschulsky, 1845 | | | Onthophagus (Palaeonthophagus) rachelis Martín Piera, 1985 | | | Onthophagus (Palaeonthophagus) ruficapillus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) scabriusculus Harold, 1873 | | | Onthophagus (Palaeonthophagus) semicornis Panzer, 1798 | Halbhörniger Kotkäfer | | Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767 | Kühlein | | Onthophagus (Palaeonthophagus) verticicornis (Laicharting, 1781) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus (Palaeonthophagus) vitulus (Fabricius, 1776) | ⇒ Onthophagus immundus Boheman, 1858 | | Onthophagus (Paleonthophagus) andalusicus Waltl, 1835 | | | Onthophagus (Paraphanaeomorphus) argyropygus Gillet, 1927 | | | Onthophagus (Paraphanaeomorphus) carinensis Boucomont, 1914 | | | Onthophagus (Paraphanaeomorphus) comottoides Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) janushevi Kabakov, 1983 | | | Onthophagus (Paraphanaeomorphus) napolovi Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) pseudojavanus Paulian, 1931 | | | Onthophagus (Paraphanaeomorphus) quadricolor Kabakov, 1983 | | | Onthophagus (Paraphanaeomorphus) semipiceus Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) strnadi Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) vaulogeri Boucomont, 1923 | | | Onthophagus (Parascatonomus) anceyi Boucomont, 1921 | | | Onthophagus (Parascatonomus) damaki Masumoto, Ha..., 2002 | | | Onthophagus (Parascatonomus) kiuchianus Masumoto, Ya..., 2004 | | | Onthophagus (Parascatonomus) koichii (Ochi & Araya, 1994) | ⇒ Caccobius jessoensis Harold, 1867 | | Onthophagus (Parascatonomus) leytensis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) miyakei Ochi & Araya, 1992 | | | Onthophagus (Parascatonomus) murasakianus Nomura, 1976 | | | Onthophagus (Parascatonomus) negrosensis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) pseudorudis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) quezonensis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) satoi Ochi, Shimad..., 2003 | | | Onthophagus (Parascatonomus) shizumui Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) takedai Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) takeshii Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) tricornis (Wiedemann, 1823) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus (Parascatonomus) yukae Masumoto, Oc..., 2002 | | | Onthophagus (Parentius) kabakovi Martin-Piera, 1985 | | | Onthophagus (Paronthophagus) hidakai Ochi & Kon, 1995 | | | Onthophagus (Phanaeomorphus) ater Waterhouse, 1875 | | | Onthophagus (Phanaeomorphus) cernyi (Balthasar, 1935) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus (Phanaeomorphus) johkii Ochi & Kon, 1994 | | | Onthophagus (Phanaeomorphus) lenzi Harold, 1875 | | | Onthophagus potanini hiurai Ochi, 1984 | | | Onthophagus (Phanaeomorphus) zetteli Masumoto, Oc..., 2002 | | | Onthophagus (Pseudophanaeomorphus) parachandrai Ochi, Kon & ..., 2009 | | | Onthophagus (Serrophorus) avocettoides Kabakov, 1983 | | | Onthophagus (Serrophorus) laevis (Harold, 1880) | ⇒ Onthophagus laevissimus d´Orbigny, 1905 | | Onthophagus laevis asiaticus Boucomont, 1919 | | | Onthophagus laevis laevis | | | Onthophagus laevis lampromelas Fairmaire, 1891 | | | Onthophagus (Serrophorus) oblongus Zhang, 1997 | | | Onthophagus (Serrophorus) rectecornutus Lansberge, 1883 | | | Onthophagus (Serrophorus) sagittarius Fabricius, 1775 | | | Onthophagus (Sinonthophagus) nampatensis Tarasov & Ka..., 2010 | | | Onthophagus (Strandius) changshouensis Zhang, 1997 | | | Onthophagus (Strandius) japonicus Harold, 1875 | | | Onthophagus (Strandius) kuraruanus (Matsumura, 1938) | ⇒ Onthophagus lenzi Harold, 1875 | | Onthophagus (Strandius) potanini R. C. Brusca..., 1990 | | | Onthophagus potanini hiurai Ochi, 1984 | | | Onthophagus potanini potanini | | | Onthophagus (Strandius) taiwanus Nomura, 1973 | | | Onthophagus (Sunenaga) streltsovi Tarasov & Ka..., 2010 | | | Onthophagus abei Ochi & Kon, 2015 | | | Onthophagus academus Balthasar, 1946 | | | Onthophagus aceroides Krikken & Hu..., 2013 | | | Onthophagus acrisius Balthasar, 1946 | | | Onthophagus aemulus Gillet, 1930 | | | Onthophagus aequiperus Matsumura & ..., 1937 | | | Onthophagus aereomaculatus Boucomont, 1914 | | | Onthophagus aereopictus Boucomont, 1914 | | | Onthophagus aereopictus aereopictus | | | Onthophagus aereopictus rubropictus Boucomont, 1924 | | | Onthophagus agilis (Matsumura, 1938) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus akhaus Masumoto, Oc..., 2007 | | | Onthophagus akirai (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus alfuricus Huijbregts &..., 2012 | | | Onthophagus allojavanus Huijbregts &..., 2011 | | | Onthophagus aloysiellus Zunino, 1977 | | | Onthophagus amamiensis Nomura, 1965 | | | Onthophagus amamiensis amamiensis | | | Onthophagus amamiensis matsumurai Ochi, Kawaha..., 2019 | | | Onthophagus ambang Krikken & Hu..., 2011 | | | Onthophagus ambang ambang | | | Onthophagus ambang morowali Krikken & Hu..., 2011 | | | Onthophagus amenthus Matsumura, 1937 | | | Onthophagus amphicoma Boucomont, 1914 | | | Onthophagus amphioxus Arrow, 1933 | | | Onthophagus amycoides Kabakov, 2014 | | | Onthophagus andonarensis Boucomont, 1914 | | | Onthophagus angkhanensis Masumoto, Oc..., 2013 | | | Onthophagus annulopunctatus Krikken & Hu..., 2017 | | | Onthophagus aphodioides Lansberge, 1883 | | | Onthophagus apomontanus Ochi, Kon & ..., 2023 | | | Onthophagus apunneea Masumoto, Oc..., 2007 | | | Onthophagus arayai Ochi & Kon, 2007 | | | Onthophagus ardoini Frey, 1971 | | | Onthophagus armatus Blanchard, 1853 | | | Onthophagus asiaticus Endrödi, 1973 | | | Onthophagus atropolitus d´Orbigny, 1902 | | | Onthophagus aureopilosus Boucomont, 1914 | | | Onthophagus australis Guérin-Ménev..., 1830 | | | Onthophagus avocetta Arrow, 1933 | | | Onthophagus axanicus Ochi, Kon & ..., 2021 | | | Onthophagus azusae Ochi & Kon, 2006 | | | Onthophagus babirussa Eschscholtz, 1822 | | | Onthophagus baenzigeri Masumoto, Oc..., 2007 | | | Onthophagus baeri Boucomont, 1914 | | | Onthophagus bakeri (Boucomont, 1919) | ⇒ Panelus bakeri Boucomont, 1921 | | Onthophagus balthasari Vsetecka, 1939 | | | Onthophagus bangueyensis Boucomont, 1914 | | | Onthophagus baolocensis Ochi & Kon, 2015 | | | Onthophagus baramtagal Ochi, Kon & ..., 2016 | | | Onthophagus basicruentatus Goidanich, 1926 | | | Onthophagus batillifer Harold, 1875 | | | Onthophagus batui Huijbregts &..., 2009 | | | Onthophagus begoniophilus Krikken & Hu..., 2017 | | | Onthophagus bellus d´Orbigny, 1905 | | | Onthophagus bicolensis Ochi & Kon, 2006 | | | Onthophagus bicolensis asin Ochi & Kon, 2006 | | | Onthophagus bicolensis bicolensis | | | Onthophagus bicuneus Kabakov, 2008 | | | Onthophagus bifasciatus Fabricius, 1781 | Zweibindiger Pillenkäfer | | Onthophagus biscarinulatus Huijbregts &..., 2012 | | | Onthophagus bison Boucomont, 1919 | | | Onthophagus bisscrutator Krikken & Hu..., 2017 | | | Onthophagus bistiniocelloides Krikken, 1986 | | | Onthophagus bituberoculus Krikken & Hu..., 2013 | | | Onthophagus bivertex Heyden, 1887 | | | Onthophagus blanchardi Harold, 1869 | | | Onthophagus blumei (Lansberge, 1883) | ⇒ Onthophagus avocetta Arrow, 1933 | | Onthophagus bongkudai Krikken & Hu..., 2017 | | | Onthophagus bonorae Zunino, 1976 | | | Onthophagus borneensis Harold, 1877 | | | Onthophagus borneotagal Ochi, Kon & ..., 2016 | | | Onthophagus boucomontianus Balthasar, 1935 | | | Onthophagus bourgognei Paulian, 1945 | | | Onthophagus brutus Arrow, 1931 | | | Onthophagus bunamin Matthews, 1972 | | | Onthophagus calcaratus Boucomont, 1920 | | | Onthophagus cameloides d´Orbigny, 1900 | | | Onthophagus campestris Masumoto, Ki..., 2018 | | | Onthophagus capitosus Harold, 1867 | | | Onthophagus carinulatus Harold, 1877 | | | Onthophagus carpophilus Pereira & Ha..., 1961 | | | Onthophagus caviceps Frey, 1962 | | | Onthophagus cervicapra Boucomont, 1914 | | | Onthophagus cervus (Fabricius, 1798) | ⇒ Onthophagus falsus Gillet, 1925 | | Onthophagus chandrai Ochi, 2007 | | | Onthophagus chebaicus Boucomont, 1919 | | | Onthophagus chetroiensis Masumoto, Oc..., 2012 | | | Onthophagus cheyi Ochi & Kon, 2006 | | | Onthophagus chiangdaolekus Masumoto, Oc..., 2020 | | | Onthophagus chibanus Matsumura & ..., 1937 | | | Onthophagus chinbucephalus Ochi & Kon, 2018 | | | Onthophagus chinvigilans Ochi & Kon, 2018 | | | Onthophagus chokakurianus Matsumura, 1938 | | | Onthophagus chosensis Matsumura, 1937 | | | Onthophagus chumphonensis Masumoto, Oc..., 2008 | | | Onthophagus chuzenjianus (Matsumura & ..., 1937) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus circulifer Arrow, 1931 | | | Onthophagus citreum Boucomont, 1919 | | | Onthophagus clermonti Paulian, 1931 | | | Onthophagus clivimerus Huijbregts &..., 2011 | | | Onthophagus cognatus Boucomont, 1921 | | | Onthophagus colffsi Lansberge, 1883 | | | Onthophagus collaris (Kabakov, 1983) | ⇒ Proagoderus loricatus Klug, 1855 | | Onthophagus collinsi Krikken & Hu..., 1987 | | | Onthophagus commottoides Kabakov, 1998 | | | Onthophagus comottoi Lansberge, 1885 | | | Onthophagus consentaneus Harold, 1867 | | | Onthophagus convexicollis Boheman, 1858 | | | Onthophagus coomani Paulian, 1931 | | | Onthophagus coprimorphoides Huijbregts &..., 2012 | | | Onthophagus coracinoides Kabakov, 1983 | | | Onthophagus coracinus Boucomont, 1914 | | | Onthophagus crassicollis Boucomont, 1913 | | | Onthophagus cribratus Lansberge, 1883 | | | Onthophagus cuneus Boucomont, 1924 | | | Onthophagus cupreiceps Arrow, 1907 | | | Onthophagus cupreopastillatus Ochi & Kon, 2006 | | | Onthophagus cupreotagal Ochi, Kon & ..., 2016 | | | Onthophagus curvispina Reitter, 1893 | | | Onthophagus danumicus Huijbregts &..., 2012 | | | Onthophagus dapitanensis Boucomont, 1919 | | | Onthophagus dayacus Boucomont, 1914 | | | Onthophagus deflexicollis Lansberge, 1883 | | | Onthophagus deliensis Lansberge, 1885 | | | Onthophagus deliensis bawangicus Ochi, Kon & ..., 2009 | | | Onthophagus deliensis deliensis | | | Onthophagus denticornis Boucomont, 1914 | | | Onthophagus depressicollis Boucomont, 1914 | | | Onthophagus difficilis (Le Guillou, 1844) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus dinoderus d´Orbigny, 1913 | | | Onthophagus dissentaneus Balthasar, 1964 | | | Onthophagus doriae Harold, 1877 | | | Onthophagus dorsofasciatus Fairmaire, 1893 | | | Onthophagus drescheri Paulian, 1939 | | | Onthophagus echinus Boucomont, 1914 | | | Onthophagus egenus (Harold, 1877) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus egurianus Matsumura, 1938 | | | Onthophagus eschscholtzi Boucomont, 1924 | | | Onthophagus euplynes Bates, 1888 | | | Onthophagus excavatus (Redtenbacher, 1848) | ⇒ Liatongus phanaeoides Westwood, 1839 | | Onthophagus falculatus Boucomont, 1914 | | | Onthophagus falsus Gillet, 1925 | | | Onthophagus fissinasus Fairmaire, 1895 | | | Onthophagus fodiens Waterhouse, 1875 | | | Onthophagus foedus Boucomont, 1914 | | | Onthophagus fossor Arrow, 1931 | | | Onthophagus frankrozendaali Huijbregts &..., 2012 | | | Onthophagus frontalis Raffray, 1877 | | | Onthophagus fukuyamai Ochi, Kon & ..., 2018 | | | Onthophagus fulvus Sharp, 1875 | | | Onthophagus fuscostriatus Boucomont, 1914 | | | Onthophagus gaesatus Boucomont, 1925 | | | Onthophagus gagates Hope, 1831 | | | Onthophagus gajo Krikken, 1977 | | | Onthophagus galeatus Boucomont, 1919 | | | Onthophagus gestroi (Harold, 1877) | ⇒ Onthophagus bellus d´Orbigny, 1905 | | Onthophagus gibbicollis Lansberge, 1885 | | | Onthophagus ginyunensis Vsetecka, 1942 | | | Onthophagus gonipa Krikken & Hu..., 2011 | | | Onthophagus griseoaeneus Lansberge, 1885 | | | Onthophagus hageni Lansberge, 1883 | | | Onthophagus hanboonsongae Masumoto, Oc..., 2021 | | | Onthophagus hartiniae Ochi & Kon, 2009 | | | Onthophagus heterorrhinus Lansberge, 1885 | | | Onthophagus heyrovskyi Vsetecka, 1943 | | | Onthophagus hielkemai Krikken & Hu..., 2011 | | | Onthophagus hikidai Ochi & Kon, 2006 | | | Onthophagus hikosanus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus hiranoyasutoshii Ochi & Kon, 2018 | | | Onthophagus hiroyukii Ochi, 2007 | | | Onthophagus hirsutulus Lansberge, 1883 | | | Onthophagus histeriformis Boucomont, 1914 | | | Onthophagus hoberlandti Balthasar, 1966 | | | Onthophagus hollowayi Krikken & Hu..., 2017 | | | Onthophagus holosericeus Harold, 1877 | | | Onthophagus holosericus Harold, 1877 | | | Onthophagus holzi Boucomont, 1914 | | | Onthophagus hosomai Ochi & Kon, 2014 | | | Onthophagus ibonus (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus ifugaoensis Ochi & Kon, 2006 | | | Onthophagus imbutus Sharp, 1875 | | | Onthophagus imperator Castelnau, 1840 | | | Onthophagus incisus Harold, 1877 | | | Onthophagus incollaris Kabakov, 2008 | | | Onthophagus indosinicus Boucomont, 1921 | | | Onthophagus inermivertex Boucomont, 1921 | | | Onthophagus infucatus Harold, 1877 | | | Onthophagus insulindicus Boucomont, 1914 | | | Onthophagus isanus Masumoto, Oc..., 2008 | | | Onthophagus itoi Nomura, 1976 | | | Onthophagus itoianus Masumoto, Oc..., 2022 | | | Onthophagus itonoborui Masumoto, Oc..., 2012 | | | Onthophagus iumienus Masumoto, Oc..., 2007 | | | Onthophagus jacobeus Boucomont, 1924 | | | Onthophagus javanensis Balthasar, 1963 | | | Onthophagus javanoides Huijbregts &..., 2011 | | | Onthophagus jeannelianus Paulian, 1945 | | | Onthophagus jeannelianus jeannelianus | | | Onthophagus jeannelianus shokhini Kabakov, 2014 | | | Onthophagus jedensis Matsumura, 1937 | | | Onthophagus jingping Masumoto, Oc..., 2007 | | | Onthophagus jiupengensis Masumoto, La..., 2017 | | | Onthophagus kachinicus Kabakov, 2006 | | | Onthophagus kaengkrachangus Masumoto, Oc..., 2008 | | | Onthophagus kandai (Matsumura, 1937) | ⇒ Onthophagus olsoufieffi Boucomont, 1924 | | Onthophagus kangeanus Paulian, 1936 | | | Onthophagus kaosoidowensis Masumoto, Oc..., 2008 | | | Onthophagus kapitensis Frey, 1971 | | | Onthophagus kashizakii Kon & Ochi, 2005 | | | Onthophagus katayamai Masumoto, Oc..., 2012 | | | Onthophagus kawaharai Ochi & Kon, 2007 | | | Onthophagus kawarinus (Matsumura & ..., 1937) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus keikoae Ochi & Kon, 2014 | | | Onthophagus kiyoshii Ochi & Kon, 2007 | | | Onthophagus klapperichi Balthasar, 1952 | | | Onthophagus kleinei Balthasar, 1935 | | | Onthophagus knapperti Krikken, 1981 | | | Onthophagus koechlei Masumoto, Oc..., 2011 | | | Onthophagus kogatanus (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus kolaka Huijbregts &..., 2009 | | | Onthophagus koma Matsumura, 1937 | | | Onthophagus komabellus Matsumura & ..., 1937 | | | Onthophagus komareki Balthasar, 1935 | | | Onthophagus kongkaewensis Masumoto, Oc..., 2012 | | | Onthophagus koni Ochi, 2007 | | | Onthophagus konsarnensis Masumoto, Oc..., 2008 | | | Onthophagus kontumicus Kabakov, 1983 | | | Onthophagus kozunonis (Matsumura, 1937) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus kulti Balthasar, 1952 | | | Onthophagus kuluensis Bates, 1891 | | | Onthophagus kyokoae Masumoto & Ochi, 2015 | | | Onthophagus laevigatus Fabricius, 1798 | | | Onthophagus laevissimus d´Orbigny, 1905 | | | Onthophagus laichauensis Ochi, Kon & ..., 2021 | | | Onthophagus laminatus (MacLeay, 1864) | ⇒ Onthophagus lunatus Harold, 1868 | | Onthophagus lamyi Paulian, 1945 | | | Onthophagus langkawiensis Ochi & Kon, 2015 | | | Onthophagus laratinus Arrow, 1916 | | | Onthophagus lekmaengtibanphrao Masumoto & Ochi, 2015 | | | Onthophagus leknamnaous Masumoto & Ochi, 2015 | | | Onthophagus lenzii Harold, 1874 | | | Onthophagus leomontanus Boucomont, 1914 | | | Onthophagus leusermontis Huijbregts &..., 2011 | | | Onthophagus lindu Huijbregts &..., 2009 | | | Onthophagus liui Masumoto, Oc..., 2014 | | | Onthophagus liwagensis Ochi & Kon, 2006 | | | Onthophagus lore Huijbregts &..., 2009 | | | Onthophagus loricatus Klug, 1855 | | | Onthophagus luctuosus Boucomont, 1914 | | | Onthophagus lunatus Harold, 1868 | | | Onthophagus lunulifer Boucomont, 1914 | | | Onthophagus luteosignatus Lansberge, 1883 | | | Onthophagus luzonwallacei Ochi, Kawaha..., 2021 | | | Onthophagus maephaluangus Masumoto, Oc..., 2007 | | | Onthophagus maesalongensis Masumoto, Oc..., 2013 | | | Onthophagus magnini Paulian, 1933 | | | Onthophagus magnioculus Ochi & Kon, 2006 | | | Onthophagus magnipygus Boucomont, 1914 | | | Onthophagus malangensis Boucomont, 1914 | | | Onthophagus malasiacus Gillet, 1927 | | | Onthophagus malthinus Gillet, 1930 | | | Onthophagus manguliensis Boucomont, 1914 | | | Onthophagus maruchanus Masumoto, Oc..., 2012 | | | Onthophagus maruyamai Ochi, Kon & ..., 2014 | | | Onthophagus maryatiae Ochi & Kon, 2005 | | | Onthophagus matanyo Huijbregts &..., 2009 | | | Onthophagus matsuii Ochi & Kon, 2006 | | | Onthophagus matsuokai (Matsumura & ..., 1937) | ⇒ Onthophagus sauteri Gillet, 1924 | | Onthophagus megapacificus Ochi & Kon, 2006 | | | Onthophagus megathorax Gillet, 1921 | | | Onthophagus mekara Huijbregts &..., 2009 | | | Onthophagus mentaveiensis Boucomont, 1914 | | | Onthophagus micellus Matsumura, 1937 | | | Onthophagus midorianus (Matsumura, 1938) | ⇒ Parascatonomus anguicorius Boucomont, 1914 | | Onthophagus miichunus Masumoto, Oc..., 2020 | | | Onthophagus mimikanus Balthasar, 1969 | | | Onthophagus mindanaensis Boucomont, 1914 | | | Onthophagus mindanaoensis Frey, 1971 | | | Onthophagus minoi Matsumura & ..., 1937 | | | Onthophagus minokuchianus (Matsumura, 1934) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus misujianus Matsumura & ..., 1937 | | | Onthophagus miyabei (Matsumura, 1937) | ⇒ Onthophagus punctator Reitter, 1893 | | Onthophagus miyazakianus Matsumura & ..., 1937 | | | Onthophagus mniszechi Harold, 1869 | | | Onthophagus moajat Huijbregts &..., 2009 | | | Onthophagus mogo Krikken & Hu..., 2008 | | | Onthophagus montishannoniae Krikken & Hu..., 2008 | | | Onthophagus mordindangus Masumoto, Oc..., 2008 | | | Onthophagus morimotoi Masumoto, Oc..., 2021 | | | Onthophagus mouhoti Harold, 1875 | | | Onthophagus multipunctatus Boucomont, 1914 | | | Onthophagus mulutagal Ochi, Kon & ..., 2016 | | | Onthophagus mushanus Matsumura, 1938 | | | Onthophagus mushensis Matsumura, 1938 | | | Onthophagus nagasawai Matsumura, 1938 | | | Onthophagus nakatomii (Matsumura, 1937) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus namnaolekus Masumoto, Oc..., 2020 | | | Onthophagus namnaonus Masumoto, Oc..., 2020 | | | Onthophagus namnaous Masumoto, Oc..., 2013 | | | Onthophagus necessarius Reitter, 1893 | | | Onthophagus neervoorti Boucomont, 1914 | | | Onthophagus neofurcatus Goidanich, 1926 | | | Onthophagus nitefactus Harold, 1877 | | | Onthophagus nitidulus Klug, 1845 | | | Onthophagus nitidus Waterhouse, 1875 | | | Onthophagus obliviosus Balthasar, 1935 | | | Onthophagus obscurior Boucomont, 1914 | | | Onthophagus ocellatopunctatus Waterhouse, 1875 | | | Onthophagus ochii Masumoto, 1988 | | | Onthophagus ochromerus Harold, 1877 | | | Onthophagus ohbayashii Nomura, 1939 | | | Onthophagus ohkuboi Ochi & Kon, 2006 | | | Onthophagus oishii Matsumura & ..., 1937 | | | Onthophagus okushiranus Matsumura & ..., 1937 | | | Onthophagus okushirianus Matsumura, 1937 | | | Onthophagus oniellus (Matsumura, 1937) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus ontosatu Huijbregts &..., 2012 | | | Onthophagus opacihartiniae Ochi & Kon, 2015 | | | Onthophagus orbus Boucomont, 1924 | | | Onthophagus oshimanus Nakane, 1960 | | | Onthophagus otai Ochi & Kon, 2006 | | | Onthophagus pacificus Lansberge, 1885 | | | Onthophagus palatus Boucomont, 1914 | | | Onthophagus palavanus (Huijbregts &..., 2011) | ⇒ Onthophagus javanensis Balthasar, 1963 | | Onthophagus palawantagal Ochi, Kon & ..., 2016 | | | Onthophagus pangmaphanus Masumoto, Oc..., 2022 | | | Onthophagus paracenricornis Ochi & Kon, 2014 | | | Onthophagus paracentricornis Ochi & Kon, 2014 | | | Onthophagus parapalatus Krikken & Hu..., 1988 | | | Onthophagus parenthesis Boucomont, 1913 | | | Onthophagus paroculus Krikken & Hu..., 1987 | | | Onthophagus parryi Harold, 1869 | | | Onthophagus parvidens (Frey, 1971) | ⇒ Onthophagus clermonti Paulian, 1931 | | Onthophagus parvulus (Fabricius, 1798) | ⇒ Onthophagus frontalis Raffray, 1877 | | Onthophagus pasotagal Ochi, Kon & ..., 2016 | | | Onthophagus pastillatus Boucomont, 1919 | | | Onthophagus pauper Boucomont, 1914 | | | Onthophagus pavidus Harold, 1877 | | | Onthophagus pedator Sharp, 1875 | | | Onthophagus peninsularis Boucomont, 1914 | | | Onthophagus peninsulomerus Huijbregts &..., 2011 | | | Onthophagus perakensis Ochi, Kon & ..., 2014 | | | Onthophagus personatus Boucomont, 1914 | | | Onthophagus phahompokus Masumoto & Ochi, 2015 | | | Onthophagus phakuaiensis Masumoto, Oc..., 2013 | | | Onthophagus phanaeicollis Lansberge, 1883 | | | Onthophagus phanaeides Frey, 1956 | | | Onthophagus phanaeiformis Boucomont, 1914 | | | Onthophagus phanaeoides (Westwood, 1839) | ⇒ Proagoderus lanista Castelnau, 1840 | | Onthophagus phatoensis Masumoto, Oc..., 2008 | | | Onthophagus philippinensis Boucomont, 1919 | | | Onthophagus phillippsorum Krikken & Hu..., 1987 | | | Onthophagus phukhieoensis Masumoto, Oc..., 2007 | | | Onthophagus pimpasaleei Masumoto, Oc..., 2008 | | | Onthophagus planicollis Harold, 1880 | | | Onthophagus poeophagus Kabakov, 1979 | | | Onthophagus pollicatus Harold, 1879 | | | Onthophagus poringensis Ochi & Kon, 2005 | | | Onthophagus praedatus Harold, 1862 | | | Onthophagus procurvus Balthasar, 1935 | | | Onthophagus productus Arrow, 1907 | | | Onthophagus pseudoarai Ochi & Kon, 2018 | | | Onthophagus pseudobrutus Kabakov, 1983 | | | Onthophagus pseudoconvexicollis Bai & Yang, 2016 | | | Onthophagus pseudocoracinus Kabakov, 1983 | | | Onthophagus pseudojaponicus Balthasar, 1941 | | | Onthophagus pseudojaponicus fukiensis Balthasar, 1941 | | | Onthophagus pseudojaponicus pseudojaponicus | | | Onthophagus ptox Erichson, 1842 | | | Onthophagus pumilus Kabakov, 2014 | | | Onthophagus pupillatus Kolbe, 1886 | | | Onthophagus purpurascens Boucomont, 1914 | | | Onthophagus putealis (Matsumura, 1938) | ⇒ Onthophagus hastifer Lansberge, 1885 | | Onthophagus pygidialis Lansberge, 1883 | | | Onthophagus quadripustulatus Fabricius, 1775 | Neuholländische Vierblatter | | Onthophagus quangnamensis Ochi, Kon & ..., 2021 | | | Onthophagus quasijohkii Ochi & Kon, 2005 | | | Onthophagus quasitagal Ochi & Kon, 2005 | | | Onthophagus ramosellus Bates, 1891 | | | Onthophagus ranongensis Masumoto, Oc..., 2008 | | | Onthophagus ranongjohkii Ochi, Kon & ..., 2016 | | | Onthophagus ratchataniensis Masumoto, Oc..., 2013 | | | Onthophagus remotus Harold, 1879 | | | Onthophagus rhyncophorus | | | Onthophagus ribbei Boucomont, 1914 | | | Onthophagus rizali Ochi, Kawaha..., 2021 | | | Onthophagus robertopoggii Ochi & Kon, 2006 | | | Onthophagus rorarius Harold, 1877 | | | Onthophagus rosenbergi Krikken & Hu..., 2011 | | | Onthophagus rosenbergi divergens Krikken & Hu..., 2011 | | | Onthophagus rosenbergi rosenbergi | | | Onthophagus rotundicollis Lansberge, 1883 | | | Onthophagus rouyeri Boucomont, 1914 | | | Onthophagus rubripennis (Arrow, 1907) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus ruficauda Arrow, 1931 | | | Onthophagus rugicollis Harold, 1880 | | | Onthophagus rutilans (Sharp, 1875) | ⇒ Onthophagus borneensis Harold, 1877 | | Onthophagus rutilans aborneensis Ochi, Kon & ..., 2009 | | | Onthophagus rutilans rutilans | | | Onthophagus sabahensis Ochi & Kon, 2006 | | | Onthophagus sabai Masumoto, Oc..., 2012 | | | Onthophagus saigonensis Boucomont, 1924 | | | Onthophagus saksphus Masumoto, Oc..., 2021 | | | Onthophagus samarensis Ochi, Kon & ..., 2023 | | | Onthophagus samoengus Masumoto, Oc..., 2008 | | | Onthophagus sanggona Huijbregts &..., 2009 | | | Onthophagus sangirensis Boucomont, 1914 | | | Onthophagus sangwalus Masumoto, Oc..., 2008 | | | Onthophagus sarasinorum Krikken & Hu..., 2011 | | | Onthophagus sceptrifer Boucomont, 1924 | | | Onthophagus schaefernai Balthasar, 1935 | | | Onthophagus schillhammeri Kabakov, 2006 | | | Onthophagus sciron Balthasar, 1969 | | | Onthophagus scrutator Harold, 1877 | | | Onthophagus sembeli Krikken & Hu..., 2008 | | | Onthophagus semiaereopictus Ochi, Kawaha..., 2021 | | | Onthophagus semigranosus Lansberge, 1883 | | | Onthophagus semipacificus Ochi & Kon, 2006 | | | Onthophagus semiperakensis Ochi, Kon & ..., 2014 | | | Onthophagus semiworoae Ochi, Kon & ..., 2014 | | | Onthophagus senescens Péringuey, 1908 | | | Onthophagus senex Boucomont, 1914 | | | Onthophagus seniculus (Fabricius, 1781) | ⇒ Onthophagus senescens Péringuey, 1908 | | Onthophagus sepilokensis Ochi & Kon, 2006 | | | Onthophagus seramicus Huijbregts &..., 2012 | | | Onthophagus serdangensis Lansberge, 1886 | | | Onthophagus seseba Krikken & Hu..., 2017 | | | Onthophagus setoculus Krikken & Hu..., 1987 | | | Onthophagus sexdentatus Boucomont, 1919 | | | Onthophagus shibatai Nakane, 1960 | | | Onthophagus shigeo (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus shigeoi Matsumura, 1937 | | | Onthophagus shinanensis (Matsumura, 1934) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus shinichii Ochi & Kon, 2018 | | | Onthophagus shirakii Nakane, 1960 | | | Onthophagus shoyozanus (Matsumura, 1937) | ⇒ Onthophagus gibbulus Pallas, 1781 | | Onthophagus shunichii Ochi, Kon & ..., 2021 | | | Onthophagus shurianus Matsumura & ..., 1937 | | | Onthophagus sibela Huijbregts &..., 2012 | | | Onthophagus sibuyanus Boucomont, 1924 | | | Onthophagus sidangeaus Masumoto, Oc..., 2021 | | | Onthophagus sideki Krikken & Hu..., 1987 | | | Onthophagus sihkahonoi Huijbregts &..., 2009 | | | Onthophagus simboroni Ochi & Kon, 2006 | | | Onthophagus simplicifrons Reitter, 1893 | | | Onthophagus sinagai Krikken & Hu..., 2008 | | | Onthophagus sinicus Zhang & Wang, 1997 | | | Onthophagus smetanai Balthasar, 1952 | | | Onthophagus sobosanus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus solivagus Harold, 1886 | | | Onthophagus sonani Miwa, 1930 | | | Onthophagus sopu Krikken & Hu..., 2017 | | | Onthophagus spathatus Boucomont, 1914 | | | Onthophagus spiculatus Boucomont, 1914 | | | Onthophagus spinifex Fabricius, 1781 | | | Onthophagus spitsbergeniensis Krell, 2010 | | | Onthophagus statzi ♰ Krell, 1990 | | | Onthophagus sternax Balthasar, 1959 | | | Onthophagus strandi Balthasar, 1935 | | | Onthophagus subcornutus Boucomont, 1914 | | | Onthophagus sugihartoi Ochi, 2007 | | | Onthophagus suginokoichii Ochi & Kon, 2008 | | | Onthophagus sulawesiensis Krikken & Hu..., 2011 | | | Onthophagus sulawesijohkii Ochi, Kon & ..., 2016 | | | Onthophagus sulci Balthasar, 1935 | | | Onthophagus sumatramontanus Ochi & Kon, 2008 | | | Onthophagus sumatranus Lansberge, 1883 | | | Onthophagus sundanensis Lansberge, 1883 | | | Onthophagus surdus Boucomont, 1925 | | | Onthophagus susterai Balthasar, 1952 | | | Onthophagus sutleinensis Splichal, 1910 | | | Onthophagus sycophanta (Fairmaire, 1887) | ⇒ Euonthophagus gibbosus Scriba, 1790 | | Onthophagus tabidus Balthasar, 1935 | | | Onthophagus taeniatus Boucomont, 1914 | | | Onthophagus tagal Boucomont, 1924 | | | Onthophagus takabayashii Matsumura, 1937 | | | Onthophagus takaoi Ochi & Kon, 2014 | | | Onthophagus tambing Krikken & Hu..., 2011 | | | Onthophagus taoi Ochi & Kon, 2006 | | | Onthophagus thailaevis Masumoto, Oc..., 2008 | | | Onthophagus thainuaensis Masumoto, Oc..., 2008 | | | Onthophagus thungphraensis Masumoto, Oc..., 2020 | | | Onthophagus tibetanus Arrow, 1907 | | | Onthophagus timorensis Boucomont, 1914 | | | Onthophagus tiniocelloides Boucomont, 1925 | | | Onthophagus tonkineus Gillet, 1921 | | | Onthophagus tonywhitteni Krikken & Hu..., 2011 | | | Onthophagus toraut Krikken & Hu..., 2011 | | | Onthophagus traginus Frey, 1972 | | | Onthophagus tragoides Boucomont, 1914 | | | Onthophagus transvestitus Huijbregts &..., 2009 | | | Onthophagus tricolor Boucomont, 1914 | | | Onthophagus tridentitibialis Ochi & Kon, 2008 | | | Onthophagus trochilus Arrow, 1931 | | | Onthophagus tsubakii Ochi & Kon, 2009 | | | Onthophagus tsutomui Ochi & Kon, 2016 | | | Onthophagus tungkamangensis Masumoto, Oc..., 2007 | | | Onthophagus tungkamungensis Masumoto, Oc..., 2008 | | | Onthophagus turfanicus Balthasar, 1971 | | | Onthophagus turpidus Reitter, 1887 | | | Onthophagus ubaidillahi Krikken & Hu..., 2012 | | | Onthophagus uedai Ochi & Kon, 2006 | | | Onthophagus uedanus (Matsumura, 1937) | ⇒ Onthophagus olsoufieffi Boucomont, 1924 | | Onthophagus ulugombakensis Ochi, Kon & ..., 2014 | | | Onthophagus unidentatus Boucomont, 1927 | | | Onthophagus uniformis Heyden, 1886 | | | Onthophagus urellus Boucomont, 1919 | | | Onthophagus ushiodai (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus vanasseni Krikken & Hu..., 2008 | | | Onthophagus vanderblomi Krikken & Hu..., 2008 | | | Onthophagus vanofwegeni Krikken & Hu..., 2008 | | | Onthophagus variolaris Lansberge, 1883 | | | Onthophagus ventralis Lansberge, 1883 | | | Onthophagus vethi Krikken, 1977 | | | Onthophagus viridiperakensis Ochi, Kon & ..., 2014 | | | Onthophagus volucer Balthasar, 1959 | | | Onthophagus vulpes Harold, 1877 | | | Onthophagus vulpinus Arrow, 1931 | | | Onthophagus wallacei Harold, 1871 | | | Onthophagus wangnamkieous Masumoto, Oc..., 2008 | | | Onthophagus wangtaichieni Masumoto, Oc..., 2020 | | | Onthophagus waterstradti Boucomont, 1914 | | | Onthophagus watuwila Krikken & Hu..., 2008 | | | Onthophagus wayaua Huijbregts &..., 2012 | | | Onthophagus wiebesi Krikken, 1977 | | | Onthophagus woroae Ochi & Kon, 2006 | | | Onthophagus woroae bruneiensis Ochi, Kon & ..., 2009 | | | Onthophagus woroae woroae | | | Onthophagus yakuinsulanus Nakane, 1984 | | | Onthophagus yangmunensis Masumoto & Ochi, 2015 | | | Onthophagus yanoi Matsumura, 1938 | | | Onthophagus yaoi Masumoto, Oc..., 2014 | | | Onthophagus yasuhikoi Masumoto, Oc..., 2012 | | | Onthophagus yifer Krikken & Hu..., 2012 | | | Onthophagus yugianus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus yujii Masumoto, Oc..., 2012 | | | Onthophagus yumigatanus Matsumura, 1937 | | | Onthophagus yumotoi Ochi & Kon, 2006 | | | Onthophagus zavreli Balthasar, 1935 | | | Onthophagus zebu Boucomont, 1921 | | | Onthophagus zimmermanni Balthasar, 1959 | | | Onthophagus zymoticus Moxey, 1963 | | | Proagoderus amplexus Sharp, 1875 | | | Proagoderus gladiator Arrow, 1907 | | | Proagoderus lanista Castelnau, 1840 | | | Proagoderus loricatus Klug, 1855 | | | Proagoderus mouhoti Harold, 1875 | | | Proagoderus panoplus Bates, 1888 | | | Proagoderus schwaneri Snellen van ..., 1864 | | | Proagoderus victor Sharp, 1875 | | | Proagoderus watanabei Ochi & Kon, 2002 | | | Proagoderus yunnanus Boucomont, 1912 | |
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